[Crustacea • 2026] Cambarus emegi • A New Stream-dwelling Species of Cambarus (Decapoda: Cambaridae) from Virginia and Pennsylvania (USA) with Notes on its Distribution, Ecology, and Conservation Status

 

Cambarus emegi 

Lieb, Williams, Foltz & Loughman, 2026 

 

DOI: doi.org/10.11646/zootaxa.5857.2.5 

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Abstract

A new species of crayfish, Cambarus emegi sp. nov., is described from the upper James and Roanoke River basins in Virginia and the lower Delaware River basin in Pennsylvania, where it occurs in streams and small rivers. The species was formerly part of the Cambarus acuminatus species complex (Cambarus species C complex), a long-standing taxonomic enigma that is widely distributed across the Atlantic slope drainages of eastern North America. Cambarus emegi sp. nov. can be distinguished from Cambarus acuminatus due to the absence of a well-defined subpalmar tubercle, greater number of punctations across its areola, and differences in the mesial process of the form I male gonopod. Coloration and pattern in life differ between the two species. Additionally, the distributions of C. emegi sp. nov. and C. acuminatus are separated by hundreds of kilometers. The ranges of two other undescribed members of the Cambarus species C complex border that of C. emegi sp. nov. in Virginia. These species do not appear to overlap geographically; C. emegi sp. nov. is confined to the Ridge and Valley physiographic province, whereas the two undescribed species are found in the Blue Ridge and northern inner Piedmont regions. Cambarus emegi sp. nov. is morphologically similar but visually and genetically different from the undescribed species directly to the south in the upper Roanoke River system. Cambarus emegi sp. nov. is both visually and morphologically similar but genetically distinct from the undescribed species immediately downstream in the James River basin. Cambarus emegi sp. nov. is threatened by several potentially interacting factors including invasive crayfishes, narrow range, and urban and agricultural development. Alarmingly, the species is being rapidly replaced by invasive Faxonius species across a substantial part of its range. This paper represents an important step in providing the information needed to conserve and manage Cambarus emegi sp. nov.

Crustacea, Cambarus species C., Cambarus acuminatus, new species, Ridge and Valley, Northern Piedmont, Delaware, James, Roanoke, invasive crayfish

 Cambarus emegi sp. nov., color in life: (a) holotype, NCSM 95987; (b) allotype, NCSM 95988.

Life color and color pattern of adult  Cambarus emegi sp. nov.:
 (a) dorsal view; (b) oblique lateral view (photo credit Zackary A. Graham).

 Life color and color pattern of juvenile  Cambarus emegi sp. nov. 

Cambarus emegi sp. nov. 

Diagnosis.—Body and eyes pigmented. Rostrum weakly to moderately excavated and ventrally deflected anteriorly; rostral margins subparallel, pronounced and entire through acumen, and more thickened posteriorly than anteriorly; acumen acuminate with prominent dorsally deflected spiniform tubercle at terminus. Areola 2.2–3.6 (¯x=2.7, n=42, s=0.4) times as long as wide, with 6–10 (¯x=8.5, n=42, s= 0.7) punctations across narrowest point. Lateral portion of branchiostegal region of carapace heavily punctate; hepatic regions also punctate with welldeveloped tubercles. Strong cervical spines present; branchiostegal spine prominent; postorbital ridge terminating anteriorly in spiniform tubercle or spine. Weak to acute suborbital angle typically present, rarely absent; suborbital angle terminating anteriorly in spiniform tubercle or spine. Antennal basipodite spines prominent; ischiopodite spines present but not prominent.

Etymology.—Derived from the Turkish work “emeği” meaning “labor of” or “hard work of”, which seems appropriate given this project required many years and the collection of thousands of specimens from South Carolina to Pennsylvania. Additionally, the name honors Emily (Em) and Megan (Meg) Lieb, the senior author’s two daughters, who happily accompanied him on many crayfish sampling trips as children and later helped him collect C. emegi sp. nov. from sites in its Pennsylvania range. 

Common name.—We propose Enigma Crayfish as the common name.

David A. LIEB, Bronwyn W. WILLIAMS, David A. FOLTZ II and Zachary J. LOUGHMAN. 2026. Description of A New Stream-dwelling Species of Cambarus (Decapoda: Cambaridae) from Virginia and Pennsylvania (USA) with Notes on its Distribution, Ecology, and Conservation Status.  Zootaxa. 5857(2); 299-316. DOI: doi.org/10.11646/zootaxa.5857.2.5 [2026-07-29]

 facebook.com/WLUCrayfish/posts/1731022025695189 

[Arachnida • 2026] Kwonkan elatus & K. yorkrakine • Wishbone Spiders of the Genus Kwonkan Main, 1983 (Araneae: Mygalomorphae: Anamidae) in south-western Australia: Redescription of legacy species, Two New Species, and an assessment of agricultural zone diversity

Live spiders and burrows of Kwonkan.
A Juvenile K. eboracum. B Female K. wonganensis C Juvenile K. sp. ‘MYG978’ 
 Burrow entrance morphology of D K. eboracum (open); E K. wonganensis (closed); F K. sp. ‘MYG978’(open) 
G, H Burrow entrance morphology of K. turriger, showing both free-standing (G) and foliage-supported (H) burrow entrances. I Burrow entrance morphology (open) of an undescribed Kwonkan species from near Leinster, W.A. 

in Wilson, Urso, Rix, et Harvey, 2026. 

