[Entomology • 2026] Caloptilia augeas & C. ceryneia • Two New Species of Caloptilia (Lepidoptera: Gracillariidae) from New Caledonia inducing galls on Glochidion billardierei (Phyllanthaceae) and Redescription of C. xanthopharella (Meyrick, 1880)

[4-6] Caloptilia augeas
[11-13] Caloptilia ceryneia 

Guiguet, Lopez-Vaamonde, van Nieukerken & Ohshima, 

in Guiguet, van Nieukerken, Giron, Gravendeel, Lopez-Vaamonde et Ohshima, 2026. 

 DOI: doi.org/10.3897/zookeys.1268.173885 

Abstract

New Caledonia is a biodiversity hotspot with high levels of micro-endemism, yet its gracillariid fauna remains poorly documented. Here, two new species of Caloptilia Hübner, 1825 (Gracillariidae) are described from Glochidion J.R.Forst. & G.Forst. (Phyllanthaceae) host plants in Parc des Grandes Fougères, New Caledonia: Caloptilia augeas Guiguet, Lopez-Vaamonde, van Nieukerken & Ohshima, sp. nov., and Caloptilia ceryneia Guiguet, Lopez-Vaamonde, van Nieukerken & Ohshima, sp. nov. Both species induce leaf galls on Glochidion billardierei Baill., co-occurring on the same host species, sometimes even on the same leaf. They exhibit distinct wing patterns, but very similar male and female genitalia, and DNA barcoding supports their status as separate species. These findings provide evidence for potential within-host sympatric speciation, as documented in other gall-inducing insects. The larval biology of C. augeas and C. ceryneia reveals a unique frass disposal behaviour, whereby waste is excreted through a hole and the aperture is subsequently sealed—an adaptation not previously reported in gall-inducing Lepidoptera. Our findings double the known number of gall-inducing species in Gracillariidae, highlighting that this life history strategy may be more common than currently appreciated. We also provide new information on distribution and host plants of Caloptilia xanthopharella (Meyrick, 1880), a leaf roller found on the same host plant, G. billardierei. These findings mark the first records of the subfamily Gracillariinae in New Caledonia. This study underscores the underexplored diversity of New Caledonian gracillariids and emphasises the conservation value of Parc des Grandes Fougères. Further surveys in the Indo-Pacific region may reveal additional yet undescribed Caloptilia species associated with Phyllanthaceae and help clarify the evolutionary mechanisms underpinning their diversification.

Key words: Australia, DNA barcoding, endemism, frass disposal, host plant use, sympatry, taxonomy

Habitus of adult Caloptilia augeas sp. nov. 

4. Resting posture; 5. Paratype, male RMNH.INS.25563; 6. Holotype, male, RMNH.INS.25562.
 

Habitus of adult Caloptilia ceryneia sp. nov.
11. Resting posture, holotype male, RMNH.INS.25650; 12. Paratype, female, RMNH.INS.25565; 13. Paratype, male, RMNH.INS.25564.
Scale bars: 2 mm.

Caloptilia augeas Guiguet, Lopez-Vaamonde, van Nieukerken & Ohshima, sp. nov., 

Etymology. The specific name augeas refers to the fifth Labour of Hercules in the Greek mythology. The larval behaviour of frass cleaning in the gall reminds this Labour that consisted in cleaning the stables of king Augeas (Αὐγέας). This name comes from the word “αὐγή” that designates “sun shine” and “reflection of a shiny object” in Classic Greek (Bailly 1899), referring to the bright colour of the adult wings. The epithet is to be regarded as a noun in apposition.

Caloptilia ceryneia Guiguet, Lopez-Vaamonde, van Nieukerken & Ohshima, sp. nov.

Etymology. The specific name ceryneia refers to the ancient Greek city of Ceryneia or Keryneia (Κερύνεια), the location of the Ceryneian Hind from Greek mythology, which features in the third Labour of Hercules. The moth’s striking yellow and red wing pattern evokes the golden antlers and bronze hooves of the mythical deer. Its cryptic gall-inducing behaviour also recalls the elusive nature of the Ceryneian hind, which was known for its ability to remain hidden. The epithet is to be treated as a noun in apposition.

 Antoine Guiguet, Erik J. van Nieukerken, David Giron, Barbara Gravendeel, Carlos Lopez-Vaamonde, Issei Ohshima. 2026. Two New Species of Caloptilia (Lepidoptera, Gracillariidae) from New Caledonia inducing galls on Glochidion billardierei (Phyllanthaceae) and Redescription of C. xanthopharella (Meyrick, 1880). ZooKeys. 1268: 113-137. DOI: doi.org/10.3897/zookeys.1268.173885 [04 Feb 2026]

[Funga • 2025] Samsoniella scoliopterygis (Hypocreales: Cordycipitaceae) • A New Species of Samsoniella on the herald moth, Scoliopteryx libatrix (Noctuoidea: Erebidae), from caves in the British Isles

Samsoniella scoliopterygis  H.C. Evans & J.P.M. Araújo,

in Evans, Fogg, Buddie, Yeap et Araújo. 2025. 

