[Mollusca • 2023] Siamochlamys panhai • A New Limestone Karst‐Restricted Terrestrial Snail (Eupulmonata: Helicarionidae) from Thailand

Siamochlamys panhai Pholyotha, 

Pholyotha, Tongkerd et Sutcharit, 2023. 

DOI: doi.org/10.58837/tnh.23.7.258814 

Abstract

A new genus and new species of helicarionid snail, Siamochlamys panhai, is described from limestone outcrops in Thailand. This new genus differs from all helicarionid genera based on morphological characters of shell, radula, and genitalia. The diagnostic characters of Siamochlamys panhai gen. et sp. nov. are depressed shell, well-polished surface, umbilicus opened, and aperture less covered by reflection of columellar margin. The genitalia exhibit a straight epiphallic caecum and small flagellum but lack a dart apparatus; radula has tricuspid central and lateral teeth and bicuspid marginal teeth. The discovery of this karst-associated land snail increases the knowledge of endemic malacofauna in Thailand, and also suggests that many taxa remain undiscovered.

Keywords: endemic, Indochina, land snail, limestones, taxonomy

Superfamily Helicarionoidea Bourguignat, 1877

Family Helicarionidae Bourguignat, 1877

Subfamily Durgellinae Godwin-Austen, 1888

Living snails and mantle extensions of Siamochlamys panhai gen. et sp. nov. paratype CUMZ14211.
A. living snail during relaxed position showing the expansion of mantles B. alcohol-preserved snailshowingshell lobes and dorsal lobes and posterior body C. dorsal view of mantle extensions (shell lobes and dorsal lobes).

Genus Siamochlamys Pholyotha gen. nov.

Etymology.− The name combines ‘Siam’,which is a former name of Thailand,chosen because this snail is restricted to the country as far as we know, and the Greek word ‘chlamys’,meaning “mantle or cloak”. Therefore, the generic name means the land snail with well-developed mantle extensions from Thailand. 

Diagnosis.− Depressed, glossy shellwith open umbilicus, straight epiphallic caecum, short flagellum, no dart apparatus, radula with tricuspidcentral and lateral teeth and bicuspid marginal teeth, andfive mantle extensions well-developed.

Siamochlamys panhai Pholyotha, sp. nov.

Arthit Pholyotha, Piyoros Tongkerd, Chirasak Sutcharit. 2023. Siamochlamys panhai, A New Limestone Karst‐Restricted Terrestrial Snail from Thailand (Eupulmonata: Helicarionidae). Tropical Natural History. Supplement 7;41-50. DOI: doi.org/10.58837/tnh.23.7.258814 

[Mollusca • 2026] Pagodapalaina euphrosyne • A New open-coiled Microsnail Species of the Genus Pagodapalaina (Caenogastropoda: Diplommatinidae) from Thailand

Pagodapalaina euphrosyne Sutcharit,

 

in Sutcharit, Srisonchai, Maneewan, Jeratthitikul et Annate, 2026. 

หอยกระสวยจิ๋วคลายเกลียวยูโฟรซีน  |  DOI: doi.org/10.1556/1777.2026.00169  

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Abstract

A new species of Pagodapalaina from southern Thailand is described. Pagodapalaina euphrosyne Sutcharit, sp. nov. is characterised by a sinistral, elongate-fusiform shell with an open-coiled teleoconch, well-developed radial ribs restricted to the penultimate and last whorls, and the presence of the parietalis and columellaris. Unlike all other congeners, the detached teleoconch subsequently resumes regular coiling, with the last whorl becoming tightly reattached to the preceding whorl. This previously undocumented developmental pattern raises fundamental questions about current understanding of the developmental constraints governing shell coiling in terrestrial caenogastropods.

Keywords: limestone; endemic species; whorl detachment; ecophenotypic plasticity; micro-CT

A, approximate locations of the type locality of all Pagodapalaina species.
Numbers correspond to: 1) P. acme (Laidlaw, 1949) [= Diplommatina laemsonensis Boonmachai, Bergey & Nantarat, 2023]; 2) P. attenuata (Laidlaw, 1949); 3) P. carinaspinosa (Maassen, 2002); 4) P. diminuta (von Möllendorff, 1891); 5) Pagodapalaina euphrosyne sp. nov.; 6) P. gadutensis (Maassen, 2002); 7) P. hidagai (Panha, 1998a); 8) P. hortulanica Chen, 2025; 9) P. jirasaki (Panha, Kanchanasaka & Burch, 2002); 10) P. karoensis (Maassen, 2002); 11) P. krabiensis (Panha & Burch, 1998); 12) P. lenggongensis (Tomlin, 1941) [= Diplommatina lenggongensis nawawii Davison, 1995]; 13) P. parabates (Laidlaw, 1949); 14) P. pongrati (Panha, Kanchanasaka & Burch, 2002); 15) P. prakayangensis (Panha, 1998b); 16) P. sinistra (Tomlin, 1938) [= Diplommatina prakaiphetensis Boonmachai, Bergey & Nantarat, 2023]; 17.1) P. superba superba (Godwin-Austen & Nevill, 1879); 17.2) P. superba brevior (Laidlaw, 1949); 18) P. suratensis (Panha & Burch, 1998);
B, live snail of  Pagodapalaina euphrosyne sp. nov.