  DOI: doi.org/10.3897/asp.84.e189518 

 Photos: A, C, F, I by J. Wilson; B, E by E. Volschenk; D by V. Cruz Bedón; G, H by M. Harvey.

Abstract

The collar-door wishbone spiders of the genus Kwonkan Main, 1983 are an Australian-endemic lineage of mygalomorph spiders that often construct elaborate burrow entrances, including collars and turrets, and remain poorly documented across their range, despite museum collections indicating high local endemism and substantial undescribed diversity. Much of the existing taxonomy, including nine of the 14 currently described species, was based on limited material and lacked modern morphological or molecular approaches to species delimitation, hindering efforts to document the remaining diversity and address conservation concerns. Here, we redescribe all nine legacy species and review Kwonkan diversity within the south-western Western Australian (SWWA) agricultural region, a highly fragmented and mostly cleared landscape harbouring extensive undescribed diversity and the threatened species K. eboracum Main, 1983. In the process, we clarify species identities, present the first molecular data for several species including K. eboracum, and describe two new species (K. elatus sp. nov. and K. yorkrakine sp. nov.) that were previously attributed to legacy species. In our review of the SWWA agricultural region fauna we identify 29 putative undescribed species and a pattern of extensive sympatry, fine-scale species turnover, and extremely restricted ranges. These findings highlight the need for continued revisionary work and potential conservation listing of additional described species such as K. wonganensis (Main, 1977).

Keywords: short-range endemism, conservation systematics, mygalomorph spiders, south-western Australian biodiversity hotspot, Wheatbelt

Live spiders and burrows of Kwonkan.
A Juvenile K. eboracum from near Kellerberrin, Western Australia (W.A.). B Female K. wonganensis from near Wongan Hills, W.A. C Juvenile K. sp. ‘MYG978’ from the Avon Wheatbelt, W.A.
D Burrow entrance morphology (open) of K. eboracum from near Kellerberrin, W.A. E Burrow entrance morphology (closed) of K. wonganensis from near Wongan Hills, W.A. F Burrow entrance morphology (open) of K. sp. ‘MYG978’ from the Avon Wheatbelt, W.A.
G, H Burrow entrance morphology of K. turriger, showing both free-standing (G) and foliage-supported (H) burrow entrances. I Burrow entrance morphology (open) of an undescribed Kwonkan species from near Leinster, W.A. 

 Photos: A, C, F, I by J. Wilson; B, E by E. Volschenk; D by V. Cruz Bedón; G, H by M. Harvey.

 Jeremy D. Wilson, Arianna Urso, Michael G. Rix, Erich S. Volschenk, Valentina Cruz Bedón and Mark S. Harvey. 2026. Wishbone Spiders of the Genus Kwonkan Main, 1983 (Araneae: Mygalomorphae: Anamidae) in south-western Australia: Redescription of legacy species, Two New Species, and an assessment of agricultural zone diversity. Arthropod Systematics & Phylogeny. 84: 510-547.  DOI: doi.org/10.3897/asp.84.e189518 [08-07-2026]

[Crustacea • 2025] Cambarus ocoeensis • A New Species of Crayfish (Decapoda: Cambaridae) from the Ocoee River Basin of Tennessee, USA

 

Cambarus ocoeensis 

 Thoma & Williams, 2025 

DOI: doi.org/10.11646/zootaxa.5618.1.7  

  Researchgate.net/publication/390392403 

  facebook.com/RogerFThoma

Abstract

Cambarus ocoeensis sp. nov. is described from the Ocoee River Gorge area of Tennessee. The species is morphologically most similar to Cambarus hiwasseensis Hobbs, 1981, but differs from C. hiwasseensis in the following ways: first pereopod chelae in general less sculpted, usually with one row of palmer tubercles, if second row present very weakly developed and consisting of six or fewer tubercles, lateral margin not costate, lateral impression weak at most but usually absent; areola length 2.5–4.1 times greater than width, having 5–8 punctations in narrowest part; size at maturity smaller with MI ranging from 21.8 to 30.9 mm total carapace length (24.6–40.0 mm in C. hiwasseensis); and antennal scale broadest at midpoint. It can be distinguished from other morphologically similar species—those with acuminate rostra—from the Hiwassee River basin of northern Georgia, southeast Tennessee, and western North Carolina area in lacking lateral rostral spines/tubercles and cervical spines. Cambarus ocoeensis sp. nov. is genetically most closely related to an undescribed member of the Cambarus bartonii (Fabricius, 1798) species complex.

