[Phycology • 2026] Halymenia dracophylla & H. elegans (Rhodophyta: Halymeniales) • Systematic Revision of branched Halymenia from the Philippines

Halymenia elegans Tahil & Showe M.Lin, 

in Tahil, Lin, Dumilag et Liao, 2026. 

DOI: doi.org/10.1080/09670262.2026.2738911 

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ABSTRACT

Despite early documentation of Halymenia species in the Philippines, the species diversity and phylogenetic relationships of this genus in the region, and its connections to neighbouring areas, have remained largely unexplored. In this study, we used rbcL sequence data to infer interspecific relationships among Philippine and global Halymenia species. Molecular analyses identified at least nine Halymenia species in the Philippines, including two newly described taxa – H. dracophylla sp. nov. and H. elegans sp. nov. – alongside several species commonly found throughout the IndoPacific (e.g. H. durvillei, H. tondoana, H. malaysiana, H. maculata, H. dilatata). Phylogenetic analysis showed that H. dracophylla is a sister species to the H. durvillei complex, but it can be readily distinguished by its thallus bearing numerous strap-shaped, short bladelets and having relatively large carposporophytes (260−380 µm in diameter). Conversely, H. elegans forms a lineage sister to a larger clade of branched Indo-Pacific species and differs from the other species by possessing thin, membranous thallus blades and having smaller carposporophytes (100−115 µm in diameter). The discovery of these two new species, along with the identification of additional cryptic taxa through molecular analyses, reveals that Halymenia diversity in the Philippines is greater than previously recognized. These findings highlight the need for continued investigation into the taxonomy and biogeography of Halymenia in the Indo-Pacific region.

 KEYWORDS: Cryptic species, Halymenia dracophylla sp. nov., Halymenia elegans sp. nov., marine red algae, Philippines, taxonomy

Halymenia dracophylla Tahil & Showe M.Lin sp. nov. 
Fig. 2. Holotype specimen (NTOU002614), a female gametophyte showing densely overlapping bladelets along the thallus margins.
Fig. 3. Isotype specimen (NTOU002618), a tetrasporophyte showing a deeply branched thallus blade arising from a short stipe (arrow).

Scale bars: Figs 2, 3 = 5 cm.

Halymenia dracophylla Tahil & Showe M.Lin, sp. nov.  

ETYMOLOGY: ‘dracophylla’ refers to the thallus blades swinging underwater, resembling flying dragons.

DISTRIBUTION, HABITAT AND SEASONALITY: Only known from the province of Sorsogon in southern Luzon. Plants were found during the late dry season (May) in the Philippines, in deeper subtidal areas (6–8 m in depth), growing on coral rubble and rocks. The fringing reef habitat is characterized by strong water currents and pronounced tidal waves.

SPECIMENS EXAMINED: Luzon Island: FMB Resort, Sorsogon, coll. J. Dig & C. Pangantihon 7 May 2024 (tetrasporic and female, NTOU-002614–002616).

Halymenia elegans Tahil & Showe M.Lin sp. nov. 
 Fig. 20. Holotype specimen (NTOU002610), a female gametophyte showing a palmate-shaped blade (white arrow) and feathery branches bearing numerous small bladelets (black arrows) with dentate margins.
Fig. 21. Isotype specimen (NTOU002611), a tetrasporophyte showing broad blades bearing irregularly arranged marginal bladelets (black arrows) and surface tiny bladelets (white arrowheads).  

Scale bars: Figs 20, 21 = 5 cm.

Halymenia elegans Tahil & Showe M.Lin sp. nov. 

ETYMOLOGY: ‘elegans’ referring to its showy and beautiful thallus blades.

DISTRIBUTION, HABITAT AND SEASONALITY: Only known from the type locality. Plants were found during the early dry season (January) in the Philippines, growing on shaded rocks in the reef adjacent to an estuary and extensive mangrove area on the island of Sanga-Sanga.

SPECIMENS EXAMINED: Sanga-Sanga Island, Tawi-Tawi: Lakit-Lakit, coll. A. Tahil, 28 January 2024 (tetrasporic & female, NTOU002610–002613).

Arkady A. Tahil, Showe-Mei Lin, Richard V. Dumilag and Lawrence M. Liao. 2026. Systematic Revision of branched Halymenia (Halymeniales, Rhodophyta) from the Philippines, including the Description of H. dracophylla sp. nov. and H. elegans sp. nov. European Journal of Phycology. DOI: doi.org/10.1080/09670262.2026.2738911 [22 Oct 2026]

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[Herpetology • 2026] Megophrys (Viridophrys) motuoensis • A striking New Subgenus and Species of Asian horned toads (Anura: Megophryidae) from Xizang, China

Megophrys (Viridophrys) motuoensis Liu, Zhang, Chen & Yuan,
Viridophrys Liu, Zhang, Chen & Yuan, subgen. nov.

in Liu, Zhang, Wei, Ai, Lu, Li, Zhou, Chen et Yuan, 2026. 