Systematics

Superfamily Cyclophoroidea Gray, 1847

Family Diplommatinidae Pfeiffer, 1856

Genus Pagodapalaina Chen, 2025

Pagodapalaina Chen in Chen et al. 2025: 390, 391.

Type species: Diplommatina acme Laidlaw, 1949, by original designation.

Remark: The genus is characterised by a sinistrally coiled, elongate-fusiform shell with tightly to loosely coiled whorls and a shell surface bearing low ridges to prominent and trough-like radial ribs. The internal lamellar system comprises a well-developed columellaris, a parietalis varying from small to well developed, a vertical palatalis that may be absent, and a constriction ranging from weak to strong.

The genus currently comprises 18 recognised species (Chen et al. 2025, MolluscaBase 2026, this study). Of these, three species are known from Sumatra, Indonesia, and nine species, including the new species described herein, are recorded from Thailand. The northern distributional limit of the genus is currently considered not to extend beyond the Isthmus of Kra (Fig. 1A).

Pagodapalaina euphrosyne sp. nov., paratypes CUMZ 14506 from the type locality.
A, B, adult specimens with narrowly expanded lip;
C, juvenile specimen shows an early stage of penultimate whorl, and onset of tall trough-like radial ribs

Pagodapalaina euphrosyne sp. nov. from the type locality.
A, holotype CUMZ 14505, insets show the aperture with columellaris;
B, paratype CUMZ 14506, juvenile with penultimate and last whorl reattached but lip not yet expanded;
C, paratype CUMZ 14506, insets show the protoconch sculpture

Pagodapalaina euphrosyne Sutcharit, sp. nov.

Diagnosis: Shell small, sinistral, elongate-conical, fusiform, with ∼6 whorls. Earlier teleoconch whorls solute for about two whorls, with smooth surface. Penultimate and last whorls regularly coiled, tightly attached, sculptured with very tall, trough-like radial ribs, and fine spiral striations throughout. Peristome double, expanded. Aperture rounded, with four barriers: one parietalis, two vertical palatalis and one columellaris.

Etymology: The specific epithet, euphrosyne, is a noun in apposition derived from Euphrosyne, one of the Three Graces (Charites) in Greek mythology, the personification of joy, beauty, and elegance. It refers to the graceful and delicate shell morphology of the new species.

  Chirasak Sutcharit, Ruttapon Srisonchai, Nattaphong Maneewan, Ekgachai Jeratthitikul and Supunya Annate. 2026. A New open-coiled Microsnail Species of the Genus Pagodapalaina (Caenogastropoda: Diplommatinidae) from Thailand. Animal Taxonomy and Ecology. DOI: doi.org/10.1556/1777.2026.00169 [16 Sep 2026]

𝙋𝙖𝙜𝙤𝙙𝙖𝙥𝙖𝙡𝙖𝙞𝙣𝙖 𝙚𝙪𝙥𝙝𝙧𝙤𝙨𝙮𝙣𝙚 Sutcharit, 2026 หรือ หอยกระสวยจิ๋วคลายเกลียวยูโฟรซีน ค้นพบที่ถ้ำทะลุ อำเภอละงู จังหวัดสตูล โดยตั้งชื่อชนิดตามรูปร่างเปลือกที่ละเอียดอ่อนและสง่างาม เสมือน “ยูโฟรซีน” ผู้เป็น 1ใน 3เทพีแห่งความงามของเทพปกรณัมกรีก 

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[Crustacea • 2026] Caridina srivijaya, C. takkola, C. tambralinga, ... • A Taxonomic Review of the freshwater shrimp Genus Caridina H. Milne Edwards, 1837 (Decapoda: Atyidae) from southern Thailand, with Descriptions of Five New Species

A. Caridina cf. rangoona, from Phang Nga Province, Takuatung District, Khum Mut Stream; B. C. johnsoni, from Surat Thani Province, Mueang District, Phunphin Stream; 

C, D. C. nangroma sp. nov. from Chumphon Province, Lamae District, Lamae Stream;

E, F. C. srivijaya sp. nov., from Surat Thani Province, Phrasaeng District, Bang Sawan Spring; 

G. C. takkola sp. nov., from Surat Thani Province, Mueang District, Phunphin Stream; H. C. tambralinga sp. nov., from Songkhla Province, Ranot District, Thale Noi NHA. 