 

Crustacea, C. hiwasseensis, C. bartonii, C. howardi, Hiwassee River basin, species complex

Type material of Cambarus ocoeensis sp. nov.:
 A–C, E, G–I, holotype MI (OSUMC #10897); F, allotypic paratypefemale (OSUMC #10898); D, morphotypic paratype MII (OSUMC #10899).
A–carapace, dorsal view; B–gonopod, lateralview; C–gonopod, mesial view; D–gonopod, lateral view; E–antennal scale, dorsal view; F–annulus ventralis; G–pereopods3–5, right side ventral views; H–epistome; I–right chela, dorsal view. 

Color image of Cambarus ocoeensis sp. nov., holotype (OSUMC #10897).

Image captured in lab under artificial light.

Cambarus ocoeensis sp. nov. 

Roger F. THOMA and Bronwyn W. WILLIAMS. 2025. Cambarus ocoeensis, A New Species of Crayfish (Decapoda: Cambaridae) from the Ocoee River Basin of Tennessee, USA.  Zootaxa. 5618(1); 106-118. DOI: doi.org/10.11646/zootaxa.5618.1.7 [2025-04-01] 

 Researchgate.net/publication/390392403_Cambarus_ocoeensis_a_new_species_of_crayfish_from_the_Ocoee_River_Basin_of_Tennessee_USA

  facebook.com/RogerFThoma/posts/10228373600190065

[Entomology • 2026] Scymnus tshunsii • The First termitophilous Ladybird Beetle (Coleoptera: Coccinellidae) from Microcerotermes Nests in Thailand

 

  Scymnus (Pullus) tshunsii Seki, Liang & Maruyama, 

in Seki, Liang , Hasin, Chiu et Maruyama, 2026.

DOI: doi.org/10.14411/eje.2026.017  

Termite nests host diverse symbiotic arthropods (termitophiles), many of which exhibit specialized morphological and behavioral adaptations to life within termite colonies. While numerous beetle families include well-known termitophiles, an association with termites has not yet been recorded in the family Coccinellidae (ladybird beetles). Here, we report the discovery of Scymnus (Pullus) tshunsii Seki, Liang & Maruyama, sp. n., based on adults and larvae collected from nests of Microcerotermes crassus Snyder, 1934, in Chiang Mai and Nakhon Ratchasima, Thailand, with additional adults captured at light traps. Although the adult morphology is typical for free-living Scymnini, the larva exhibits a highly unusual, soft-bodied, glabrous body form closely resembling termite workers - an apparent case of convergent evolution also seen in termitophilous rove beetles, scarabs, and phorid flies. The stark contrast between adult and larval morphology suggests stage-specific adaptations, with larvae more intimately integrated into the termite nest environment. Collecting adults at light traps indicates nocturnal dispersal, likely to locate new host colonies. This discovery represents the first confirmed termitophilous ladybird beetle and indicates that termite nests may support previously unrecognized ecological associations within Coccinellidae.

 

Keywords: New species, predators, Scymnini, taxonomy, termite

  Scymnus (Pullus) tshunsii sp. n.
1 – dorsal view; 2 – ventral view; 3 – lateral view;
4 – alive, lateral view; 5 – alive, ventral view.

Habitat of Scymnus (Pullus) tshunsii sp. n.
 32–34 – Scymnus (Pullus) tshunsii sp. n. (32 – adult; 33–34 – larvae); 35 – soldier of Microcerotermes crassus Snyder, 1934;
36 – forest environment; 37 – mound of Microcerotermes crassus Snyder, 1934.

 

Scymnus (Pullus) tshunsii Seki, Liang & Maruyama sp. n. 

 

Diagnosis. Scymnus (Pullus) tshunsii sp. n. is similar in external coloration to many species of Scymnini (e.g., Sasaji, 1971; Chen et al., 2015a, b; Wang & Chen, 2022), but can be readily distinguished by the structure of the genitalia. This species is placed in the subgenus Pullus Mulsant, 1846 based on the combination of the following characters: antennae composed of 11 antennomeres, prosternal process with well-developed lateral carinae, abdominal postcoxal lines recurved and complete, and female genitalia with a distinct infundibulum (Chen et al., 2015a). Chen et al. (l.c.) divided the subgenus Scymnus (Pullus) from mainland China and Taiwan into S. hingstoni, S. subvillosus, S. impexus, S. perdere, and S. sodalis groups. The new species lacks a prominent dorsal keel on the penis guide of ...