DOI: doi.org/10.3114/fuse.2025.15.09 

 

Abstract

Samsoniella scoliopterygis (Cordycipitaceae: Hypocreales), pathogenic on the herald moth Scoliopteryx libatrix (Erebidae: Noctuoidea), is described as a new species, based on morphological, ecological and molecular evidence. Infected specimens of this troglophilic moth were recorded from caves, as well as from a copper mine, in southern England (Devon, Norfolk), Northern Ireland (Fermanagh) and the Republic of Ireland (Sligo). The moths were found consistently on the cave floor or rock ledges, rather than on the ceiling amongst the overwintering, healthy moth populations; producing multiple, white or non-pigmented clavae of the cordyceps-like sexual morph. Synnemata of the asexual morph occurred sporadically in situ but were produced consistently in vitro. The ecology and taxonomy are discussed in relation to previous records of entomopathogenic fungi on Scoliopteryx libatrix and other trogloxene moths.

 Keywords: Samsoniella scoliopterygis; ecology; entomopathogenic fungi; new taxon; phylogenetics; systematics 

Samsoniella scoliopterygis sp. nov., holotype IMI 507611.
A, B. Sexual morph on herald moth, (Scoliopteryx libatrix), dried specimen from floor of Boho Caves (Co. Fermanagh, Northern Ireland), showing the superficial perithecia on stromata emerging from the dorsal surface and orifices (mouth and anus). C, D. Transverse section of an ascostroma showing the superficial, flask-shaped perithecia with a wide, prominent neck region and wall. E. Perithecial tip showing the asci and ascospores oozing from the ostiole. F. Ascus showing prominent cap.

 Scale bars: A, B = 2 mm; C = 100 µm; D = 30 µm; E = 15 µm; F = 5 µm.

Samsoniella scoliopterygis, IMI 507613, on herald moths in situ in Boho Caves, July 2024.
 A. Infected moth on ledge in twilight zone (arrow), ca. 6 m from cave entrance in background. B. Close-up of stromata showing the white, superficial, crowded perithecia. C–E. Infected moths in various positions within the dark zone of the cave system, all with white stromata.

Scale bars: B = 1 mm; C, E = 8 mm; D = 5 mm.

Samsoniella scoliopterygis H.C. Evans & J.P.M. Araújo, sp. nov.

 

H.C. Evans, T. Fogg, A.G. Buddie, Y.T. Yeap and J.P.M. Araújo. 2025. A New Species of Samsoniella (Cordycipitaceae: Hypocreales) on the herald moth, Scoliopteryx libatrix (Erebidae: Noctuoidea), from caves in the British Isles. Fungal Systematics and Evolution. 15(1); 201-224. DOI: doi.org/10.3114/fuse.2025.15.09 (June 2025) 

x.com/joaofungo/status/1885628021882659155

[Entomology • 2020] Apsidophora bala & A. chandrapatyae • Two New Species of the Genus Apsidophora Diakonoff, 1973 (Lepidoptera: Tortricidae: Olethreutinae) from Thailand

[2, 5] Apsidophora bala, sp. nov. (holotype, male)
[3, 6] A. bala, sp. nov. (paratype, females)
[4, 7] A. chandrapatyae, sp. nov. (holotype, male) 

Pinkaew & Muadsub, 2020

DOI: doi.org/10.11646/zootaxa.4877.3.1 

Researchgate.net/publication/346815150

Abstract

 Two new species, Apsidophora bala sp. nov. and Apsidophora chandrapatyae sp. nov., are described. Illustrations of adult and genitalia are provided. We also present a photograph of living specimen of A. chandrapatyae sp. nov. in natural resting posture. The two new taxa increase the number of described Apsidophora species to 3.

 Key words: new species, Olethreutini, Thailand

 

 Head of Apsidophora spp. 5. Apsidophora bala, sp. nov. (holotype, male) 6. A. bala, sp. nov. (paratype, female np13274) 7. A. chandrapatyae, sp. nov. (holotype, male).

 Wing pattern of Apsidophora spp. 2. Apsidophora bala, sp. nov. (holotype, male) 3. A. bala, sp. nov. (paratype, female np6675) 4. A. chandrapatyae, sp. nov. (holotype, male), scale bars = 2 mm.

 Male hindwing of Apsidophora spp.
8–9. Apsidophora bala, sp. nov. (holotype, male) 8. Hindwing 9. Projecting lobe
10–13. A. chandrapatyae, sp. nov. (holotype, male) 10. Hindwing 11. Projecting lobe 12. Two groups of sex scales on hindwing (upperside hindwing) 13. A row of long scales (underside hindwing).

