Showing posts with label Egypt. Show all posts
Showing posts with label Egypt. Show all posts

Friday, March 27, 2026

[PaleoMammalogy • 2026] Masripithecus moghraensis • An Early Miocene Ape from the Biogeographic Crossroads of African and Eurasian Hominoidea


Masripithecus moghraensis
Al-Ashqar, Seiffert, El-Sayed, Salem, Gohar, El-Saka, Amin & Sallam, 2026

Illustration by Mauricio Antón

Abstract
The Early Miocene fossil record documenting hominoid evolution has long been restricted primarily to sites in East Africa, whereas contemporaneous North African sites have only yielded remains of cercopithecoid monkeys. Here, we describe a fossil ape from North Africa, a new genus (Masripithecus) from the Early Miocene (~17 million to 18 million years) of northern Egypt, on the basis of mandibular remains. A combined molecular-morphological Bayesian tip-dating analysis positions Masripithecus closer to crown hominoids than coeval fossil apes from East Africa, thereby filling a phylogenetic and biogeographic gap in the evolution of stem hominoids. This evidence suggests that crown Hominoidea might have originated during the Early Miocene in the underexplored northeastern part of Afro-Arabia, rather than in eastern Africa or Eurasia.
 

Masripithecus moghraensis


Illustration of Masripithecus moghraensis by Mauricio Antón

Masripithecus moghraensis and the dispersal of crown hominoids in the Miocene. The map highlights Wadi Moghra, Egypt (star), which is the discovery site of Masripithecus—the first definitive North African ape—alongside key Miocene hominoid localities (see table S1) across Afro-Arabia and Eurasia. Arrows indicate inferred dispersal routes based on the phylogenetic and biogeographic analyses presented here. The inset phylogeny places Masripithecus as the closest sampled sister taxon of crown Hominoidea. At lower left, a life reconstruction of Masripithecus based on the Masripithecus mandible combined with the facial morphology of the middle Miocene hominoid Pierolapithecus. Illustration by Mauricio Antón
 

SHOROUQ F. AL-ASHQAR, ERIK R. SEIFFERT, SANAA EL-SAYED, BELAL S. SALEM, ABDULLAH S. GOHAR, HOSSAM EL-SAKA, MOHAMED AMIN, AND HESHAM M. SALLAM. 2026. An Early Miocene Ape from the Biogeographic Crossroads of African and Eurasian Hominoidea. SCIENCE. 391(6792); 1383-1386. DOI: doi.org/10.1126/science.adz4102 [26 Mar 2026]
Editor’s summary: The vast majority of early hominoid fossil hunting has occurred in East Africa, where a trove of early fossils and lineages have been found. Other regions in Africa have been less explored for various reasons, inspiring the question of whether a focus on East Africa has shaped opinions about where early hominoid evolution occurred. Al-Ashqar et al. now describe a Miocene ape from Egypt with crown hominoid affinities suggesting both that this lineage diverged before entering Eurasia and that a focus on one African region may have shaped our ideas about where hominoids first emerged (see the Perspective by Alba and Arias-Martorell). —Sacha Vignieri

Monday, October 27, 2025

[Paleontology • 2025] Wadisuchus kassabi • An early dyrosaurid (Pseudosuchia: Crocodyliformes) from the Campanian of Egypt sheds light on the Origin and Biogeography of Dyrosauridae

 
Wadisuchus kassabi
Saber, Salem, Ouda, Gohar, El-Sayed, O’Connor & Sallam, 2025 

Paleoart by Nathan Dehaut. 

Abstract
Dyrosauridae are a clade of crocodyliforms characterized by diverse cranial morphologies and a broad palaeogeographic distribution from the Late Cretaceous to the Palaeogene. However, their early evolutionary history remains poorly understood due to a significant fossil gap during the Campanian. Here, we describe Wadisuchus kassabi gen. et sp. nov., an early-diverging dyrosaurid from the middle Campanian Quseir Formation of Egypt, based on two partial skulls and three partial mandibles. This new taxon displays transitional cranial features—including reduced premaxillary alveoli, modified occlusion patterns, and dorsally positioned external nares—that clarify aspects of cranial evolution related to longirostry in early dyrosaurids. Phylogenetic analyses consistently recover Wadisuchus as the earliest-diverging dyrosaurid, closely related to Chenanisuchus and distinct from Elosuchus, supporting a transition from dyrosauroids to dyrosaurids. Its Campanian age extends the temporal range of the clade and suggests that transatlantic dispersal from Africa to South America occurred earlier than previously recognized. Alongside Brachiosuchus kababishensis from Sudan, the new Egyptian taxon also implies that reverse dispersal into Africa preceded the Maastrichtian. Wadisuchus provides critical insights into the early diversification, palaeobiogeography, and cranial evolution of Dyrosauridae, confirming longirostry as an early-acquired trait and highlighting North Africa as a key region in their origin.