Etymology. The species is named after Tshun-Sî, the son of Wei-Ren Liang, the second author. His name, meaning “sowing in spring,” reflects the hope for the flourishing of future studies on termitophily in Thailand following this groundbreaking discovery. 

Distribution. Thailand (Chiang Mai, Nakhon Ratchasima)  

Ryōta SEKI, Wei-Ren LIANG, Sasitorn HASIN, Chun-I CHIU and Munetoshi MARUYAMA. 2026. The First termitophilous Ladybird Beetle: Scymnus (Pullus) tshunsii sp. n. (Coleoptera: Coccinellidae) from Microcerotermes nests in Thailand. Eur. J. Entomol. 123: 165-174. DOI: doi.org/10.14411/eje.2026.017  [May 14, 2026]

[Entomology • 2026] Montana tomorri & M. dani • Two New Species of Bush-crickets (Orthoptera: Tettigoniidae) from the Mountains of south Albania

 

(A–D) Montana tomorri and (G–H) Montana dani 

Puskás, Lemonnier-Darcemont, Willemse, Chobanov, Heller, Halimi, Darcemont, Kotitsa& Szövényi, 2026

DOI: doi.org/10.11646/zootaxa.5828.2.3

Abstract

Two new taxa of Platycleidini have been discovered in southern Albania, with one of them extending its range to the Greek border. In this paper, we analyse their morphological features, the characteristics of their acoustic signals, and phylogenetic relationships. These taxa clearly belong to the cluster of species comprising the four genera Montana, Amedegnatiana, Parnassiana, and Metrioptera s. str. The results of this study suggest their placement within the genus Montana, and we describe two new species: Montana tomorri sp. nov. and Montana dani sp. nov. Our findings highlight the urgent need for a reassessment of the tribe Platycleidini using integrative approaches that incorporate molecular and bioacoustic data, in order to better understand monophyletic groups and evaluate the consistency of current generic classifications.

Orthoptera, balkans, mountain endemism, Platycleidini, bioacoustics, phylogeny

habitus and colour variation of Montana tomorri sp. nov. from Tomorr Mt. (A–D) 

 

habitus and colour variation of M. dani sp. nov. from Trebeshina Mt. (E–H).

GELLÉRT PUSKÁS, MICHÈLE LEMONNIER-DARCEMONT, LUC WILLEMSE, DRAGAN CHOBANOV, KLAUS-GERHARD HELLER, ELTJON HALIMI, CHRISTIAN DARCEMONT, NEFELI KOTITSA, GERGELY SZÖVÉNYI. 2026. Montana tomorri and Montana dani (Orthoptera: Tettigoniidae), Two New Species of Bush-crickets from the Mountains of south Albania. Zootaxa. 5828(2); 225-255. DOI: doi.org/10.11646/zootaxa.5828.2.3 [2026-06-09]

[Chilopoda • 2026] Hainanthereua albilineata • Integrative Morphological and Molecular Evidence reveals A New Genus of scutigerid centipede (Scutigeromorpha: Scutigeridae) from Hainan, China, with implications for its evolution and biogeography

Hainanthereua albilineata   

Li, Edgecombe & Jiang, 2026

 

DOI: doi.org/10.3897/asp.84.e186815

Abstract

Hainanthereua albilineata gen. nov. et sp. nov. is described and illustrated based on specimens from Hainan Province, China. Morphological examination showed that these specimens belong to Thereuoneminae and share certain similarities with Thereuopodina Verhoeff, 1905. Phylogenetic reconstruction based on five genes (nuclear 18S and 28S rRNA, mitochondrial 12S and 16S rRNA, and cytochrome c oxidase subunit I) indicated that the specimens form a distinct and well-supported clade that is sister group to Thereuonema Verhoeff, 1904, so a new genus is accordingly established. Combining the evolutionary history and biogeographic framework of Thereuoneminae, this study revealed the evolutionary significance of Hainanthereua gen. nov. in the context of the Peninsular Indian Plate as a biotic ferry implicated in the origin of East and Southeast Asian lineages. Morphological similarities between Hainanthereua and Thereuopodina are symplesiomorphies of a clade that includes these two genera, Thereuonema and Thereuopoda Verhoeff, 1904.

Key words: Chilopoda, morphology, new species, phylogeny, taxonomic key, Thereuoneminae

Hainanthereua albilineata sp. nov., holotype.
A Habitus, dorsal view, body length 23 mm; B head, dorsal view; C T1, dorsal view; D T2, dorsal view.

Abbreviations: lmd, longitudinal median depression; ap, anterior projection of the (cephalic) transverse suture; To, Tömösváry’s organ; t, (cephalic) transverse suture; ce, compound eye.

 Hainanthereua gen. nov.

Hainanthereua albilineata gen. nov. et sp. nov. 