Nantasak Pinkaew and Sopita Muadsub. 2020. Two New Species of the Genus Apsidophora Diakonoff, 1973 (Lepidoptera: Tortricidae: Olethreutinae) from Thailand. Zootaxa. 4877(3); 401–412. DOI: doi.org/10.11646/zootaxa.4877.3.1 

Researchgate.net/publication/346815150_Two_new_Apsidophora_from_Thailand

[Entomology • 2026] Stygioides hecate • A Golden Shadow in the Mountains: A New Stygioides Bruand, 1853 (Lepidoptera: Cossidae) from Turkey

 Stygioides hecate Japaridze & Junnilainen, 

in Japaridze, Junnilainen et Sihvonen, 2026.  

 DOI: doi.org/10.11646/zootaxa.5821.2.8

 

Abstract

A new species of the genus Stygioides Bruand, 1853 (Lepidoptera: Cossidae) is described from central Turkey. Stygioides hecate Japaridze & Junnilainen sp. nov. is based on material collected in the Sultan Daðlarý Mountains (Konya Province). The new species is characterised by its distinct wing coloration in females. Adults are diurnal, as in other members of the genus, and were collected by daytime netting. DNA barcode data (COI) are provided and compared with available sequences in the Barcode of Life Data Systems (BOLD Systems), supporting the distinctiveness of the new taxon. The discovery of this species increases the number of Stygioides species to 10 and those from Turkey to three, thus contributing to a better understanding of the diversity and distribution of the genus in the Palaearctic realm.

Lepidoptera, carpenter moths, DNA barcoding, Heteroneura, Middle East

Adults of Stygioides hecate Japaridze & Junnilainen sp. nov.
 A–B. Paratype male, dorsal and ventral view and associated labels.
C–D. Holotype female, dorsal and ventral view and associated labels.

Scale bar: 10 mm

Stygioides hecate Japaridze & Junnilainen sp. nov. 

Diagnosis: Males of S. hecate Japaridze & Junnilainen sp. nov. differ from S. nupponenorum by the presence of a distinct white tip of the male abdomen and white scales on the antennae, the absence of brown piliform scales on the abdomen, a narrower apex of the valvae, and a shorter saccus (Figs 1A–1B, 2 C; cf. Lingenhöle et al. 2016: fig. 6; Yakovlev 2011: fig. 35). Females are readily distinguished from all congeners by the presence of golden-yellow scales on the antennae, head, thorax and wings (Figs 1C–1D; cf. Lingenhöle et al. 2017: figs 1–3, 9; Japaridze et al. 2025b: fig. 22).

Etymology: The species name is derived from Hecate, the ancient Greek goddess associated with the night and commonly depicted carrying yellow or golden torches. This name alludes to the coloration of the holotype female, which is predominantly black with yellow-golden scales on the wings, thorax, head, and antennae.

LASHA-GIORGI JAPARIDZE, JARI JUNNILAINEN and PASI SIHVONEN. 2026. A Golden Shadow in the Mountains: A New Stygioides Bruand, 1853 (Lepidoptera: Cossidae) from Turkey.  Zootaxa. 5821(2); 274-280. DOI: doi.org/10.11646/zootaxa.5821.2.8 [2026-05-29]

[Entomology • 2026] Pyralis papaleonei • A New Species (Lepidoptera: Pyralidae) from Crete (Greece)

[2, 3, 6, 9] Pyralis papaleonei sp. nov.

Huemer, Kaila & Segerer, 2026;

[4] P. kacheticalis;    [5] P. regalis  

DOI: doi.org/10.3897/nl.49.185483

Abstract 

Pyralis papaleonei sp. nov. is described based on diagnostic phenotypic and genital morphological characters as well as unique DNA barcodes (COI gene – cytochrome c oxidase subunit 1) and compared with closely related taxa occurring in the Eastern Mediterranean, namely P. kacheticalis and P. regalis. The new species is endemic to the island of Crete and currently only known from the White Mountains (Lefka Ori) in the western part of the island. Pyralis kacheticalis is recorded from Iraq and Armenia for the first time.

Pyralis spp., habitus.
2. Pyralis papaleonei sp. nov., holotype, ♂; 3. P. papaleonei sp. nov., paratype, ♂;
4. P. kacheticalis, ♂, Georgia; 5. P. regalis ♂, Italy (TLMF). 

Pyralis papaleonei sp. nov.

Diagnosis. Pyralis papaleonei sp. nov. is distinguished from all congeners by its narrow white forewing markings, featuring an almost uniformly slender white median band extending across the entire wing (Figs 2, 3). In all other species of the group (Figs 4, 5), except for specimens of P. kacheticalis from Cyprus, the white pattern elements are distinctly broader, and in some cases abruptly narrowed medially or terminating in the middle of the wing. Also characteristic are the nearly uniform pale greyish-purple hindwings, which are divided into three sections by two white lines; the basal section is only slightly darker than the outer sections. In contrast, in ...