Macharoenboon, von Rintelen, Klotz, Sutcharit, Siriwut & Jeratthitikul, 2026

DOI: doi.org/10.3897/zookeys.1292.193168

Abstract

A comprehensive taxonomic review of the genus Caridina in southern Thailand is presented, integrating extensive field surveys with comprehensive morphological and molecular analyses to document the species diversity. Phylogenetic tree reconstructed from partial sequences of two mitochondrial genes (16S rRNA and COI) indicated 18 distinct lineages, recognized as species in our study, belonging to six species groups, from southern Thailand. Five of these are new to science and are described as Caridina kuehnei sp. nov., C. nangroma sp. nov., C. srivijaya sp. nov., C. takkola sp. nov., and C. tambralinga sp. nov. Including historical records, there are 21 species of Caridina found in southern Thailand of which almost half are endemic to the region. Updated distribution maps and a key to all known species in the region are also provided. Our findings highlight that while morphological identification is challenging due to significant intraspecific variation and interspecific similarity, molecular data is essential for helping taxonomists to delineate species boundaries.

Key words: Amphidromous, crustacean, freshwater fauna, landlocked, Malay Peninsula, molecular phylogeny, Thailand

Living specimens of Caridina species.
 A. C. thai, topotype MUMNH-CAR861-F1 from Nakhon Ratchasima Province, Pak Chong District, Muak Lek Waterfall; B, C. C. gracillima, specimens; B. MUMNH-CAR626-P1 from Trang Province, Mueang District, Trang River; C. MUMNH-CAR631-P1 from Krabi Province, Mueang District, Tha Hindarn Stream; D. C. gracilipes, specimen MUMNH-CAR462-P1 from Nakhon Si Thammarat Province, Khanom District, Khao Phlai Dam;
E. C. macrophora, specimen MUMNH-CAR800 from Nakhon Si Thammarat Province, Bang Khan District, Lam Dang Stream; F. C. peninsularis, specimen MUMNH-CAR633-P1 from Phang Nga Province, Khum Mut Stream; G. C. excavatoides, specimen MUMNH-CAR616-F2 from Nakhon Sri Thammarat Province, Thung Song District, Tha-Luang Stream; H. C. temasek, specimen MUMNH-CAR835-P1 from Pattani Province, Mueang District, Pattani River. Scale bar: 4 mm.

Living specimens of Caridina species.
A. C. cf. rangoona, specimen MUMNH-CAR434-P1 from Phang Nga Province, Takuatung District, Khum Mut Stream; B. C. johnsoni, specimen MUMNH-CAR471-P1 from Surat Thani Province, Mueang District, Phunphin Stream; C, D. C. nangroma sp. nov.; C. paratype MUMNH-CAR404-P1 and D. specimen MUMNHCAR474-P1 from Chumphon Province, Lamae District, Lamae Stream;
E, F. C. srivijaya sp. nov., paratypes; E. MUMNH-CAR502-P1; F. MUMNH-CAR502-M1; G. C. takkola sp. nov., paratype MUMNH-CAR469-P1; H. C. tambralinga sp. nov., specimen MUMNH-CAR489-P1 from Songkhla Province, Ranot District, Thale Noi Non-Hunting Area. Scale bar: 3 mm.

Caridina kuehnei sp. nov. 

C. nangroma sp. nov. 

C. srivijaya sp. nov. 

C. takkola sp. nov. 

C. tambralinga sp. nov. 

 

 Kongkit Macharoenboon, Thomas von Rintelen, Werner Klotz, Chirasak Sutcharit, Warut Siriwut and Ekgachai Jeratthitikul. 2026. A Taxonomic Review of the freshwater shrimp Genus Caridina H. Milne Edwards, 1837 (Decapoda, Atyidae) from southern Thailand, with Descriptions of Five New Species. ZooKeys. 1292: 1-126. DOI: doi.org/10.3897/zookeys.1292.193168 [15 Sep 2026]

[Mollusca • 2025] Songkhlanaia songkhramensis • An integrative approach to A Revision of the freshwater mussel Genus Songkhlanaia (Bivalvia: Unionidae), with the Description of A New Species

 Songkhlanaia songkhramensis sp. nov.

Jeratthitikul, Sutcharit & Prasankok, 2025

S. tamodienica, Songkhla Lake Basin, Thailand; 
 S. moreleti, Tonle Sap Basin, Thailand  

DOI: doi.org/10.3897/zookeys.1224.140549  

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Abstract

Mainland Southeast Asia, also known as Indochina, is recognized as a global biodiversity hotspot for freshwater mussels, hosting a significant number of species and exhibiting high levels of endemism. Recently, the monotypic genus Songkhlanaia was described from the Songkhla Lake Basin in southern Thailand. In this study, two additional lineages are revealed, S. moreleti comb. nov. and S. songkhramensis sp. nov., from the Mekong Basin through an integrative taxonomic approach combining morphological characters and molecular phylogenetic analyses. The multi-locus phylogenetic inference supported the monophyly of the genus and further elucidated the sister relationship between S. moreleti and the new species, and with S. tamodienica positioned as a basal lineage. Pairwise uncorrected COI p-distances among these three species also supported the species validity and ranged from 4.2% to 8.24%. Notably, S. songkhramensis sp. nov. and S. moreleti exhibit similarities in shell morphology; however, the new species can be differentiated by more robust pseudocardinal teeth. Both species are distinguishable from S. tamodienica by their approximately twice larger size, more inflated shells, and more prominent, roughened, irregular growth lines on the shell surface. Furthermore, based on the current data, these three species are recognized as endemic and are restricted to disjunct biogeographic areas in Indochina: S. tamodienica in the Songkhla Lake Basin in southern Thailand, S. moreleti in the Tonle Sap and Lower Mekong basins, and S. songkhramensis sp. nov. in the Songkhram Basin and its nearby tributaries of the Middle Mekong Basin.