Habitat of Hainanthereua albilineata sp. nov. (from type locality: Mingfenggu Valley, Jianfengling, Ledong Li Autonomous County, Hainan Province, China).
A Panoramic view of type locality; B woodland on the mountain of type locality shown in A; C interior habitat of the woodland shown in B, the species was collected in relatively dry leaf litter or under stones; D individual of the species found under a stone; E live habitus of the species.

 A–D provided by Mr. Quanyu Ji, E provided by Mr. Jiazhou Lu.

Qing Li, Gregory D. Edgecombe and Chao Jiang. 2026. Integrative Morphological and Molecular Evidence reveals A New Genus of scutigerid centipede from Hainan, China, with implications for its evolution and biogeography (Scutigeromorpha: Scutigeridae). Arthropod Systematics & Phylogeny. 84: 447-464. DOI: doi.org/10.3897/asp.84.e186815

 

[PaleoEntomology • 2026] Cretolala kachinensis (Hemiptera: Fulgoromorpha: Delphacoidea) • the First lalacid from mid-Cretaceous Kachin Amber: Morphological and Taphonomic Implications

Cretolala kachinensis 

Tang, ŠMÍDOVÁ, Ding, BODERAU, Fabrikant, Mähler, Deng, Nyunt, Fu, Jouault, Tomaschek, Szwedo & Jiang, 2026

 

 DOI: doi.org/10.11646/palaeoentomology.9.2.5

Abstract 

We report Cretolala kachinensis gen. et sp. nov., the first formally described representative of the family Lalacidae from mid-Cretaceous (~99 Ma) Kachin amber. The morphology and taphonomy of the specimen were studied using optical microscopy, micro-computed tomography (micro-CT), confocal laser scanning microscopy (CLSM), and scanning electron microscopy coupled with energy-dispersive X-ray spectroscopy (SEM‒EDS). Cretolala kachinensis gen. et sp. nov. exhibits a unique combination of forewing characters, including a strengthened, broad, and wrinkled costal margin; ScP+R fork proximal to both the Pcu+A1 fusion and the CuA fork; MP with five terminals; and CuA1 and CuA2 each with two terminals. Crossvein patterns differ between the left and right forewings, indicating that crossvein expression alone is not diagnostic. The forewing bears distinct maculation, with a face-like pattern dorsally and an extensive lateral pattern that may indicate aposematism, mimicry, or plant-like camouflage in an arboreal herbivore. Taphonomic analyses based on optical micrographs and material-dependent X-ray attenuation in micro-CT show that minerals are distributed throughout the fossil, coating anatomical surfaces and infilling pre-existing cavities and voids. These phases help stabilize morphology and influence the micro-CT reconstruction of soft-bodied structures. SEM–EDS analyses indicate that the body-coating and void-filling minerals are dominated by quartz and pyrite, with feldspars and aluminosilicate mixtures also present, locally enriched in K, Na, Mg, or Ca. This assemblage and its textures point to a polyphase paragenetic sequence, with early detrital infill and later authigenic mineralization. Our results suggest that minerals and some chemicals derived from the host sediments can be transported into and, in some cases, precipitated within amber inclusions, infilling voids and providing structural support that influences fossil preservation. Collectively, these phases occlude porosity, stabilize morphology, and enhance X-ray contrast, although they locally obscured primary tissue boundaries. Our results highlight multistage diagenesis in Kachin amber and underscore variability among inclusions in both mineral assemblages and the timing of mineralization/diagenetic events.

Hemiptera, Fulgoromorpha, Delphacoidea, Mesozoic, Burmese amber, Planthopper, taxonomy, taphonomy, mineral infilling, wing maculation

Cretolala kachinensis gen. et sp. nov., holotype (HMJ–2016–019).
A–D, Reflected-light micrographs. A, Right lateral view. B, Left lateral view. C, Dorsal view. D, Ventral view. E, F, Micro-computed tomography (micro-CT) reconstructions. E, Dorsal view. F, Ventral view. Scale bars = 1 mm.

Systematic palaeontology

Order Hemiptera Linnaeus, 1758

Suborder Fulgoromorpha Evans, 1946

Superfamily Delphacoidea Leach, 1815

Family Lalacidae Hamilton, 1990

Cretolala Jiang, Boderau, Jouault & Szwedo gen. nov.

  

Etymology. The generic name is formed from the combination of “Cretaceous” and generic name “Lalax”; the latter is derived from the Greek λάλαξ (“croaker”, “caller”), in reference to a lalacid planthopper from the Cretaceous. Gender is feminine.

Diagnosis (provisional, pending confirmation from additional specimens and in combination with specieslevel diagnostic characters). Tegmen with narrow base and distinctly widened apical portion; costal margin strengthened, broad and wrinkled. Pc+CP shifted away from anterior margin near tegmen base, parallel to CA, forming a narrow, sclerotized costal area. ScP+R forks basad both CuA fork and Pcu+A1 fusion. RA with two terminals; RP with three terminals; MP with five terminals; CuA1 and CuA2 with two terminals each. 