Etymology. The new species is dedicated to the head of the Catholic Church, Pope Leo XIV. The Pontiff is a strong advocate of climate and environmental protection, and we hope that his voice may serve as an example for humanity. Furthermore, due to its distinctive coloration and overall appearance, the new species belongs to a group of Pyralidae whose species names refer to high secular or ecclesiastical offices, including Pyralis regalis, Pyralis imperialis (= P. kacheticalis), Pyralis princeps, and Pyralis cardinalis.

 Peter Huemer, Lauri Kaila and Andreas H. Segerer. 2026. Pyralis papaleonei sp. nov. from Crete (Greece) (Lepidoptera, Pyralidae). Nota Lepidopterologica. 49: 63-74. DOI: doi.org/10.3897/nl.49.185483 [28 Apr 2026]

[Entomology • 2026] Teulisna breveprocessa & T. nicobara • Two New Species and A New Species Record of Genus Teulisna Walker (Lepidoptera: Erebidae: Arctiinae) from India

[1-2]. Teulisna breveprocessa sp. nov., Devikulam, Kerala, India;
[3] T. ruma (Swinhoe. 1889), Nilgiri, Tamil Nadu, India; [4] T. basigera (Walker [1865]), Tamil Nadu, India (NZCZSI); [5] T. inducta (Walker 1864), Naduvattam, Tamil Nadu, India;
[6-8]. T. nicobara sp. nov., Great Nicobar Islands, Govind Nagar, India.

S. Singh, N. Singh & Joshi, 2026 

DOI: doi.org/10.1016/j.japb.2026.03.007

Abstract

Two new species of the genus Teulisna Walker, 1862 (Lepidoptera: Erebidae: Lithosiina) are described from India: T. (Cossa) breveprocessa sp. nov. (from the Western Ghats of Kerala) and T. (Tegulata) nicobara sp. nov. (from the Great Nicobar Island). The former is diagnosed with its close congeners in the subgenus Cossa Walker: T. (C.) ruma (Swinhoe, [1890] 1889), T. (C.) munnara Volynkin, 2023, T. (C.) basigera (Walker, [1865]) and T. (C.) inducta (Walker, [1865]), whereas the latter is diagnosed with T. (Tegulata) chiloides chiloides (Walker 1862). T. (Poikilothosia) sayinyingpo Huang, Volynkin & Černý, 2025, from Arunachal Pradesh, which is a new record to India.

Keywords: Kerala, Lithosiina, Nicobar Islands, NorthEast Himalaya, Tegulata, Western Ghats

Habitus of Teulisna spp.
1. Teulisna breveprocessa sp. nov., m# HT, Devikulam, Kerala, India (NZCZSI); 2. T. breveprocessa sp. nov., m# PT, Devikulam, Kerala, India (NZCZSI);
3. T. ruma (Swinhoe. 1889) m# Avalanchi, Nilgiri, Tamil Nadu, India (NZCZSI); 4. T. basigera (Walker [1865]) m# Pykara, Tamil Nadu, India (NZCZSI);
5. T. inducta (Walker 1864), m#, Naduvattam, Tamil Nadu, India (NZCZSI);
6. T. nicobara sp. nov., m# HT Great Nicobar Islands, Govind Nagar, India (NZCZSI); 7. T. nicobara sp. nov., m# PT Great Nicobar Islands, Govind Nagar, India (NZCZSI); 8. T. nicobara sp. nov., f# PT, Great Nicobar Islands, Govind Nagar, India (NZCZSI);
9. T. sayinyingpo Huang, Volynkin & Černý, 2025 m#, Sela, Arunachal Pradesh, India (NZCZSI); 10. T. sayinyingpo Huang, Volynkin & Černý, 2025 f#, Sela, Arunachal Pradesh, India (NZCZSI).

Teulisna (Cossa) breveprocessa sp. nov. (from the Western Ghats of Kerala) 

T. (Tegulata) nicobara sp. nov. (from the Great Nicobar Island).

Santosh Singh, Navneet Singh and Rahul Joshi. 2026. Two New Species and A New Species Record of Genus Teulisna Walker (Lepidoptera: Erebidae: Arctiinae) from India. Journal of Asia-Pacific Biodiversity. In Press. DOI: doi.org/10.1016/j.japb.2026.03.007 [20 April 2026]

[Entomology • 2026] Pseudodelta melas • A New Wasp-mimicking Clearwing Moth (Lepidoptera: Sesiidae: Sesiini) from Uganda

Pseudodelta melas

Bartsch & Sáfián, 2026

DOI: doi.org/10.11646/zootaxa.5807.1.9 

Abstract

In this study a new genus of Sesiini, Pseudodelta gen. nov., and its type species Pseudodelta melas sp. nov., are described from the Ziika (Zika) Forest, Uganda. This highly mimetic species bears a striking resemblance to Delta emarginatum (Linnaeus, 1758), the black mud wasp widespread in Africa, which is probably its model.

Lepidoptera, Cossoidea, new species, Afrotropical region, taxonomy

Pseudodelta melas holotype, male (1) upperside, (2) underside.