Key words: Freshwater mussels, Indochina, Mekong Basin, multi-locus phylogeny, new taxa, Pseudodontini, taxonomic revision, Thailand

Family Unionidae Rafinesque, 1820

Subfamily Gonideinae Ortmann, 1916

Tribe Pseudodontini Frierson, 1927

Subtribe Pseudodontina Frierson, 1927

Genus Songkhlanaia Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov, 2023

Songkhlanaia Konopleva et al., 2023: 13, 14. Bolotov et al. 2023: 12.

Type species: Songkhlanaia tamodienica Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al. 2023 (by original designation).

Species included: Songkhlanaia currently consists of three species: S. tamodienica (type species), S. moreleti comb. nov., and S. songkhramensis sp. nov.

Shells of Songkhlanaia songkhramensis sp. nov. 
A holotype MUMNH-UNI3024 and B paratype MUMNH-UNI3023 from Prahang River, Songkhram Basin, Thailand C specimen MUMNH-UNI0925 from Songkhram River, Songkhram Basin, Thailand.

Pseudocardinal teeth, left valve on the left-hand side, and right valve on the right-hand side
A S. tamodienica, topotype MUMNH-UNI2956 from Tamot Stream, Songkhla Lake Basin, Thailand B S. moreleti, specimen MUMNH-UNI0301 from Phrom Hot Stream, Tonle Sap Basin, Thailand C S. songkhramensis sp. nov., holotype MUMNH-UNI3024 from Prahang River, Songkhram Basin, Thailand. Scale bars: 10 mm.

 Ekgachai Jeratthitikul, Chirasak Sutcharit and Pongpun Prasankok. 2025. An integrative approach to A Revision of the freshwater mussel Genus Songkhlanaia (Bivalvia, Unionidae), with the Description of A New Species. ZooKeys. 1224: 187-209. DOI: doi.org/10.3897/zookeys.1224.140549

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[Botany • 2026] Didiclis pseudobisulcata (Selaginellaceae) • A New Species of the Bisulcata group from northern Thailand and southwest China

  

Didiclis pseudobisulcata  X.M.Zhou & Liang Zhang,

in Gong, Hu, Pollawatn, Chantanaorrapint, Wang, Zhang et Zhou. 2026. 

 DOI: doi.org/10.3897/phytokeys.278.204624

Abstract

Didiclis pseudobisulcata (Lycopodioidoideae, Selaginellaceae), a new lycophyte species from northern Thailand and southwest China, is described and illustrated. Morphologically, it is similar to D. bisulcata but can be reliably distinguished by its dorsal leaves on branches with an arista often more than 3/4 of the leaf length, the densely ciliolate base of the dorsal sporophyll pteryx, and megaspores with subreticulate exospore ornamentation. Molecular phylogenetic analysis based on ITS and rbcL sequences supports this species as an independent clade. The ITS sequences revealed more than 42 nucleotide differences between the new species and other members of the Bisulcata group, providing clear molecular evidence for its separation from its closest relatives.

Key words: Asia, Lycopodioidoideae, megaspore and microspore, phylogeny

Field photographs of Didiclis pseudobisulcata.
 A, B. Plant in its natural habitat; C. Dorsal view of stems and branches with strobili; D. Ventral view of stems and branches with strobili; E. Dorsal view of strobilus; F. Ventral view of strobilus; G. Dorsal view of a portion of the main stem, showing ventral and dorsal leaves; H. Dorsal view of a portion of a branch, showing ventral and dorsal leaves; I. Ventral view of a portion of the main stem, showing axillary and ventral leaves; J. Ventral view of a portion of a branch, showing axillary and ventral leaves.

(from voucher: Liang Zhang, Xin-Mao Zhou, Lu-Yao Jiang, & Xu-Ke Han 5734).

Didiclis pseudobisulcata X.M.Zhou & Liang Zhang, sp. nov.