墨脱绿角蟾  |  DOI: doi.org/10.3897/zse.102.183945

Abstract

Motuo County, located in the eastern Himalayas and within the lower section of the Yarlung Tsangpo Grand Canyon, is characterized by steep elevational gradients and complex topography, supporting high levels of biodiversity and endemism that remain insufficiently explored. Based on integrated evidence from morphological comparisons and molecular phylogenetic analyses, a new subgenus and a new species of Asian horned toads from Motuo County, Xizang, China, are described. The new subgenus, Viridophrys subgen. nov., with its type species Megophrys (Viridophrys) motuoensis sp. nov., represents an independent evolutionary lineage within the broadly defined Megophrys, as supported by mitochondrial and nuclear DNA analyses. Although its exact sister-group relationship is not consistently resolved among different datasets, the new lineage is phylogenetically close to the subgenera Brachytarsophrys and Atympanophrys. Morphologically, the new subgenus can be distinguished from other major megophryine lineages by a combination of morphological characters. The discovery of Viridophrys subgen. nov. and M. (Viridophrys) motuoensis sp. nov. highlights the underestimated diversity of megophryids in the Yarlung Tsangpo Grand Canyon region and emphasizes the need for continued biodiversity surveys and conservation efforts in the eastern Himalayas.

Key Words: Himalayas, Megophrys (Viridophrys) motuoensis sp. nov., taxonomy, Viridophrys subgen. nov., Yarlung Tsangpo Grand Canyon region

Megophryinae Bonaparte, 1850

Genus Megophrys Kuhl & van Hasselt, 1822

Subgenus Viridophrys Liu, Zhang, Chen & Yuan, subgen. nov.

 Type species. Megophrys (Viridophrys) motuoensis
Liu, Zhang, Chen & Yuan, sp. nov.

Diagnosis. The subgenus Viridophrys subgen. nov. can be distinguished from other subgenera within the broadly defined Megophrys by the following diagnostic characteristics: (1) medium body size, mean SVL < 50 mm in adult females; (2) dorsal skin rough, with vivid green dorsal coloration and reddish markings, and ventral surfaces covered with irregular blotches; (3) head wider than long; (4) snout obtusely rounded and without rostral appendage; (5) maxillary teeth present; (6) vomerine teeth and vomerine ridges absent; (7) palpebral appendages absent, with minute tubercles along the outer edge of the upper eyelid; (8) temporal region short, tympanum large, distinct, close to posterior corner of eye, supratympanic fold thick and slightly curved over tympanum; (9) occipital fold absent; (10) dorsolateral fold absent; (11) presence of V-shaped parietoscapular ridge and arcuate central dorsal ridge; (12) flanks bearing scattered white-tipped tubercles, with a few tubercles occasionally present along the ventrolateral boundary; (13) lower jaw smooth, asperities absent; (14) subarticular tubercles on fingers absent; (15) webbing between toes is absent, toe fringe is lacking (Figs 4, 5).

Megophrys (Viridophrys) motuoensis sp. nov. adult female holotype SWU 0015893 in life.
A. Dorsal view; B. Ventral view; C. Dorsolateral view; D. Profile of head; E. Cloacal area.

Photographs by Ren-Da Ai.

Habitat of Megophrys (Viridophrys) motuoensis sp. nov. at the type locality, Beibeng Township, Motuo County, Nyingchi City, Xizang Autonomous Region, China. A. Broadleaf and coniferous mixed forest; B. Megophrys (Viridophrys) motuoensis sp. nov. in the wild (SWU 0015893).

Habitat photograph by Wen-Zhu Lu. Species photograph by Ren-Da Ai.

Etymology. The subgeneric name Viridophrys subgen. nov. is derived from the Latin viridis, meaning green, and the Greek ophrys, meaning eyebrow, in reference to the extensive green markings on the dorsum of the new taxon. The gender of the name is feminine.

Common name. Green horned toad subgenus /
Lǜ jiǎo chán yà shǔ (绿角蟾亚属).

Megophrys (Viridophrys) motuoensis Liu, Zhang, Chen & Yuan, sp. nov.

Etymology. The name “motuoensis” originated from the type locality: Motuo County, Nyingchi City, Xizang Autonomous Region, China.

Common name. Motuo green horned toad /
Mò tuō lǜ jiǎo chán (墨脱绿角蟾).

 

Xiao-Long Liu, Yin-Peng Zhang, Ping-Fan Wei, Ren-Da Ai, Wen-Zhu Lu, Xian-Qi Li, Wei-Wei Zhou, Jin-Min Chen and Zhi-Yong Yuan. 2026. A striking New Subgenus and Species of Asian horned toads (Anura, Megophryidae, Megophryinae) from Xizang, China.  Zoosystematics and Evolution. 102(5): 1295-1313. DOI: doi.org/10.3897/zse.102.183945 [21 Oct 2026]

[Ichthyology • 2024] Turcinoemacheilus ekmekciae • A New dwarf loach (Cypriniformes: Nemacheilidae) from upper Tigris and Euphrates

Turcinoemacheilus ekmekciae

Kaya, Yoğurtçuoğlu, Aksu, Bayçelebi & Turan, 2024

DOI: doi.org/10.1111/jfb.15578 

 

Abstract

Turcinoemacheilus ekmekciae, new species, from upper Euphrates and Tigris drainages is distinguished from other species of Turcinoemacheilus in Western Asia by having a dark stripe broader than the eye diameter along the lateral line, rarely possessing roundish blotches, 5–6 mandibular pores in mandibular canal, a comperatvely smaller head, a deeper body, and a greater pre-pelvic distance. Our specimens collected from the upper Great Zab, near the type locality of Turcinoemacheilus kosswigi, showed notable genetic divergence (a minimum K2P of 3.3%) from sequences reported as T. kosswigi in previous studies. Despite morphological similarities, this molecular difference suggests that the populations analysed in previous studies may represent a potential new species of Turcinoemacheilus, which we tentatively named as Turcinoemacheius cf. kosswigi. Molecular data also suggest that T. ekmekciae is characterized by a minimum K2P distance of 3.5% from Turcinoemacheilus minimus and T. cf. kosswigi. The three methods for species delimitation (assemble species by automatic partitioning [ASAP], Poisson tree processes [PTP], and multi-rate PTP [mPTP]) that were utilized for testing species assignments consistently identified our test group as a distinct species.