Pseudodelta melas holotype, genitalia structure.

Pseudodelta gen. nov. 

Etymology. The name refers to ancient Greek ψευδο [pseudo] = false, pretended, not real, and the genus Delta Fabricius, 1804 (Vespidae); gender is neuter.

Pseudodelta melas sp. nov. 

Etymology. Ancient Greek μέλας [mélas] = black, dark.

DANIEL BARTSCH and SZABOLCS SÁFIÁN. 2026. A New Wasp-mimicking Clearwing Moth from Uganda (Lepidoptera: Sesiidae: Sesiini).  Zootaxa. 5807(1); 184-188. DOI: doi.org/10.11646/zootaxa.5807.1.9  [2026-05-08]

[Entomology • 2026] Euthalia zubeengargi • A New Species of Euthalia Hübner, 1819 subgenus Limbusa Moore, [1897] (Lepidoptera: Nymphalidae: Limenitidinae) from Arunachal Pradesh, Northeast India

 

Euthalia (Limbusa) zubeengargi 

Sadasivan & Upadhaya, 

in Upadhaya et Sadasivan, 2026. 

  http://www.entomon.in

 Researchgate.net/publication/403195003

 

ABSTRACT 

A new species of Euthalia Hübner, 1819 (subgenus Limbusa Moore, [1897]) (Lepidoptera, Nymphalidae, Limenitidinae, Adoliadini) is described from Arunachal Pradesh, northeastern India. This species, Euthalia (Limbusa) zubeengargi, Sadasivan & Upadhaya sp. nov., is assigned to the Patala group of Limbusa based on its wing pattern, venation, and male genital morphology. A detailed description of the male and its genitalia is presented, along with a revised key to the males of Subtype A2 of the Patala group sensu Yokochi (2011). 

KEY WORDS: New taxon, butterfly, duke, Patala group, morphology, Basar, Eastern Himalayas, taxonomic key

Euthalia (Limbusa) zubeengargi, Sadasivan & Upadhaya sp. nov.

 Roshan Upadhaya and Kalesh Sadasivan. 2026. A New Species of Euthalia Hübner, 1819 subgenus Limbusa Moore, [1897] (Lepidoptera, Nymphalidae, Limenitidinae, Adoliadini) from Arunachal Pradesh, Northeast India. ENTOMON. 51(1): 1-10. 

Researchgate.net/publication/403195003_A_new_species_of_Euthalia_from_ArunachalPradesh_NE_India

  http://www.entomon.in/index.php/Entomon/forthcoming

[Entomology • 2026] Staphylus neideae, S. ricardoi, ... • Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera: Hesperiidae: Pyrginae), with Descriptions of Four New Species

[M–P] Staphylus (C.) nicoleae Lemes, sp. nov.; [Q, R] S. (C.) neivae sp. nov.; 
[S, T] S. (C.) ricardoi Lemes, sp. nov.; [U–X] S. (C.) neideae Lemes, sp. nov.: 

in Lemes, Siewert, Mielke, Casagrande et Warren, 2026. 

DOI: doi.org/10.3897/asp.84.e179328 

Abstract

The taxonomy of the subgenus Staphylus (Capilla) is reviewed, including redescriptions of known species, identification keys, and detailed distribution maps. The female genitalia of Staphylus (Capilla) azteca (Scudder, 1872), S. (C.) caribbea (Williams & Bell, 1940), S. (C.) corumba (Williams & Bell, 1940), S. (C.) eryx Evans, 1953, S. (C.) tucumanus (Plötz, 1884) and S. (C.) tyro (Mabille, 1878) are described and illustrated for the first time. Four new species are described: Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, S. (C.) ricardoi Lemes, sp. nov. from Peru, S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil. Pholisora imperspicua Hayward, 1940 is a syn. nov. of Staphylus (C.) lizeri lizeri (Hayward, 1938) and Hesperia melangon epicaste Mabille, 1903 is a syn. nov. of Staphylus (C.) melangon melangon (Mabille, 1883). Neotypes are designated for Staphylus epicaste Mabille, 1903, Nisoniades tucumanus Plötz, 1884 and Staphylus fascia Hayward, 1933. Lectotypes are designated for Pholisora azteca Scudder, 1872, Bolla machuca Schaus, 1913, Helias tyro Mabille, 1878, Staphylus anginus Schaus, 1902, Hesperia melangon 1883, Hesperia musculus Burmeister, 1875 and Helias aurocapilla Staudinger, 1876.