 Sheng-Rong Gong, Yu-Ping Hu, Rossarin Pollawatn, Sahut Chantanaorrapint, Wen Wang, Liang Zhang and Xin-Mao Zhou. 2026. Didiclis pseudobisulcata (Selaginellaceae), A New Species of the Bisulcata group from northern Thailand and southwest China. PhytoKeys. 278: 233-246. DOI: doi.org/10.3897/phytokeys.278.204624

[Botany • 2026] Thismia daemona (Thismiaceae) • A New Species of Thismia sect. Geomitra from Thailand supported by morphological and molecular evidence

 

Thismia daemona Seelanan & Chantanaorr.,

in Seelanan, Chantapram, Taosiri, Chuchuea et Chantanaorrapint, 2026.

 DOI: doi.org/10.3897/phytokeys.278.202868

Abstract

A new species of Thismia from Thailand, Thismia daemona is described based on morphological characters and molecular phylogenetic analyses. Morphological characters were examined from fresh and spirit materials using a stereo microscope. Phylogenetic analyses were conducted based on three nuclear (SSU, ITS, LSU) and two mitochondrial (atpA, matR) markers. The new species is characterized by coralliform roots, blackish flowers, reddish orange and fleshy outer tepals, inner tepals forming a mitre with three slender claviform appendages, inner surface of floral tube possessing irregular transverse bars in lower half and finely reticulations in upper half, epidermis of transverse bars and reticulations covered with white translucent trichomes, and translucent blue stamens with recurved apex. Phylogenetic analyses support its placement within Thismia sect. Geomitra, with a sister relationship to T. betung-kerihunensis. In addition, the new species is assessed as Critically Endangered according to the IUCN Red List categories and criteria.

Key words: Achlorophyllous plants, conservation status, coralliform root, endemic species

Thismia daemona Seelanan & Chantanaorr.
A. Underground part showing hairy roots; B. Above-ground shoots with flowers; C. Top view of mitre showing irregular papillose surface; D. Side view of flower showing outer lobe; E. Longitudinal section of floral tube showing inner surface, three stamens, stylar column and styles; F. Epidermal cells of transversal bar; G. Stamens outer view. H. Stylar column and styles.

Photographed by N. Taosiri (B, C, E, G, H) and C. Chuchuea (A, D, F).

Locality and habitats of Thismia daemona Seelanan & Chantanaorr.
A. Distribution map; B–D. Habitats; arrowheads indicate plants of T. daemona; E. Plants in natural habitat. Photographed by C. Chuchuea (B–D) and N. Taosiri (E).

Thismia daemona Seelanan & Chantanaorr., sp. nov.

Diagnosis. Thismia daemona is somewhat similar morphologically to T. limkokthayi, but differs in having reddish orange and fleshy outer tepals, outer surface of mitre covered with irregular papillae, inner surface of floral tube possessing irregular transverse bars in lower half and finely reticulations in upper half, epidermis of transverse bars and reticulations covered with translucent white trichomes, translucent blue stamens, and absence of mitre foveae.

Habitat and ecology. Thismia daemona was found growing amongst leaf litter, under shade of lower montane forest with small bamboo, at elevations 950–970 m.

 Tosak Seelanan, Isareyakorn Chantapram, Neeranuch Taosiri, Chanachon Chuchuea and Sahut Chantanaorrapint. 2026. Thismia daemona (Thismiaceae), A New Species from Thailand supported by morphological and molecular evidence. PhytoKeys. 278: 177-189. DOI: doi.org/10.3897/phytokeys.278.202868 [17-08-2026]

[Diplopoda • 2026] Enghoffosoma chiangmaiense, E. minutum, E. chanabunae • Integrative Taxonomy of the Millipede Genus Enghoffosoma Golovatch, 1993 (Polydesmida: Paradoxosomatidae) in Thailand and Myanmar, with Descriptions of Three New Species

Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn; 
E. minutum Srisonchai & Likhitrakarn; 
 E. chanabunae Srisonchai & Likhitrakarn, 
 

 in Srisonchai, Benchapong, Golovatch, Sutcharit, Panha et Likhitrakarn, 2026. 

DOI: doi.org/10.37828/em.2026.99.4 

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Abstract

Although comprehensive studies on the Southeast Asian flat-backed millipede genus Enghoffosoma Golovatch, 1993 in Thailand have begun quite recently, in 2014, continuous field surveys have since revealed a steadily increasing species diversity. Based on extensive new material collected from Thailand and Myanmar, three new species are described using an integrative taxonomic approach that combines morphological characters and mitochondrial COI DNA barcoding: Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn sp. n. from northern Thailand, Enghoffosoma minutum Srisonchai & Likhitrakarn sp. n. from northeastern Thailand, and Enghoffosoma chanabunae Srisonchai & Likhitrakarn sp. n. from the Tanintharyi Region, southern Myanmar. COI sequence comparisons, available for E. chanabunae Srisonchai & Likhitrakarn sp. n. and E. chiangmaiense Srisonchai & Likhitrakarn sp. n., reveal significant interspecific genetic divergences (K2P distances) from their congeners, ranging from 13.53–16.75% and 13.53–18.14%, respectively. While DNA data were unavailable for E. minutum Srisonchai & Likhitrakarn sp. n., its distinctive morphological characters, particularly the gonopod structure, clearly support its status as a new species. To provide a clear morphological baseline for comparison, the type species, Enghoffosoma spinipleurum (Carl, 1941), from Myanmar, originally described as Sundanina spinipleura Carl, 1941, is redescribed and illustrated in detail based on the lectotype. With these additions, the genus Enghoffosoma now comprises 21 valid species, confirming its status as a highly diverse and widespread diplopod lineage in the Indochinese region. An updated identification key to all known species and a distribution map are provided.

Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn sp. n. 

 Enghoffosoma minutum Srisonchai & Likhitrakarn sp. n. 

 Enghoffosoma chanabunae Srisonchai & Likhitrakarn sp. n. 

Ruttapon Srisonchai, Theemaporn Benchapong, Sergei I. Golovatch, Chirasak Sutcharit, Somsak Panha, Natdanai Likhitrakarn. 2026. Integrative Taxonomy of the Millipede Genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae) in Thailand and Myanmar, with Descriptions of Three New Species. Ecologica Montenegrina. 99; 40-71. DOI: doi.org/10.37828/em.2026.99.4 [2026-06-24] 

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[Botany • 2026] Asplenium paucipinnatum (Aspleniaceae) • A New fern Species from southern Thailand, based on morphological and molecular data

Asplenium paucipinnatum  K.W.Xu, Li Bing Zhang & Pollawatn, 

in Xu, Pollawatn, L. Zhang, Zhou et L.-B. Zhang, 2026.   

DOI: doi.org/10.3897/phytokeys.272.173645

Abstract

Asplenium paucipinnatum, a new fern species from Nakhon Si Thammarat, Thailand, is described. Morphologically, it closely resembles A. micantifrons in having short-creeping and radial rhizome steles, 1-pinnate laminae, deeply incised pinnae on the basiscopic margin, and sori arranged in two rows close to the costae. However, A. paucipinnatum is readily distinguished by having fewer lateral pinnae (< 14 pairs), obliquely rhombic pinnae with acute apices and obtuse marginal teeth, contrasting the more numerous pinnae (> 15 pairs), lanceolate pinnae with acuminate apices, and acute teeth in A. micantifrons. Phylogenetic analyses based on multiple plastid DNA regions (atpB, rbcL, rps4, rps4-trnS, trnL, and trnL-F) support A. paucipinnatum as a distinct lineage, forming a sister relationship with a clade containing A. lepturus and A. contiguum. The combination of morphological and molecular evidence supports the recognition of this new taxon. Ecological notes, geographic distribution, and a comparison with related species are also provided.

Key words: Asplenium aethiopicum clade, fern phylogeny, Southeast Asian fern flora, taxonomy

Asplenium paucipinnatum. A. Habit; B. Abaxial view of lamina; C. Abaxial view of pinnae showing the venation on the pinnae; D. Showing the lamina apices deeply pinnatifid and becoming gradually decrescent upwards; E. Adaxial view of Pinnae at middle portion of the laminae; F. Short-creeping rhizome; G. Rhizome scales.

(Voucher specimen: Li Bing Zhang, Liang Zhang, R. Pollawatn & X. M. Zhou 10976).

Asplenium paucipinnatum K.W.Xu, Li Bing Zhang & Pollawatn, sp. nov.

 

Description. Asplenium paucipinnatum resembles A. micantifrons (Tuyama) Tuyama ex H.Ohba by its short creeping and radial rhizomes, 1-pinnate laminae, pinnae deeply incised at basiscopic side of margin, and sori borne in 2 rows closely set along to the costa, but the former has lateral pinnae fewer than 14 pairs, pinnae oblique rhombic, pinna apex acute, and marginal teeth obtuse. In contrast, A. micantifrons has lateral pinnae more than 15 pairs, pinnae lanceolate, pinna apex acuminate, and marginal teeth acute.

Ke-Wang Xu, Rossarin Pollawatn, Liang Zhang, Xin-Mao Zhou and Li-Bing Zhang. 2026. Asplenium paucipinnatum (Aspleniaceae), A New fern Species from southern Thailand, based on morphological and molecular data. PhytoKeys. 272: 159-167. DOI: doi.org/10.3897/phytokeys.272.173645 [02 Apr 2026]

[Mollusca • 2025] Chamalycaeus aduncus • The First Record of an Operculate Micro Land Snail from the Alycaeinae (Caenogastropoda: Cyclophoridae) in Cambodia

 

Chamalycaeus aduncus Jirapatrasilp,   

in Jirapatrasilp, Thi, Chantha, Chourn, Sophea, Jeratthitikul, Pholyotha, Siriwut, Srisonchai, Panha et Sutcharit, 2025. 

www.ruthenica.net/node/6016

 Researchgate.net/publication/390396808 

 facebook.com/ASRUChula

 

This study reports the first record of the operculate micro land snail from the Alycaeinae, Chamalycaeus aduncus Jirapatrasilp sp. nov., also the first new species of the genus in Cambodia, based on a recent survey in Battambang. This new species is similar to Chamalycaeus excisus (Möllendorff, 1887) and C. excisus sublimus Páll-Gergely et Auffenberg, 2019 from the Philippines, in shell size and shape, and the strongly downwards coiling of R3, but differs in having a shorter R2 without any blunt keel, a round aperture without any incision, and a beak- or hook-like basal edge of the outer peristome. 