Keywords: cytochrome c oxidase I, freshwater, fish, taxonomy, Western Asia

Turcinoemacheilus ekmekciae,
from the top, FFR 3608, holotype, 36 mm standard length (SL); Kaynarca stream, Euphrates:
FFR 3616, paratype, 41 mm SL, Nerduş stream, Tigris:
Not preserved, ~35 mm SL; not preserved, ~40 mm SL; not preserved, ~40 mm SL; Kaynarca stream, Euphrates.

 Turcinoemacheilus ekmekciae, new species

Diagnosis: Turcinoemacheilus ekmekciae is distinguished from its congeners from upper Tigris and Euphrates drainages by a combination of characters. The new species is distinguished from T. minimus from Göksu and Kahta drainages by having a deeper caudal peduncle (7%–9% SL v. 6–7), and 5–6 mandibular pores in mandibular canal (v. 4–5). T. ekmekciae is distinguished from T. kosswigi from upper Greater Zab by having a greater pre-pelvic distance (50%–53% SL v. 47–50), a shorter upper caudal fin lobe (14%–18% SL v. 18–21), and a shorter maxillary barbel (15%–21% HL v. 21–26). The new species is further distinguished from T. kosswigi by having a dark stripe broader than the eye diameter along the lateral line, only in one population possessing roundish blotches (v. a dark-brown mid-lateral stripe narrower than the eye diameter).

Turcinoemacheilus ekmekciae is distinguished from the two Iranian Turcinoemacheilus, T. bahaii and T. hafezi, by the position of the ...

Etymology: The species is named for Fitnat Güler Ekmekçi for her contribution to the knowledge of the ichthyofauna of Türkiye. A noun in genitive, indeclinable.

 

Cüneyt Kaya, Baran Yoğurtçuoğlu, İsmail Aksu, Esra Bayçelebi and Davut Turan. 2024. Turcinoemacheilus ekmekciae, A New dwarf loach from upper Tigris and Euphrates (Teleostei: Nemacheilidae). Journal of Fish Biology. DOI: doi.org/10.1111/jfb.15578 [10 October 2023]

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[Arachnida • 2026] Songthela fengxiang, S. nanqiao, S. parallela, S. yanhe, S. yinchiba, ... • Species Diversity in the Segmented Spider Genus Songthela (Araneae: Mesothelae: Liphistiidae), with Descriptions of 12 New Species from Guizhou Province, China

[G, J] Songthela nanqiao sp. nov.;
[H, K] S. yanhe sp. nov.;
[I, L] S. yinchiba sp. nov. 

Xu, Zhang & Li, 2026

DOI: doi.org/10.3897/zse.102.216113

G–I. Female; J–L. Male, Scale bar: 5 mm.

 

Abstract

Twelve new species of the primitively segmented spider genus Songthela Ono, 2000, from Guizhou Province, China, are described based on the genital morphology of both sexes: S. chenjiapian sp. nov. (♂♀), S. cuantang sp. nov. (♂♀), S. fengxiang sp. nov. (♂♀), S. fuyan sp. nov. (♂♀), S. nanqiao sp. nov. (♂♀), S. parallela sp. nov. (♂♀), S. qingganglin sp. nov. (♂♀), S. tianba sp. nov. (♂♀), S. tudiao sp. nov. (♂♀), S. yanhe sp. nov. (♂♀), S. yinchiba sp. nov. (♂♀), and S. yuhu sp. nov. (♂♀). All new species are assigned to the multidentata-group based on the morphology of the male palp and female genitalia. Detailed diagnoses, descriptions, photographs, and a distribution map are provided. Mitochondrial cytochrome c oxidase subunit I (COI) sequences are also provided for the holotypes to facilitate future species identification and molecular studies.

Key Words: COI, description, morphology, taxonomy, trapdoor spiders

Microhabitat (A–F) and general somatic morphology (G–L) of new Songthela species.
G–I. Female; J–L. Male;
G, J. Songthela nanqiao sp. nov.; H, K. S. yanhe sp. nov.; I, L. S. yinchiba sp. nov.

Scale bar: 5 mm.

S. chenjiapian sp. nov., S. cuantang sp. nov., S. fengxiang sp. nov.,
S. fuyan sp. nov., S. nanqiao sp. nov., S. parallela sp. nov.,
S. qingganglin sp. nov., S. tianba sp. nov., S. tudiao sp. nov.,
S. yanhe sp. nov., S. yinchiba sp. nov., S. yuhu sp. nov.