Keywords: Butterflies, Cytochrome oxidase subunit 1, Neotropical, Skippers

Species of Staphylus (Capilla) in dorsal and ventral views.  
A–D S. (C.) ascalon: A, B male (Brazil, São Paulo, Teodoro Sampaio, Morro do Diabo, OM 17.996); C, D female (Brazil, Minas Gerais, Passa Quatro, DZ 46.813).
E, F S. (C.) buena: male (Bolivia, Santa Cruz, Buenavista, holotype, CMNH 7188).
G–J S. (C.) caribbea: G, H male (Costa Rica, Limón Province, La Bomba, MGCL 1112082); I, J female (Costa Rica, Limón Province, Germania, MGCL 1112078).
K–N S. (C.) eryx: K, L male (Brazil, Pará, Santarém, DZ 46.566); M, N female (Brazil, Pará, Santarém, MGCL 1112109).
O–R S. (C.) corumba: O, P male (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.033); Q, R female (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.005).
S, T S. (C.) lizeri album: male (Brazil, Minas Gerais, Carmo do Rio Claro, OM 2.788). U–X S. (C.) lizeri lizeri: U, V male (Peru, Madre de Dios, Tambopata Reserve, DZ 46.176); W, X female (Colombia, Meta, Villavencio, Bosque Bavaria, DZ 45.526).

 Scale bar = 1 cm.

Species of Staphylus (Capilla) in dorsal and ventral views.
A–D S. (C.) tucumanus: A, B male (Argentina, Salta, Pichanal, DZ 45.125); C, D female (Paraguay, Presidente Hayes, Estancia Victoria, Rio Norte Lindo, OM 37.963).
E–H S. (C.) azteca: E, F male (El Salvador, La Libertad, Santa Tecla, MGCL 1112133); G, H female (Mexico, Chiapas, San Jerónimo, DZ 45.085).
I–L S. (C.) tyro: I, J male (Venezuela, Aragua, Ocumare de la Costa, DZ 22.893); K, L female (Venezuela, Aragua, Ocumare de la Costa, DZ 46.608).

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944).
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944). 
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, 

S. (C.) ricardoi Lemes, sp. nov. from Peru, 

S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil.  

 José Ricardo Assmann Lemes, Ricardo Russo Siewert, Olaf Hermann Hendrik Mielke, Mirna Martins Casagrande and Andrew David Warren. 2026. Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera, Hesperiidae, Pyrginae, Carcharodini), with Descriptions of Four New Species. Arthropod Systematics & Phylogeny. 84: 123-173. DOI: doi.org/10.3897/asp.84.e179328 [24 Feb 2026]

[Entomology • 2026] Covellana niomalan, Sosxetra mamanina & S. thutakanay • Revision of the Comose Flame Moths of the Genus Sosxetra Walker (Lepidoptera: Noctuidae: Dyopsinae), with Descriptions of A New Genus and Three New Species

 [16, 17, 27] Covellana niomalan Martinez, Homziak, Plotkin & Castillo-Argaez gen. et sp. nov.; 

[5, 6, 26] Sosxetra grata Walker, 1862; 

[9, 10] Sosxetra mamanina Martinez, Homziak, & Castillo-Argaez, sp. nov. 

[11, 12] S. thutakanay Martinez, Homziak, & Castillo-Argaez, sp. nov.

in Martinez, Homziak, Pierson, Campo, Plotkin et Castillo-Argaez, 2026. 

DOI: doi.org/10.3897/zookeys.1268.138260

Abstract

As part of our Neotropical Dyopsinae Project, a revision of the Neotropical genus Sosxetra Walker is proposed. Morphological and molecular evidence challenge its previous monotypic classification. The genus is redescribed and a neotype is designated for Sosxetra grata Walker, previously considered the only species in the genus. Two new species are described: Sosxetra mamanina Martinez, Homziak, & Castillo-Argaez, sp. nov. and Sosxetra thutakanay Martinez, Homziak, & Castillo-Argaez, sp. nov. Finally, Covellana Martinez, Homziak, Plotkin & Castillo-Argaez, gen. nov., is established based on Covellana niomalan Martinez, Homziak, Plotkin & Castillo-Argaez, sp. nov., previously misidentified as Sosxetra grata.

Key words: Barcoding, cryptic species, Lepidoptera, morphology, neotropics, systematics, taxonomy

Adult head structure. 2. Sosxetra grata, ♂, Yucatan, Mexico, MGCL; 3. S. grata, ♀, Alajuela, Costa Rica, MGCL;
4. Covellana niomalan, Paratype, ♂, Madre de Dios, Peru, MGCL.

Adult habitus of Sosxetra and Covellana.
5. Sosxetra grata, ♂, neotype, Espirito Santo, Brazil, MPM; 6. S. grata, syntype of S. agatha, ♂, Suriname, MfN; 7. S. grata, ♂, Napo, Ecuador, MGCL; 8. S. grata, ♂, Yucatan, Mexico, MGCL;
9. S. mamanina, ♂, holotype, Madre de Dios, Peru, MGCL; 10. S. mamanina, ♂, paratype, Madre de Dios, Peru, MGCL; 11. S. thutakanay, ♂, holotype, Napo, Ecuador, MGCL; 12. S. thutakanay, ♂, paratype, French Guiana, MGCL;
13. S. grata, syntype of S. agatha, ♀, Suriname, MfN; 14. S. grata, ♀, Pichincha, Ecuador, MGCL; 15. S. grata, ♀, paratype, Alajuela, Costa Rica, MGCL;
16. Covellana niomalan, ♂, holotype, Canal Zone, Panama, MGCL; 17. C. niomalan, ♂, paratype, Madre de Dios, Peru, MGCL.