Keywords: Battambang; Cyclophoroidea; Gastropoda; Mollusca; taxonomy

Chamalycaeus aduncus sp. nov.
A paratype CUMZ 14462/1, and B paratype CUMZ 14462/2, showing a scraping hole at the umbilical side.

. Representative SEM images of Chamalycaeus aduncus sp. nov., holotype CUMZ 14461
 A whole shell in different positions, and B Close-up images of the aperture; white arrows pointing to the beak- or hook-like basal edge of the outer peristome.

Chamalycaeus aduncus Jirapatrasilp sp. nov.

 

Parin JIRAPATRASILP, Sothearen THI, Nasak CHANTHA, Phyroum CHOURN, Chhin SOPHEA, Ekgachai JERATTHITIKUL, Arthit PHOLYOTHA, Warut SIRIWUT, Ruttapon SRISONCHAI, Somsak PANHA and Chirasak SUTCHARIT. 2025. The First Record of an Operculate Micro Land Snail from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Cambodia, with Description of A New Species of Chamalycaeus von Möllendorff, 1897. Ruthenica Russian Malacological Journal. 35(2); 65-72. https://www.ruthenica.net/node/6016

 Researchgate.net/publication/390396808_The_first_record_of_an_operculate_micro_land_snail_from_the_Alycaeinae_Cyclophoridae_in_Cambodia

 facebook.com/ASRUChula/posts/1081408774003851

[Diplopoda • 2026] Rhopalomeris dulcia, R. lentiginosa, R. verhoeffi, ... • Five New Species of the Pill Millipede Genus Rhopalomeris Verhoeff, 1906 (Glomerida: Glomeridae) from Thailand

Rhopalomeris dulcia, 
R. verhoeffi  

Sapparojpattana & Likhitrakarn,

in Sapparojpattana, Jeratthitikul, Siriwut, Srisonchai, Wesener, Sutcharit et Likhitrakarn, 2026.  

DOI: doi.org/10.1163/18759866-bja10090 

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Abstract

An integrative approach based on morphological characteristics and DNA data from the mitochondrial COI gene has revealed five new species of the pill millipede genus Rhopalomeris Verhoeff, 1906 from Thailand: Rhopalomeris muka Sapparojpattana & Likhitrakarn, sp. nov., R. lentiginosa Sapparojpattana & Likhitrakarn, sp. nov., R. dulcia Sapparojpattana & Likhitrakarn, sp. nov., R. punctata Sapparojpattana & Likhitrakarn, sp. nov., and R. verhoeffi Sapparojpattana & Likhitrakarn, sp. nov. The five new species are distinguishable from their congeners by distinctive color patterns and telopod structure. The interspecific genetic divergences between the new species and other Rhopalomeris species ranged from 7.86 to 13.71%. Intraspecific genetic divergences within these five species ranged from 0 to 4.53%, with the highest divergences found in R. carnifex and R. verhoeffi sp. nov. This is consistent with both the wide distribution range and the significant intraspecific morphological variations observed in these latter two species. Furthermore, the taxonomic scope of R. carnifex in Thailand is reviewed, and intraspecific morphological variations within R. carnifex and R. verhoeffi sp. nov. are discussed. A distribution map, morphological illustrations from SEM, and an updated key to all known Rhopalomeris species are also provided.

Keywords: DNA barcoding; Indochina; integrative taxonomy; phylogeny; systematics

Rhopalomeris dulcia sp. nov., (A, B) paratype (MUMNH-GLO188), habitus, live coloration (C–E) ♀ paratype (MUMNH-GLO188), alcohol preserved, in dorsal, ventral, and lateral views (A) unrolled in sublateral view (B) rolled in sublateral view.

(A, B) image not to scale (C–E) scale bars: 2 mm

Rhopalomeris verhoeffi sp. nov., (A, B) paratype (MUMNH-GLO201), habitus, live coloration of color morph 2 (C–E) ♀ paratype (MUMNH-GLO201), alcohol preserved specimen of color morph 2 in dorsal, ventral, and lateral views (A) unrolled in sublateral view (B) rolled in sublateral view.

(A, B) image not to scale (C–E) scale bars: 2 mm

 Rhopalomeris muka Sapparojpattana & Likhitrakarn, sp. nov.