Xin Xu, Yan Zhang and Daiqin Li. 2026. Species Diversity in the Segmented Spider Genus Songthela (Mesothelae, Liphistiidae), with Descriptions of 12 New Species from Guizhou Province, China. Zoosystematics and Evolution. 102(5): 1265-1293.  DOI: doi.org/10.3897/zse.102.216113 [15-10-2026]

[Ichthyology • 2026] Peckoltia kahxi • A New Species of Peckoltia (Siluriformes: Loricariidae) from the rapids of rio Trombetas Basin, Brazi

 

 Peckoltia kahxi

Dopazo, Araújo, Akama & Wosiacki, 2026

 

DOI: doi.org/10.1590/1982-0224-2026-0053

Abstract

A new species of Peckoltia is described from the rio Trombetas basin, Pará State, Brazil. The new species was investigated by morphological and molecular dataset and it can be distinguished from its congeners by: (1) mottled appearance, without defined spots or patches on the head and body, (2) partially naked abdomen with clusters of plates on the lateral sides of the abdomen from the pectoral to the pelvic region, (3) higher numbers of teeth on both jaws, (4) short odontodes in the cheek, (5) dark spots on the interradial membranes on all fins, (6) four saddles along the body, (7) lower lips with rounded papillae, (8) yellow stripes of the head obliquely positioned on head, not larger than eye orbit, (9) upper lip completely dark, and (10) dorsal margin of the hyomandibula concave. Genetic analysis revealed a genetic divergence of 2.3 to 15% between the new species and other species of the genus. Additionally, phylogenetic analysis based on a mitochondrial gene (Cytb) was conducted and the species was recognized as a monophyletic unit. The description of the new species contributes to the knowledge of the Trombetas River basin, a little-known tributary on the left bank of the Amazon River.

Keywords: Amazon basin; Ancistrini; Armoured catfish; Rheophilic species; Taxonomy

 Peckoltia kahxi, new species

 Manuela Dopazo, lFelipe Arian de Andrade Araújo, Alberto Akama and Wolmar Benjamin Wosiacki. 2026. A New Species of Peckoltia (Siluriformes: Loricariidae) from the rapids of rio Trombetas Basin, Brazil. Neotrop. ichthyol. 24(3); DOI: doi.org/10.1590/1982-0224-2026-0053

 

[Fungi • 2026] Dianjunus rex gen. et sp. nov. • Unearthing a fungal Giant: Dianjunaceae fam. nov., A Novel Paleocene Lineage of Xylariales (Ascomycota: Sordariomycetes)

 

Dianjunaceae fam. nov.
Dianjunus rex F.Q. Yu, Pérez-Moreno & Senan.,

in Song, Yan, Pérez-Moreno, Zhang, Yang, Xie, Su, J. Liu, Wang, D. Liu, Z. Yang, C. Yang, W. Liu, Shi, Wan, Cheewangkoon, Dai, Senanayake et Yu, 2026.  

DOI: doi.org/10.3389/ffunb.2026.1940464

 

Abstract

During mycological surveys in Yunnan Province, China, specimens of a fungus producing massive, upright stromata up to 50 cm high and individually 2.2 kg in weight were sampled. Through an integrative taxonomic approach combining detailed morphology, multilocus phylogeny [internal transcribed spacer (ITS), large subunit ribosomal DNA (LSU), RNA polymerase II subunit 2 (RPB2), and beta-tubulin (TUB2)], and phylogenomic analyses, this fungus is proposed as the new species Dianjunus rex gen. et sp. nov., the type of the new family Dianjunaceae (Xylariales). Phylogenetic analyses robustly place Dianjunaceae as a distinct sister clade to Graphostromataceae. Divergence time estimation dates the origin of this family to the early Paleocene (~65 million years ago), coinciding with the period following the Cretaceous–Paleogene (K–Pg) extinction, when an estimated 75% of all plant and animal species went extinct, and a significant ecological reorganization of life on Earth happened. The stromata of D. rex represent the largest fructifications documented within the Ascomycota, significantly expanding the known morphological range of the Xylariales. The study provides a comprehensive description, including a nodulisporium-like anamorph with periconiella-like branching patterns, and discusses the taxon’s phylogenetic placement and distinctive morphology. This discovery highlights the unexplored fungal diversity in East Asian forests.

Keywords: divergence time, genome sequences, large stromata, morphology, novel taxa

Stromata of Dianjunus rex in the field.
(a, c, d, f) Stromata attached to host roots. (b, e, g, h) Stromata around host plant. (a) Collectors. (i) Total weight of largest bilobedstromata.

Dianjunus rex (HKAS148941, holotype).
(a) Stromata. (b) Yellow powdery layer on outer crust covering ostioles. (c) Ostiole opening on stromatic surface. (d) Vertical cross section of ascoma. (e) Peridium. (f) Stromatic tissues in 5% (w/v) KOH. (g) Horizontal section through the stromata. (h) Apical ring in water. (i) Apical ring in Melzer’s reagent. (j) Apical ring in Lugol’s reagent after pretreatment with 3% (w/v) KOH. (k) Apical ring in Lugol’s reagent. (l) S-shaped germ-slits in ascospores in Melzer’s reagent. (m-p) Asci in water. (q-zf) Ascospores in water.

Scale bars: a = 15 cm, b, c, e = 100 µm, d = 750 µm, g = 250 µm, l = 20 µm, m-p = 50 µm, h-k, q-zf = 10 µm.

Dianjunaceae F.Q. Yu, Pérez-Moreno & Senan., fam. nov.

Etymology: Based on the type genus Dianjunus.

Diagnosis: Dianjunaceae is distinguished from other families in Xylariales by having large stromata with persistent powdery-fibrous surface, unique S-shaped germ slit in ascospores, the inverted-trapezoid, amyloid apical ring in Melzer’s reagent, and nodulisporium-like anamorph with periconiella-like branching patterns produce distantly-packed branchlets bearing cluster of cylindrical, conidiogenous cells.

 

Dianjunus F.Q. Yu, Pérez-Moreno & Senan., gen. nov.

Etymology: The generic name is derived from the “Dian Kingdom”, a historical society situated in the region of present-day Yunnan Province in southwestern China where the samples were collected. The epithet is combined with the Chinese word “jūn”, meaning ‘mushroom’ or ‘fungus’.