Resting position of Sosxetra and Covellana.
 26. S. grata, ♂, Cayo District, Belize photo by Thomas Shahan;
27. Covellana niomalan Barro Colorado Island, Panama photo by Maxim Larrivée.

Sosxetra mamanina Martinez, Homziak, & Castillo-Argaez, sp. nov. 

Sosxetra thutakanay Martinez, Homziak, & Castillo-Argaez, sp. nov. 

 Covellana Martinez, Homziak, Plotkin & Castillo-Argaez, gen. nov., 

Covellana niomalan Martinez, Homziak, Plotkin & Castillo-Argaez, sp. nov.

 Jose I. Martinez, Nicholas T. Homziak, Taylor L. Pierson, Rhys J. L. Campo, David M. Plotkin and Raiza J. Castillo-Argaez. 2026. Revision of the Comose Flame Moths of the Genus Sosxetra Walker (Noctuidae, Dyopsinae), with Descriptions of A New Genus and Three New Species. ZooKeys. 1268: 227-248.  DOI: doi.org/10.3897/zookeys.1268.138260 [06 Feb 2026]

[Entomology • 2025] Pterotopteryx ivonini, P. elenaivshinae & P. ismailovi • First Records of Many-plumed Moths (Lepidoptera: Alucitidae) from Laos

 

[3, 3a] Pterotopteryx ivonini
[8, 8a] P. ismailovi 

Kovtunovich & Ustjuzhanin, 2025

 DOI: doi.org/10.37828/em.2025.93.14

Abstract

Three new species of many-plumed moths are described from Laos: Pterotopteryx ivonini Kovtunovich & Ustjuzhanin, sp. n., Pterotopteryx elenaivshinae Kovtunovich & Ustjuzhanin, sp. n., and Pterotopteryx ismailovi Kovtunovich & Ustjuzhanin, sp. n. The first image of the male genitalia of Pterotopteryx toxophila (Meyrick, 1906), comb. n. This work reports the first records of many-plumed moths from Laos.

Pterotopteryx ivonini Kovtunovich & Ustjuzhanin, sp. n.
3. Adult, male (Holotype, ZISP);  3a. Reconstructed image of adult in dorsal view.  

Pterotopteryx ismailovi Kovtunovich & Ustjuzhanin, sp. n.
 8. Adult, female (Holotype, ZISP);
8a. Reconstructed image of adult in dorsal view.

 Pterotopteryx ivonini Kovtunovich & Ustjuzhanin, sp. n. 

 Pterotopteryx elenaivshinae Kovtunovich & Ustjuzhanin, sp. n. 

 Pterotopteryx ismailovi Kovtunovich & Ustjuzhanin, sp. n.

Vasily Kovtunovich, Petr Ustjuzhanin. 2025. First Records of Many-plumed Moths (Lepidoptera: Alucitidae) from Laos. Ecologica Montenegrina. 93; 157-167.  DOI: doi.org/10.37828/em.2025.93.14

[Entomology • 2024] Actinote pyrrhosticta • A New Species and eight new subspecies of high elevation Actinote (Nymphalidae: Heliconiinae: Acraeini) from Colombia, Ecuador and Peru

 

Actinote pyrrhosticta Lamas, Willmott & Hall,

C-H, Actinote  pyrrhosticta apurimac n. ssp., Peru, Cuzco; 

   I) Actinote pyrrhosticta n. ssp.?, Peru, La Libertad, 

J) A. hilaris arcoiris n. ssp. female, Ecuador, Morona-Santiago,   

K, L) Actinote hilaris sourakovi n. ssp. female, Peru, Amazonas,   
   

in Willmott, Lamas, Hall, Boyer, Pyrcz et Florczyk, 2024.  

Tropical Lepidoptera Research. 34(1) 

  journals.flvc.org/TropLep/article/view/134801

Abstract  

We describe one new species and eight new subspecies of high elevation Actinote Hübner, [1819] (Nymphalidae, Heliconiinae, Acraeini) from Colombia, Ecuador and Peru: Actinote pyrrhosticta Lamas, Willmott & Hall, n. sp., Actinote pyrrhosticta quintecocha Lamas & Willmott, n. ssp., Actinote pyrrhosticta apurimac Lamas & Willmott, n. ssp., Actinote pyrrhosticta alfamayo Lamas & Willmott, n. ssp., Actinote eresia albesia Lamas & Willmott, n. ssp., Actinote eresia canyaris Pyrcz & Lamas, n. ssp., Actinote hilaris arcoiris Willmott & Hall, n. ssp., Actinote hilaris sourakovi Willmott & Lamas, n. ssp., and Actinote trinacria alegria Boyer & Willmott, n. ssp. We treat Actinote binghamae Dyar, 1913 as a new synonym of Actinote eresia eresina (Hoffer, 1874) n. syn., and treat Altinote santamarta Winhard, 2017 as a subspecies, Actinote trinacria santamarta rev. stat. We illustrate adult specimens of all described taxa of A. pyrrhosticta n. sp., A. eresia (C. Felder & R. Felder, 1862) and A. hilaris Jordan, 1910, and representatives of A. trinacria (C. Felder & R. Felder, 1862). We also provide illustrations of male and female genitalia for select taxa, distribution maps, and a revised, annotated synonymic list for all four species.