Campanian, Quseir Formation, Egypt, North Africa, Dyrosauridae, phylogeny

Life reconstruction of Wadisuchus kassabi from the Late Cretaceous of Egypt, showing an adult capturing a lungfish in a wetland, with a juvenile nearby. The scene highlights the ancient ecosystem, including turtles and lush vegetation, based on fossil evidence from the Quseir Formation.
Paleoart by Nathan Dehaut. / MUVP—Scientific supervision

Wadisuchus kassabi gen. et sp. nov.

Sara Saber, Belal S Salem, Khaled Ouda, Abdullah S Gohar, Sanaa El-Sayed, Patrick M O’Connor and Hesham M Sallam. 2025. An early dyrosaurid (Wadisuchus kassabi gen. et sp. nov.) from the Campanian of Egypt sheds light on the Origin and Biogeography of Dyrosauridae. Zoological Journal of the Linnean Society. 205(2) zlaf134. DOI: doi.org/10.1093/zoolinnean/zlaf134 [27 October 2025]

Friday, March 14, 2025

[Entomology • 2025] Davidlivingstonia mooseri • A New Species and A New Record of the Afrotropic Genus Davidlivingstonia Yakovlev, 2020 (Lepidoptera: Cossidae) in the Middle East


[A-B]. Davidlivingstonia lenzi Yakovlev, 2020;
[C-D] Davidlivingstonia mooseri Yakovlev, Müller, Saldaitis & Prozorov, 2025 
 

 Abstract
A new species, Davidlivingstonia mooseri sp. n. is described from Oman and compared with the closely related D. boisduvalii (Herrich-Schäffer, 1854), which is known from Sierra Leone. Additionally, Davidlivingstonia lenzi Yakovlev, 2020 is recorded for Egypt for the first time. These new records significantly extend the known distribution of the Afrotropical genus Davidlivingstonia Yakovlev, 2020, demonstrating its range reaches much farther north and northeast than previously documented.
 
Keywords: Cossoidea, Egypt, Oman, Saudi Arabia, new species, taxonomy

Davidlivingstonia, adult males and their genitalia:
A. D. lenzi, male, S. Egypt; B. D. lenzi, male genitalia, slide Prozorov 2022 0457;
C. Davidlivingstonia mooseri, Holotype; D. D. mooseri, male genitalia, slide Prozorov 2022 0458.
 

Roman V. Yakovlev, Günter C. Müller, Aidas Saldaitis and Alexey M. Prozorov. 2025. A New Species and A New Record of the Afrotropic Genus Davidlivingstonia Yakovlev, 2020 (Lepidoptera: Cossidae) in the Middle East. Zoology in the Middle East. 71(1); 63-67. DOI: doi.org/10.1080/09397140.2025.2444693 [09 Jan 2025]


Yakovlev, R.V. 2020. On the taxonomy of Zeuzera boisduvalii Herrich-Schäffer, 1854 (Cossidae, Zeuzerinae) species complex with description of A New Genus and Three New Species. Ecologica Montenegrina. 38, 215–226. DOI: doi.org/10.37828/em.2020.38.31

Tuesday, February 18, 2025

[PaleoMammalogy • 2025] Cranial Anatomy of the hypercarnivore Bastetodon syrtos gen. nov. (Hyaenodonta: Hyainailourinae) and A Reevaluation of Pterodon in Africa

  

Bastetodon syrtos (Holroyd, 1999)
Bastetodon 
gen. nov.

Al-Ashqar, Borths, El-Desouky,  Heritage, Abed, Seiffert, El-Sayed & Sallam, 2025
artwork by Ahmed Morsi

ABSTRACT
During the Paleogene in Afro-Arabia, most terrestrial mammalian carnivores belonged to Hyaenodonta, an extinct lineage bearing a pair of carnassials between each set of molars. The Fayum Depression of Egypt preserves multiple lineages of hyaenodonts across the Eocene–Oligocene boundary. Here, we describe one of the most complete hyaenodont crania ever recovered from the lower Oligocene of the Fayum (Jebel Qatrani Formation, Quarry I, ∼30 Ma). The cranium is about the size of a hyena’s and preserves the complete upper tooth row. The long, shearing metastyle and reduced and mesially shifted protocones indicate a hypercarnivorous diet, and robust zygomatic arches suggest strong chewing muscles. Based on dental comparisons, we refer the cranium to Bastetodon syrtos (previously “Pterodonsyrtos). Bastetodon syrtos has three premolars and two molars—a reduced dental formula compared with other Fayum hyainailourines such as Akhnatenavus and European hyainailourines such as Pterodon. The cranium preserves the clover-shaped lambdoidal crest and long pharyngeal tube that unites Hyainailouroidea (Apterodontinae, Teratodontinae, and Hyainailourinae). The new Fayum cranium allows us to reevaluate the genus Pterodon and explore characters that distinguish Afro-Arabian and Eurasian species placed in this genus. Our phylogenetic analysis recovers a paraphyletic Pterodon, with B. syrtos as the sister taxon of Falcatodon schlosseri. We erect Sekhmetops to describe the large Fayum hyainailourine species S. phiomensis and S. africanus. This effort clarifies the biogeographic history of the clade that includes Pterodon, revealing multiple Tethys Seaway dispersals during the Paleogene followed by endemic radiations in Eurasia and Afro-Arabia.