Dianjunus rex F.Q. Yu, Pérez-Moreno & Senan., sp. nov. 

Etymology: The species epithet rex (Latin = king) refers to the large size and majestic appearance of the stromata.

Jing Song, Zhuyue Yan, Jesús Pérez-Moreno, Fengming Zhang, Shimei Yang, Tao Xie, Luofeng Su, Jianwei Liu, Yanliang Wang, Dong Liu, Zhenyan Yang, Chengmo Yang, Wei Liu, Xiaofei Shi, Shanping Wan, Ratchadawan Cheewangkoon, Dongqin Dai, Indunil C. Senanayake and Fuqiang Yu. 2026. Unearthing a fungal Giant: Dianjunaceae fam. nov., A Novel Paleocene Lineage of Xylariales harboring Dianjunus rex gen. et sp. nov. Front. Fungal Biol. 7; 1940464. DOI: doi.org/10.3389/ffunb.2026.1940464 [29 September 2026]

[Paleontology • 2026] Yuezhuornis yangyuanensis • Jurassic flightless Bird reveals early Ecological Diversity of avialans

Yuezhuornis yangyuanensis

L. Xu, Wang, G. Lin, Dong, Deng, X. Xu, Du, M. Lin, Chen & Zhou, 2026

DOI: doi.org/10.1038/s41586-026-10972-z

Life reconstruction by Min Wang.

 Abstract

The origin and early diversification of birds remain highly debated in vertebrate evolution. Although estimates of the evolutionary timescales suggest a Jurassic period radiation, a profound scarcity of fossils from this period has hampered how the distinctive bird bauplan was assembled. Selection for powered flight has been viewed as the primary driver of early avialan skeletal modifications, attesting to the independent occurrences of aerodynamic features among paravian lineages. Here we report on the discovery of a Late Jurassic avialan, Yuezhuornis yangyuanensis, gen. et sp. nov., from the Zhenghe Fauna of southeastern China. As one of the earliest diverging avialans, Y. yangyuanensis exhibits a striking mosaic of features, combining a mixture of avialan- and troodontid-like cranial characters, paired with derived postcranial traits and a deinonychosaur-like hyperextensible second pedal digit. Phylogenetically informed analyses show that Y. yangyuanensis possesses markedly short forelimbs—a condition present widely among non-avialan theropods. These findings provide strong evidence for a terrestrial, flightless lifestyle of Y. yangyuanensis. Our study demonstrates that evolution works intensively as exemplified by the occurrence of several, divergent ecological pathways being exploited in the initial diversification of birds.

 

Yuezhuornis yangyuanensis, gen. et sp. nov.

Life reconstruction of the Late Jurassic bird Yuezhuornis yangyuanensis.

Artwork by Min Wang.

 

Liming Xu, Min Wang, Ganmin Lin, Liping Dong, Ke Deng, Xing Xu, Honggang Du, Min Lin, Runsheng Chen and Zhonghe Zhou. 2026. Jurassic flightless Bird reveals early Ecological Diversity of avialans. Nature. 658; 690–697. DOI: doi.org/10.1038/s41586-026-10972-z [14 October 2026]

Liming Xu  (徐立明), Min Wang  (王敏), Ganmin Lin  (林淦旻), Liping Dong  (董丽萍), Ke Deng  (邓克), Xing Xu  (徐星), Honggang Du  (杜红岗), Min Lin  (林敏), Runsheng Chen  (陈润生) & Zhonghe Zhou  (周忠和)

[PaleoIchthyology • 2022] Dianodipterus huizeensis • A New dipnoan Genus (Sarcopterygii: Dipnoi) from the Middle Devonian of Huize, Yunnan, China

Dianodipterus huizeensis

Luo, Cui, Qiao & Zhu, 2022

DOI: doi.org/0.1080/14772019.2022.2042409 

 Researchgate.net/publication/359575707

 

Abstract

A new genus and species of tooth-plated lungfishes, Dianodipterus huizeensis gen. et sp. nov., is described from the Qujing Formation (late Eifelian, Middle Devonian) of Huize County, Qujing, north-eastern Yunnan, China. It differs from other lungfishes in the small proportion of the tooth-bearing part on the pterygoid (less than one-quarter), the extensively developed anteromedial edentulous portion of the pterygoid, and tooth rows few in number (only four). Phylogenetic analysis reveals that Dianodipterus is more plesiomorphic than Dipterus, yet crownward of Melanognathus. As the ninth Devonian lungfish genus from China, the new taxon displays a novel schema of lungfish dentition and increases the diversity of early lungfishes.

Keywords: Dipnoi, Middle Devonian, dentition, vomer, phylogeny, China

Dianodipterus huizeensis gen. et sp. nov.

Yanchao Luo, Xindong Cui, Tuo Qiao and Min Zhu. 2022. A New dipnoan Genus from the Middle Devonian of Huize, Yunnan, China. Journal of Systematic Palaeontology. 19(18); 1303-1315 DOI: doi.org/0.1080/14772019.2022.2042409 [29 Mar 2022]

 Researchgate.net/publication/359575707_A_new_dipnoan_genus_from_the_Middle_Devonian_of_Huize_Yunnan_China

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[Botany • 2026] Primula octopoda (Primulaceae) • A stoloniferous New Species from western Sichuan, China

 

Primula octopoda W.B.Ju & Q.Huang,  

in Q. Huang, X. Wang, W.-J. Huang, Yang, Pan, Ru, J.-P. Wang et Ju, 2026.