  

Images of new Actinote taxa in nature.
A, B) Mating pair of Actinote pyrrhosticta n. sp., female in foreground (A), and male (B), resting on rocks and low vegetation near edge of stream gully at type locality;
C-H, Actinote  pyrrhosticta apurimac n. ssp., Peru, Cuzco, above Mollepata (photographs by David Geale). C) Last instar; D) Pupal case with freshly emerged adult adjacent; E) Cluster of pupae; F,G) Males nectaring on Asteraceae flowers; H) Female nectaring on Asteraceae flowers;
I) Actinote pyrrhosticta n. ssp.?, Peru, La Libertad, Cochorco (photograph by Jonathan Newman);
J) A. hilaris arcoiris n. ssp. female, puddling on damp gravel, Ecuador, Morona-Santiago, Cebadas-Macas road;
K, L) Actinote hilaris sourakovi n. ssp. female, dorsal (K) and ventral (L), Peru, Amazonas, Abra Patricia (photographs by David Geale).  

Habitats of new Actinote taxa. A) Ecuador, Loja, Jimbura-San Andrés road, type locality of Actinote pyrrhosticta n. sp., stream gully and road where numerous individuals were observed on the morning of 12 June 2014; B) Peru, Cajamarca, El Pargo, habitat of A. pyrrhosticta n. sp. C) Peru,Apurímac, Santuario Nacional Ampay, Laguna Angasocha, type locality of A. pyrrhosticta apurimac n. ssp.D) Peru,Apurímac, Santuario Nacional Ampay, Laguna Uspacocha, habitat of A. pyrrhosticta apurimac n. ssp.E) Peru,Lambayeque, Cañaris, type locality of A. eresia canyaris n. ssp.F) Ecuador, Zamora-Chinchipe, San Francisco, type locality of Actinote hilaris arcoiris n. ssp.  

Keith R. Willmott, Gerardo Lamas, Jason P. W. Hall, Pierre Boyer, Tomasz Pyrcz and Klaudia Florczyk. 2024. A New Species and eight new subspecies of high elevation Actinote from Colombia, Ecuador and Peru (Nymphalidae, Heliconiinae, Acraeini). Tropical Lepidoptera Research. 34(1); 1-20.

  journals.flvc.org/TropLep/article/view/134801

 inabio.biodiversidad.gob.ec/2023/10/27/una-nueva-especie-y-ocho-nuevas-subspecies-de-mariposas-son-descritas-en-colombia-ecuador-y-peru/

En este trabajo describimos una especie nueva y ocho nuevas subspecies altoandinas de Actinote Hübner, [1819] (Nymphalidae, Heliconiinae, Acraeini) encontradas en Colombia, Ecuador y Perú: Actinote pyrrhosticta Lamas, Willmott & Hall, n. sp., Actinote pyrrhosticta quintecocha Lamas & Willmott, n. ssp., Actinote pyrrhosticta apurimac Lamas & Willmott, n. ssp., Actinote pyrrhosticta alfamayo Lamas & Willmott, n. ssp., Actinote eresia albesia Lamas & Willmott, n. ssp., Actinote eresia canyaris Pyrcz & Lamas, n. ssp., Actinote hilaris arcoiris Willmott & Hall, n. ssp., Actinote hilaris sourakovi Willmott & Lamas, n. ssp., y Actinote trinacria alegria Boyer & Willmott, n. ssp. Consideramos a Actinote binghamae Dyar, 1913 como un nuevo sinónimo de Actinote eresia eresina (Hoffer, 1874) n. syn., y a Altinote santamarta Winhard, 2017 como una subespecie, Actinote trinacria santamarta rev. stat. Ademas, ilustramos especímenes adultos de todos los taxa descritos de A. pyrrhosticta n. sp., A. eresia (C. Felder & R. Felder, 1862) y A. hilaris Jordan, 1910, y algunos representantes de A. trinacria (C. Felder & R. Felder, 1862). También presentamos ilustraciones de las genitalias masculinas y femeninas de algunos taxones seleccionados, junto con mapas de distribución, y una lista de sinonimos revisada para las cuatro especies. 

Shinichi Nakahara, Albert Thurman, Gordon B. Small. 2024. A new species of Pseudodebis Forster, 1964 from Panama (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera Research. 34(1); 21-28.