Cretolala kachinensis Jiang, Boderau, Jouault & Szwedo sp. nov.  

Material. Holotype: HMJ–2016–019, adult female in Kachin (Burmese) amber from northern Myanmar. Repository: Myanmar Gems Museum, Nay Pyi Taw, Myanmar. Condition: the specimen is incomplete, with the head and the dorsal part of the thorax missing. 

Etymology. The specific epithet is derived from “Kachin”, the name of the state in northern Myanmar where the amber originates, and is Latinized with the adjectival suffix -ensis, meaning from “Kachin”. 

Diagnosis. As for the genus, due to monotypy. Additional features include forewing coloration and venation details: costal margin with a continuous dark band along costal area; membrane bearing nearly continuous aligned patches and short transverse bands; a discrete insular macula on ScP+RA; a sub-elliptical isolated macula near distal Pcu; distad of forks of RP and MP, pigment is vein-following, tracking adjacent longitudinal and crossveins; postnodal cell centers unpigmented, ...

 Life reconstruction of Cretolala kachinensis gen. et sp. nov.

 (illustrated by Lucia Šmídová).

Ya-Ni TANG, Lucia ŠMÍDOVÁ, Ying-Ying DING, Mathieu BODERAU, Dolev FABRIKANT, Bastian MÄHLER, Jun-Chen DENG, Thet Tin NYUNT, Yan-Zh FU, Corentin JOUAULT, Frank TOMASCHEK, Jacek SZWEDO, Hui JIANG. 2026. Cretolala kachinensis gen. et sp. nov. (Hemiptera: Fulgoromorpha: Delphacoidea), the First lalacid from mid-Cretaceous Kachin Amber: Morphological and Taphonomic implications. Palaeoentomology. 9(2); 125-138. DOI: doi.org/10.11646/palaeoentomology.9.2.5 [2026-04-29]

[Entomology • 2026] Leptogenys distincta & L. gastrolucida • Two New Species of Ant Genus Leptogenys (Hymenoptera: Formicidae) from India, with an updated key to the known Oriental species

Leptogenys distincta  
 Leptogenys gastrolucida

Baidwan, Kapoor & Bharti, 2026

 

DOI: doi.org/10.1080/00222933.2026.2657496

 

ABSTRACT

Two new species of the genus Leptogenys are described based on the worker caste, namely Leptogenys distincta sp. n. and Leptogenys gastrolucida sp. n. from India. The male of L. gastrolucida sp. n. is also reported and described. An updated dichotomous key to the 60 known species of Leptogenys from the Oriental region is provided.

 

KEYWORDS: Taxonomy, Indo-Burma, biodiversity, description

 

 Leptogenys distincta sp. n.  

Leptogenys gastrolucida sp. n.  

Parvinder Singh Baidwan, Rakeshwar Kapoor and Himender Bharti. 2026. Two New Species of Ant Genus Leptogenys (Hymenoptera: Formicidae) from India, with an updated key to the known Oriental species. Journal of Natural History. 60(25-28); 1357-1397. DOI: doi.org/10.1080/00222933.2026.2657496 [08 Jun 2026] 

[Entomology • 2026] Pseudocapritermes novus • A New Species of Soil-dwelling Termites (Blattodea: Termitidae: Mirocapritermitinae) from the Indian Subcontinent

Pseudocapritermes novus Rituparna, Baraik & Rajmohana, 

in Sengupta, Baraik, Rajmohana, Debnath, Dinesh et Chinu, 2026. 

https://jibs.modares.ac.ir 

 Researchgate.net/publication/404799169

Abstract

The present study reports a new species, Pseudocapritermes novus Rituparna, Baraik & Rajmohana sp. nov. from West Bengal, India. Species description followed an integrative taxonomic approach by including a mitochondrial 16S rRNA genetic sequence with comprehensive morphological characteristics of the soldier caste. Additionally, the first mt 16S rRNA gene sequence for the recently identified Pseudocapritermes kunjepu Mathew, 2020, is presented in the study. Digital photos of the new species are also included, along with an identification key to the soldier castes of all species of the genus reported from the Indian sub-continent. Association of Pericapritermes semarangi (Holmgren, 1913) with the new species is also recorded in the study.

 

Keywords: Inquiline, mitochondrial 16S rRNA, molecular phylogeny, Pericapritermes, Pseudocapritermes, taxonomy

Pseudocapritermes novus Rituparna, Baraik & Rajmohana sp. nov., Soldier:
A. General habitus, lateral view; B. Postmentum; C. Mandibles; D. Head capsule, dorsal view; E. Head capsule, ventral view; F. Labrum marked with an arrow (the extended process of the anterolateral corner of the labrum is broken on the right side); G. Antennae.