巴塘粉报春 | DOI: doi.org/10.1002/ece3.74088 

 

ABSTRACT

Primula octopoda W.B.Ju & Q.Huang, a new species of Primula (Primulaceae) in sect. Aleuritia, is herein described and illustrated from Batang County, Sichuan Province, southwestern China. Morphologically, P. octopoda is most similar to P. stenocalyx, but it can be readily distinguished by its dwarf habit, the presence of stolons 2–7 cm in length ending in small rosettes occasionally bearing a single flower, elliptic to broadly obovate leaf blades, a very short or absent scape bearing one or two flowers, and a smaller, annulate corolla with a prominent yellow eye. The complete chloroplast genome of P. octopoda spans 154,164 bp with a GC content of 36.97% and contains 125 genes. Phylogenetic analyses using plastid and ITS data both place P. octopoda within sect. Aleuritia while supporting its distinct species status. Given the current lack of adequate information on its population size and geographic range, the species is provisionally assigned a status of Data Deficient (DD) according to the IUCN Red List criteria.

Keywords: molecular phylogeny, morphology, new species, Primula, taxonomy

Habitat of the Primula octopoda.
 (A) Habitat; (B–D) Plants.

Morphology of Primula octopoda.
 (A) Plant; (B) roots; (C) leaves; (D) inflorescence; (E) corolla; (F) bracts; (G) calyx; (H) thrum flower and pin flower; (I) capsule.

Primula octopoda W.B.Ju & Q.Huang, sp. nov. 

Diagnosis. The new species most closely resembles P. stenocalyx, sharing narrowly lanceolate bracts, a tubular calyx with similarly divided oblong or lanceolate lobes, stamens and style included within the corolla tube, and an oblong capsule. However, it differs in its shorter stature, the presence of 2–7 cm long stolons each ending in a small rosette and occasionally bearing a solitary flower, elliptic to broadly obovate leaves, a scape up to 2 cm long or absent with 1–2 flowers, and an annulate corolla with a yellow eye and smaller flowers (Figures 2 and 3).

Etymology. The specific epithet of the new species refers to its distinctive feature of producing multiple stout, fleshy stolons that resemble the tentacles of an octopus; hence the name octopoda, which vividly reflects the plant's numerous stolons extending like “eight legs.”

Vernacular Name. Chinese mandarin: “bā táng fĕn bào chūn” (巴塘粉报春), meaning “Batang Farinose Primrose” in English.

 

Qin Huang, Xin Wang, Wen-Jun Huang, Jie-Yi Yang, Hong-Li Pan, Jian Ru, Jiang-Ping Wang and Wen-Bin Ju. 2026. Primula octopoda, A stoloniferous New Species from Western Sichuan, China. Ecology and Evolution. DOI: doi.org/10.1002/ece3.74088 [27 August 2026]

[PaleoIchthyology • 2026] Yipetalichthys maxiongensis • A New quasipetalichthyid placoderm from the Middle Devonian of China

 

 Yipetalichthys maxiongensis

 Pan, Dupret, Ahlberg, Zhao, Wang, Wang & Zhu, 2026

DOI: doi.org/10.1002/spp2.70124  

 www.IVPP.ac.cn 

Abstract

We describe Yipetalichthys maxiongensis gen. et sp. nov., a quasipetalichthyid placoderm from the Eifelian Shangshuanghe Formation (Qujing, Yunnan, China). High-resolution micro-CT reveals an endocranium bearing cranial nerve grooves and inner ear structures. The dermal neck joint permits a quantitative character, the angle between the ventral laminar ridge (v.l.r) and the skull-roof midline. Current specimens yield two non-overlapping bins (state 0: ≤15°; state 1: ≥28°), separating Macropetalichthyidae and Quasipetalichthyidae; the 15–28° interval is unsampled and the threshold remains testable with new material. Expanded parsimony analysis, adding Yipetalichthys and six additional ptyctodont taxa, recovers Petalichthyida as sister to Ptyctodontida and places Yipetalichthys within Quasipetalichthyidae. The diagnosis includes large unenclosed elliptical orbits, slender unbifid postorbital processes, an elongate occipital region, and a shallow posterolateral vagus canal. Yipetalichthys shows a high v.l.r angle, supporting its quasipetalichthyid placement and providing a reproducible metric for future comparisons across articulated specimens. Our analysis of the neck joint helps to clarify petalichthyid phylogeny and identifies distinct, recurring architectures among major placoderm groups: in Antiarcha, a reverse sliding joint; in Petalichthyida, an elaborate sliding joint guided by the ventral laminar ridge with laminar cranial articular facets forming a longitudinal sliding track opposing the articular flanges of the anterior dorsolateral plate (ADL); in Ptyctodontida, a sliding joint with spoon-shaped cranial articular facets opposing ADL articular flanges; in Actinolepidoidei, a simple sliding condition; in Brachythoraci, a ginglymoid joint. These distinct neck joint architectures demonstrate the high evolutionary plasticity of this structure and its multiple, independent pathways toward cranial mobility in placoderms.

Yipetalichthys maxiongensis gen. et sp. nov.