R. lentiginosa Sapparojpattana & Likhitrakarn, sp. nov. 

 R. dulcia Sapparojpattana & Likhitrakarn, sp. nov. 

 R. punctata Sapparojpattana & Likhitrakarn, sp. nov. 

 R. verhoeffi Sapparojpattana & Likhitrakarn, sp. nov. 

Distributions of currently known Rhopalomeris species in Thailand and Myanmar

(A) Bayesian inference tree focusing on the detailed topology of Rhopalomeris carnifex (Pocock, 1889) and R. verhoeffi sp. nov., with colored drawings illustrating the color variations within each species. All millipede drawings are not to scale.
(B, C) Maps of southern Thailand showing the distributions of color morphs for (B) R. carnifex and (C) R. verhoeffi sp. nov. The different shaded colors in each pie chart represent the proportion of the corresponding color morphs in that locality.

 Pichsinee Sapparojpattana, Ekgachai Jeratthitikul, Warut Siriwut, Ruttapon Srisonchai, Thomas Wesener, Chirasak Sutcharit, and Natdanai Likhitrakarn. 2026. Five New Species of the Pill Millipede Genus Rhopalomeris Verhoeff, 1906 (Diplopoda, Glomerida) from Thailand. Contributions to Zoology. 95(1); 52–102. DOI: doi.org/10.1163/18759866-bja10090 [20 Feb 2026]

 facebook.com/ASRUChula/posts/1351095590368500

***งานวิจัยใหม่: กิ้งกือกระสุนลูกกวาด 5 ชนิดใหม่ของโลกจากประเทศไทย
และการมีรูปแบบสีที่หลากหลายซึ่งพบในบางชนิด***

...

รายชื่อชนิดใหม่ดังนี้

.

1. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙢𝙪𝙠𝙖 Sapparojpattana & Likhitrakarn, 2026
กิ้งกือกระสุนลูกกวาดมหิดลกาญจน์
 ค้นพบที่ มหาวิทยาลัยมหิดล วิทยาเขตกาญจนบุรี อำเภอไทรโยค จังหวัดกาญจนบุรี

2. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙡𝙚𝙣𝙩𝙞𝙜𝙞𝙣𝙤𝙨𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลท่าขนุน อำเภอทองผาภูมิ จังหวัดกาญจนบุรี

3. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙙𝙪𝙡𝙘𝙞𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลบ้านนา อำเภอเมืองชุมพร จังหวัดชุมพร

4. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙥𝙪𝙣𝙘𝙩𝙖𝙩𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลไร่เก่า อำเภอสามร้อยยอด จังหวัดประจวบคีรีขันธ์

5. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙫𝙚𝙧𝙝𝙤𝙚𝙛𝙛𝙞 Sapparojpattana & Likhitrakarn, 2026
ตั้งชื่อเป็นเกียรติแก่ Karl Wilhelm Verhoeff ผู้ตั้งสกุลกื้งกือกระสุนลูกกวาด 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨
ค้นพบครั้งแรกที่ น้ำตกกะทู้ อำเภอกะทู้ จังหวัดภูเก็ต และกระจายตัวหลายพื้นที่ในภาคใต้ของประเทศไทย 

[Crustacea • 2024] Caridina maeklongensis • A New landlocked freshwater Shrimp Species (Decapoda: Atyidae) from the Mae Klong Basin, Thailand

[A–D] Caridina maeklongensis  

   Macharoenboon, Manonai & Jeratthitikul. 2024

[E, F] C. sumatrensis  De Man, 1892

RAFFLES BULLETIN OF ZOOLOGY. 72

Abstract. A newly discovered landlocked shrimp, Caridina maeklongensis, new species, is described from the Mae Klong Basin, western Thailand. The morphology of the new species resembles C. sumatrensis, from which it differs in possessing a shorter rostrum with fewer dorsal teeth, a small number of very large-sized eggs, and a greatly reduced posteromedian projection on the distal end of the telson. The molecular evidence of the mitochondrial 16S rRNA gene also supports the species status of the newly described species. This is the fifth endemic species of Caridina known from Thailand. 

Key words. crustacean, landlocked freshwater shrimp, molecular phylogeny, taxonomy, Southeast Asia

Living habitus specimens of Caridina species from the Mae Klong Basin.
A–D, Caridina maeklongensis, new species: A, paratype male, MUMNH-CAR064-4; B, paratype female, MUMNH-CAR064-1; C, paratype female, MUMNH-CAR064-15; and D, paratype female, MUMNH-CAR064-13.
 E, F, C. sumatrensis: E, male, MUMNH-CAR706-P2; and F, female, MUMNH-CAR706-P1.

Family Atyidae De Haan, 1849 

Genus Caridina H. Milne Edwards, 1837 

Caridina maeklongensis, new species

Etymology. The specific name is from its type locality, the Mae Klong Basin.

Kongkit Macharoenboon, Varitsara Manonai and Ekgachai Jeratthitikul. 2024. Caridina maeklongensis, A New landlocked freshwater Shrimp Species (Crustacea: Decapoda: Atyidae) from the Mae Klong Basin, Thailand. RAFFLES BULLETIN OF ZOOLOGY. 72: 450–468.