Pseudocapritermes novus Rituparna, Baraik & Rajmohana sp. nov. 

  

Rituparna Sengupta, Balmohan Baraik, Rajmohana K., Rupam Debnath, K. P. Dinesh and Ipe Chinu. 2026. Description of A New Species of Soil-dwelling Termites (Blattodea: Termitidae: Mirocapritermitinae) from the Indian subcontinent. The Journal of Insect Biodiversity and Systematics. DOI: 10.48311/jibs.12.03.537 [13 May 2026] 

https://jibs.modares.ac.ir/article_28644.html

 Researchgate.net/publication/404799169_a_new_species_of_Pseudocapritermes_from_the_Indian_subcontinent

[Arachnida • 2026] Trogloraptor tulishpun • A New northern Species of Trogloraptor (Araneae: Trogloraptoridae), Genetic Diversity and Natural History

 

 Trogloraptor tulishpun Jones & Binford,

in Jones, Watson, Hedin et Binford, 2026.

DOI: doi.org/10.11646/zootaxa.5828.1.5 

photos by M. Hedin

Abstract

We present a morphological description of a recently discovered species of spider in the family Trogloraptoridae from the Columbia River Gorge in northwestern Oregon. The family was previously monotypic (Trogloraptor marchingtoni) and only known from populations near the southwestern Oregon—northern California border. Trogloraptor tulishpun sp. nov. retains the key family synapomorphy, distinctive subsegmented raptorial tarsi, and an oblique membranous division of the basal segment of the anterior lateral spinnerets. Trogloraptor tulishpun is distinguished from T. marchingtoni by its color pattern, clypeal height, vulvar and palp structure. We have found T. tulishpun in four localities in the Columbia River Gorge, which show little mitochondrial sequence divergence from one another, but are highly genetically distinct from T. marchingtoni. Trogloraptor tulishpun is found in basalt features, including lava tubes and shallow talus caves, and has been observed to eat arachnids and moths, making them top predators in these environments.

Araneae, arachnophagy, caves, invertebrate conservation, mitochondrial divergence, subterranean biodiversity

Habitus of live Trogloraptor tulishpun from Herman Creek Talus Cave.
A male in captivity, B female in captivity,
C male in Herman Creek Talus Cave, D female in Herman Creek Talus Cave on sparse web,
 E male on non Trogloraptor web, F female in Herman Creek Talus Cave.

(all photos by M. Hedin)

Habitus of male Trogloraptor tulishpun (holotype, USNMENT02418340) from Herman Creek Talus Cave.
A, B dorsal views, C, E ventral views, D anterior view.

Trogloraptor tulishpun, Jones and Binford, sp. nov.  

Diagnosis. Trogloraptor tulishpun resembles T. marchingtoni (Griswold et al., 2012) by its similar carapace and abdominal shape and coloration, its subsegmented raptorial tarsi, and the partly sclerotized genital region (Figs 2, 3, 5, 7). It is clearly distinguished by its conspicuous scalloped patterning along the lateral and posterior edges of the carapace which is absent in T. marchingtoni. The abdominal patterning is different from the chevron pattern of T. marchingtoni, with oval-shaped spots down the center of the dorsal side that have a thin line of beige down the ...

Etymology. Tulishpun (pronounced too-lish-pun) is a word, simplified for ease of pronunciation, from the Sahaptin River dialect meaning “cave predator, owner of the domain”. The Sahaptin River dialect is from the Columbia River area, the type locality of this species, and the name was given to us by elders from the Columbia River Inter-Tribal Fish Commission. The specific epithet is to be treated as a noun in apposition.

 

MADELINE M. JONES, FINN WATSON, MARSHAL HEDIN and GRETA J. BINFORD. 2026. Beneath the Surface: A New northern Species of Trogloraptor (Araneae: Trogloraptoridae), Genetic Diversity and Natural History.  Zootaxa. 5828(1); 103-116. DOI: doi.org/10.11646/zootaxa.5828.1.5 [2026-06-08]

[Entomology • 2026] Kongobatha spinosistyla, K. serpens & K. rufilinea • Striking, Slender, and Secretly Spinose: A Revision of the Snake Mantises of the Genus Kongobatha (Mantodea: Nanomantidae: Fulciniinae: Neomantini)

Kongobatha spinosistyla Connors, 

in Connors, Yeeles, Lach et  Rentz, 2026

DOI: doi.org/10.11646/zootaxa.5807.1.2

 

Abstract

Kongobatha is one of the most commonly encountered of all Australian mantis genera, and yet despite this, very little is known about the taxonomy or biology of the genus. Described to include a single Australian species, K. diademata, the only subsequent work on the genus has been the description of a second species from New Guinea, K. papua. We here describe three additional species, K. spinosistyla Connors sp. nov., K. serpens Connors sp. nov., and K. rufilinea Connors sp. nov., and redescribe K. diademata and K. papua in detail, the latter of which is recorded from Australia for the first time. We also describe for the first time the unusual, heavily spinose styli of male Kongobatha. These are apparently unique among Mantodea as a whole, but their function remains unknown. 