 

Zhaohui Pan, Vincent Dupret, Per E. Ahlberg, Wenjin Zhao, Jianhua Wang, Mengjun Wang and Min Zhu. 2026. A New quasipetalichthyid placoderm from the Middle Devonian of China. Papers in Palaeontology. DOI: doi.org/10.1002/spp2.70124 [24 September 2026] 

DeepBone团队发现瓣甲鱼新属种，揭示颈胸关节演化新线索

https://www.IVPP.ac.cn/xwdt/kyjz/202609/t20260928_8287312.html

[Entomology • 2026] Anchiale mabiensis & A. robusta • A Revision of Australian Anchiale Stål, 1875 (Phasmida: Phasmatidae), with the Description of Two New Species

  Anchiale mabiensis Coupland & Emmott, sp. nov. female, in situ

in Jones, Brock, Coupland, Emmott et Lo, 2026.

DOI: doi.org/10.1111/aen.70096

 (photo by Angus J. Emmott).

ABSTRACT 

The Australian species of the Phasmida genus Anchiale Stål, 1875 (Phasmatinae: Phasmatini) are updated to include five species: Anchiale albopunctata (Kirby, 1896) stat. rev. (removed from synonymy with A. briareus (Gray, 1834)); A. briareus Gray, 1834 (= Ctenomorpha tessulata Gray, 1835 syn. nov.; = A. austrotessulata Brock & Hasenpush, 2007, replacement name for C. tessulata Gray, 1835), A. mabiensis Coupland & Emmott, sp. nov.; A. robusta Brock & Jones, sp. nov.; and A. spinicollis Gray, 1833. Lectotypes are designated for Ctenomorpha albopunctatum Kirby, 1896, C. nigrovaria Stål, 1875 (= A. briareus) and C. tessulata Gray, 1835. These taxonomic decisions are supported by Bayesian and Maximum Likelihood phylogenetic analyses based on the mitochondrial COXI gene fragment of all species except A. spinicollis. Anchiale albopunctata is currently known from mideastern Queensland, A. briareus from north-eastern New South Wales and south-eastern Queensland, A. mabiensis sp. nov. and A. robusta sp. nov. from tropical eastern Queensland, and A. spinicollis from the Northern Territory. A dichotomous key to all Australian species of Anchiale and their eggs is provided.

Keywords: Acrophylla, distribution, DNA, genus, insects, molecular, Phasmida, phylogenetic, specimens, taxonomy

Anchiale albopunctata (Kirby, 1896) stat. rev. 

(removed from synonymy with A. briareus (Gray, 1834)); 

A. briareus Gray, 1834 

(= Ctenomorpha tessulata Gray, 1835 syn. nov.; 

= A. austrotessulata Brock & Hasenpush, 2007,
replacement name for C. tessulata Gray, 1835), 

A. mabiensis Coupland & Emmott, sp. nov.; 

A. robusta Brock & Jones, sp. nov.; 

A. spinicollis Gray, 1833. 

  Anchiale mabiensis sp. nov. female, in situ

 (photo by Angus J. Emmott).

Anchiale mabiensis sp. nov. in situ.
(a) Female holotype (photo by Ross M. Coupland).
(b) paratype male (photo by Ross M. Coupland).
(c) Montane rainforest habitat (photo by Angus J. Emmott).

Anchiale mabiensis Coupland & Emmott, sp. nov. 

Anchiale robusta sp. nov. in situ.
 (a) Male. (b) Female. (c) Female defence display, inset: wings of preserved paratype female. (d) Coastal habitat, Bramston Beach, Qld. (e) Garden habitat, Kuranda, Qld

(all photos by Paul D. Brock).

Anchiale robusta Brock & Jones, sp. nov.

Braxton R. Jones, Paul D. Brock, Ross M. Coupland, Angus Emmott and Nathan Lo. 2026. A Revision of Australian Anchiale Stål, 1875 (Phasmida: Phasmatidae), with the Description of Two New Species. Austral Entomology. 65(4); e70096. DOI: doi.org/10.1111/aen.70096 [21 September 2026]

[Herpetology • 2026] Leptobrachella quangninhensis • A New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Quang Ninh Province, Vietnam

Leptobrachella quangninhensis 

Luong, Hoang, Pham, Ziegler & Nguyen, 2026

 

 http://rjh.folium.ru/index.php/rjh/article/view/2250 

 facebook.com/HoangChung2019

Abstract 

A new species of the genus Leptobrachella is described from Quang Ninh Province, Vietnam based on genetic divergence and morphological differences. Leptobrachella quangninhensis sp. nov. is characterized by a combination of the following characters: size medium (SVL 28.1 – 28.7 mm (n = 4) in adult males and 36.0 mm in one adult female); head longer than wide; tympanum distinct; belly creamy white; toe webbing rudimentary and without lateral fringes; supratympanic fold edged by a distinct black line; iris bicolored, copper in upper part, fading to golden in lower part; interorbital region with distinct black triangle; dorsal skin smooth with fine scattered tubercles. In terms of genetic distance, the new species differs from other congeners for which comparable sequences are available by at least 6.15% (16S rRNA gene). This species description brings the total number of known species in the genus Leptobrachella from Vietnam to 38.

Keywords: Leptobrachella quangninhensis sp. nov.; morphology; molecular phylogeny; taxonomy

Leptobrachella quangninhensis sp. nov.  

Anh Mai Luong, Chung Van Hoang, Cuong The Pham, Thomas Ziegler and Truong Quang Nguyen. 2026. A New Species of Leptobrachella Smith 1925 (Amphibia: Anura: Megophryidae) from Quang Ninh Province, Vietnam. Russian Journal of Herpetology. 33(3); 197-216. [September 30, 2026] 

 http://rjh.folium.ru/index.php/rjh/article/view/2250 

 facebook.com/HoangChung2019/posts/28710851798545157

[Paleontology • 2026] Cranial Anatomy of the Middle Eocene Egyptian crocodylian Herugavialis cairensis provides Nnew Insights into gavialoid diversity in the Mediterranean Region

 

Life reconstruction of Herugavialis cairensis depicted with The Great Sphinx and pyramids of Giza in the background. 

