Monday, April 27, 2020

[Herpetology • 2020] Cnemaspis tubaensis • A New Species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia

 Cnemaspis tubaensis
Quah, Wood, Anuar & Muin, 2020

 Tuba Island Rock Gecko  ||  DOI: 10.11646/zootaxa.4767.1.6  

A new, diminutive species of Rock Gecko Cnemaspis tubaensis sp. nov. of the C. kumpoli group, is described from Tuba Island, Langkawi Archipelago, Kedah, Peninsular Malaysia and is differentiated from all other species in the kumpoli group by having a unique combination of morphological and color pattern characteristics, including a maximum SVL of 37.0 mm; 10 or 11 supralabials; eight or nine infralabials; 15–18 semi-linearly arranged paravertebral tubercles; lateral caudal furrow present; lateral caudal tubercles on the anterior portion of the tail; caudal tubercles not encircling tail; five or six precloacal pores; 28 or 29 subdigital lamellae on the fourth toe; smooth ventrals; smooth subcaudals with an enlarged median row of scales; subcaudal region light-grey and speckled with yellow; absence of light-colored ocelli on the shoulder; no yellow postscapular band; dorsum light-brown with sage-green blotches and black spots; flanks with scattered yellow spots; absence of black gular markings in both sexes; and 13.0–22.0% uncorrected pairwise sequence divergences in the NADH dehydrogenase subunit 2 gene (ND2). Cnemaspis tubaensis sp. nov. is the fourth species of Cnemaspis to be described from the Langkawi Archipelago and underscores the underestimated biodiversity of the islands which is in need of more thorough herpetological inventories.

Keywords: Reptilia, Integrative taxonomy, phylogeny, molecular systematics, karst, island, conservation, herpetology, reptile, amphibian, discovery 

Cnemaspis tubaensis sp. nov.  
(A) Dorsal and (B) ventral views of the male holotype (USMHC 2541). 
(C) Dorsal view of the male paratype (USMHC 2527).

Cnemaspis tubaensis sp. nov. 
 Tuba Island Rock Gecko

Distribution. Cnemaspis tubaensis sp. nov. is only known from the northeastern corner of Tuba Island, Langkawi Archipelago, Kedah on the karst towers where the Gua Wang Buluh Cave is located. It may be wider ranging on the island where other karst outcrops occur. 

Etymology. The specific epithet tubaensis is in reference to the type locality of this species on Tuba Island of the Langkawi Archipelago, Kedah, Peninsular Malaysia (Fig. 1). 

Evan S.H. Quah, Perry L. Jr. Wood, M.S. Shahrul Anuar and Mohd Abdul Muin. 2020. A New Species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia with An Updated Checklist of the Herpetofauna of Tuba Island. Zootaxa. 4767(1); 138–160. DOI: 10.11646/zootaxa.4767.1.6