Mantodea, mantis, snake mantis, Australia, New Guinea, taxonomy, styli, citizen science

Male and female Kongobatha spinosistyla (Rainforest Snake Mantis), Kuranda, North Queensland.

photo: Maurice Allan

MATTHEW G. CONNORS, PETER YEELES, LORI LACH and DAVID C.F. RENTZ. 2026. Striking, Slender, and Secretly Spinose: A Revision of the Snake Mantises of the Genus Kongobatha (Mantodea: Nanomantidae: Fulciniinae: Neomantini).  Zootaxa. 5807(1); 45-84. DOI: doi.org/10.11646/zootaxa.5807.1.2 [2026-05-08]

[Entomology • 2026] Varitrella sulu & V. alternata • New Species and Bioacoustics of Varitrella (Cantotrella) (Orthoptera: Ensifera: Oecanthidae: Podoscirtinae) from Sulu Archipelago and Borneo’s Sabah

 

Varitrella (Cantotrella) sulu Haibil, Nuñeza & Tan, 

in Haibil, Nuñeza, Damit, Japir, Chung et Tan, 2026.

DOI: doi.org/10.11646/zootaxa.5821.1.4 

facebook.com: Arthur CHUNG 

 

Abstract

Two new species of Varitrella (Cantotrella) (Oecanthidae: Podoscirtinae) are described from Sulu Archipelago: Varitrella (Cantotrella) sulu Haibil, Nuñeza & Tan, sp. nov. and Varitrella (Cantotrella) alternata Haibil, Nuñeza & Tan, sp. nov. The calling songs of Varitrella (Cantotrella) sulu Haibil, Nuñeza & Tan, sp. nov. and Varitrella (Cantotrella) trusmadi Gorochov, 2014 are also described.

Orthoptera, Calling songs, East Malaysia, Philippines, Podoscirtini, Southeast Asia

Varitrella (Cantotrella) sulu Haibil, Nuñeza & Tan, sp. nov. 

Varitrella (Cantotrella) alternata Haibil, Nuñeza & Tan, sp. nov. 

HELEN H. HAIBIL, OLGA MACAS NUÑEZA, DAYANG FAZRINAH BINTI AWG DAMIT, RAZY JAPIR, ARTHUR Y.C. CHUNG and MING KAI TAN. 2026. New Species and Bioacoustics of Varitrella (Cantotrella) (Oecanthidae: Podoscirtinae) from Sulu Archipelago and Borneo’s Sabah.  Zootaxa. 5821(1); 61-77. DOI: doi.org/10.11646/zootaxa.5821.1.4 [2026-05-28]

 facebook.com/ArthurCHUNG123799/posts/3545681898914705

[PaleoEntomology • 2026] Qiongqi multispurous • A New Cricket (Orthoptera: Trigonidiidae) from mid-Cretaceous Kachin Amber in Northern Myanmar

Qiongqi multispurous 

Ji, Nel, Xiao & Xu, 2026

DOI: doi.org/10.54103/2039-4942/29769

Abstract

A new species of Trigonidiidae, †Qiongqi multispurous sp. nov., is described from mid-Cretaceous Kachin amber. It can be definitely attributed to the family Trigonidiidae, but excluded from the two extant subfamilies. It also exhibits a different morphology from all reported trigonidiids genera and species from the Mesozoic, including subapical/apical spurs on the hind tibia and setae-like processes on the hind basitarsomere. This new discovery provides novel morphological information of Cretaceous Trigonidiidae, and highlights the potential of biodiversity of Cretaceous crickets.

Keywords: Trigonidiidae; mid-Cretaceous; new species; fossil; morphology

Qiongqi multispurous sp. nov., holotype NIGP210185.
A) Photograph of  habitus in dorsal view. B) Drawing of  habitus in dorsal view.
C) Drawing of  habitus in ventral view. D) Photograph of  habitus in ventral view.

Scale bars = 1 mm.

Qiongqi multispurous sp. nov.

XIA JI, ANDRÉ NEL, CHUANTAO XIAO and CHUNPENG XU. 2026. A NEW CRICKET (ORTHOPTERA: TRIGONIDIIDAE) FROM MID-CRETACEOUS KACHIN AMBER IN NORTHERN MYANMAR. RIVISTA ITALIANA DI PALEONTOLOGIA E STRATIGRAFIA. 132(1); DOI: doi.org/10.54103/2039-4942/29769 [06-03-2026]