Herugavialis gen. nov.
Herugavialis cairensis n. comb. (Müller, 1927)

in Burke, Spiekman, Salem, Saber, Sallam et Mannion, 2026. 

DOI: doi.org/10.1098/rsos.261173

Artwork by Karim Zanaty.

Abstract

Gavialoidea is presently composed of only two extant crocodylian species, the Indian gharial Gavialis gangeticus, and the false gharial Tomistoma schlegelii, both from southeast Asia. However, Gavialoidea displays a rich and near-cosmopolitan diversity in the fossil record, with over 40 species currently recognized. Several of these gavialoids are recovered from Cenozoic deposits in the Mediterranean region, with many assigned historically to the genus Tomistoma. However, recent phylogenetic analyses do not typically recover these species as closely related to the false gharial, with several fossil species in need of taxonomic revision. Here, we present a detailed re-evaluation of one of these species, Tomistoma cairense from the Middle Eocene Mokattam Formation of Egypt, based on the holotype skull and several referred specimens. Supplemented by endocranial anatomy revealed via computed tomography (CT), we identify numerous anatomical differences with T. schlegelii and other gavialoids, including the approximately contemporaneous Egyptian species, Eogavialis africanum and Paratomistoma courti. These differences are reinforced through phylogenetic analysis under maximum parsimony, in which T. cairense is recovered as a gavialoid that does not cluster with T. schlegelii or Mediterranean taxa. We erect a new genus name for T. cairense, with the resultant new combination of Herugavialis cairensis diagnosed by several characteristics, including telescoped orbits. An apparent decline in Mediterranean gavialoid diversity during the Late Eocene–Oligocene is potentially driven by a reduction in habitat, as a result of global cooling and declining eustatic sea level. Warming in the Miocene might have facilitated a subsequent rebound in gavialoid diversity, prior to the extirpation of crocodylians in the Mediterranean region in the Late Miocene, which might have been caused by the Messinian Salinity Crisis. The taxonomic revision of additional Mediterranean gavialoid species should facilitate greater insight into their evolutionary history and demise.

Keywords: Gavialoidea, Mediterranean, Crocodylia, Tomistoma, neuroanatomy, phylogeny

Eusuchia Huxley, 1875

Crocodylia Gmelin, 1788

Gavialoidea Hay, 1930

Herugavialis gen. nov.,

 

Herugavialis cairensis n. comb. Müller, 1927 

Tomistoma cairense (Müller, 1927).

Etymology: Heru, after the Egyptian god (also known as Horus), is often associated with, and possibly the inspiration for, the Great Sphinx of Giza. The latter was sculpted from the Mokattam Formation, from which the type specimen of H. cairensis was recovered, with both sharing the absence of the anterior tip of their nose/snout. The genus suffix, -gavialis, reflects the relationship of H. cairensis to Gavialis gangeticus within Gavialoidea. Accordingly, the original species epithet, cairense, is modified to cairensis to agree with the masculine gender of Herugavialis.

  

Locality and horizon: The holotype and referred specimens were collected from Gebel Mokattam, east of Cairo, in northeastern Egypt. They were recovered from the Mokattam Formation, which is a nummulitic marine limestone dated to the Lutetian, Middle Eocene.

 Diagnosis: Within Gavialoidea, H. cairensis is characterized by a unique combination of characters: (i) lacrimal with an anteroposterior length more than twice that of the prefrontal; (ii) telescoped orbit formed by raised margins of the lacrimal, prefrontal and frontal; (iii) a squamosal wherein the dorsal and ventral margins do not diverge anteriorly; (iv) the distance between the distal tip of the paraoccipital process and the distal end of the quadrate condyle is greater than the maximum mediolateral width of the latter; and (v) a blunt dural peak on the encephalic endocast.

Life reconstruction of Herugavialis cairensis in left lateral view. Herugavialis is depicted with The Great Sphinx and pyramids of Giza in the background.

 Artwork by Karim Zanaty.

 

 Conclusions: 

The redescription of the holotype and referred specimens of the Middle Eocene crocodylian species ‘Tomistoma’ cairense demonstrates numerous morphological differences from extant T. schlegelii. Coupled with the recognition of several autapomorphies, as well as its phylogenetic separation from T. schlegelii, we erect the new genus, Herugavialis cairensis. Herugavialis can be differentiated from approximately contemporaneous Egyptian gavialoids by numerous anatomical features, exemplifying the diversity of gavialoids in the Mediterranean region during this period. Global cooling and a drop in eustatic sea level during the Late Eocene–Oligocene potentially limited suitable habitat for gavialoids in the Mediterranean region, with subsequent increases in eustatic sea level and temperature potentially facilitating the return to high gavialoid diversity observed in the Miocene. The taxonomic classification of some of these European species awaits re-evaluation, which will facilitate greater insight into the evolutionary history of the Mediterranean gavialoids.

Paul M. J. Burke; Stephan N. F. Spiekman; Belal S. Salem; Sara Saber; Hesham M. Sallam and Philip D. Mannion. 2026. Cranial Anatomy of the Middle Eocene Egyptian crocodylian Herugavialis cairensis provides Nnew Insights into gavialoid diversity in the Mediterranean Region. R. Soc. Open Sci 13 (10): 261173. DOI: doi.org/10.1098/rsos.261173 [07 Oct 2026]