Genus  Bastetodon gen. nov.

Pterodon syrtos Holroyd, 1999:8, fig. 4 (original description).
 
Etymology—Bastetodon, from “Bastet,” one transliteration of the name of the ancient Egyptian, cat-headed goddess of protection, pleasure, and bringer of good health. She was also associated with the lion-headed goddess Sekhmet. And ancient Greek “odon” meaning tooth. Literally, “Teeth like the cat-headed goddess.”


 

 Genus SEKHMETOPS gen. nov.

Type Species—Sekhmetops africanus (Andrews, 1904)
Included Species—Sekhmetops phiomensis (Osborn, 1909)

Etymology—From Sekhmet, the wrathful lion-headed goddess of Ancient Egypt associated with war and pestilence. And “ops” meaning “face” in ancient Greek. Literally, “Having the face of the lion-headed goddess.”


Shorouq F. Al-Ashqar, Matthew Borths, Heba El-Desouky, Steven Heritage, Mohamed Abed, Erik R. Seiffert, Sanaa El-Sayed and Hesham M. Sallam. 2025. Cranial Anatomy of the hypercarnivore Bastetodon syrtos gen. nov. (Hyaenodonta, Hyainailourinae) and A Reevaluation of Pterodon in Africa. Journal of Vertebrate Paleontology. e2442472. DOI: doi.org/10.1080/02724634.2024.2442472

Notably, this study refutes the monophyly of “Pterodon.” Based on the morphological features and the phylogenetic analyses conducted within this study, we suggest that Pterodon has never been found in Afro-Arabia. “Pterodon syrtos is now placed in Bastetodon syrtos and “Pterodon africanus and “Pterodon phiomensis are placed in the new genus Sekhmetops. Under this taxonomic scheme, Pterodon is now restricted to the Eocene of Europe.


Wednesday, January 15, 2025

[Paleontology • 2025] Tameryraptor markgrafi • Re-evaluation of the Bahariya Formation carcharodontosaurid (Theropoda: Carcharodontosauridae) and its implications for allosauroid phylogeny


Tameryraptor markgrafi
Kellermann, Cuesta & Rauhut, 2025
 
Life reconstruction by Joschua Knüppe.

Abstract
The first partial skeleton of a carcharodontosaurid theropod was described from the Egyptian Bahariya Oasis by Ernst Stromer in 1931. Stromer referred the specimen to the species Megalosaurus saharicus, originally described on the basis of isolated teeth from slightly older rocks in Algeria, under the new genus name Carcharodontosaurus saharicus. Unfortunately, almost all of the material from the Bahariya Oasis, including the specimen of Carcharodontosaurus was destroyed during World War II. In 1996, a relatively complete carcharodontosaurid cranium was described from similar aged rocks in Morocco and designated the neotype of the species Carcharodontosaurus saharicus in 2007. However, due to the destruction of the original material, comparisons of the neotype to the Egyptian fossils have so far only been done cursorily. A detailed reexamination of the available information on the Egyptian carcharodontosaurid, including a previously undescribed photograph of the exhibited specimen, reveals that it differs from the Moroccan neotype in numerous characters, such as the development of the emargination of the antorbital fossa on the nasals, the presence of a horn-like rugosity on the nasal, the lack of a dorsoventral expansion of the lacrimal contact on the frontals, and the relative enlargement of the cerebrum. The referability of the Egyptian specimen to the Algerian M. saharicus is found to be questionable, and the neotype designation of the Moroccan material for C. saharicus is accepted here under consideration of ICZN Atricle 75, as it both compares more favorably to M. saharicus and originates from a locality closer to the type locality. A new genus and species, Tameryraptor markgrafi gen. et sp. nov., is proposed for the Egyptian taxon. The theropods of the Bahariya Oasis and the Moroccan Kem Kem Group are thus not as closely related as previously thought, and the proposed faunal similarities between these two strata need further examination.

The holotype of Tameryraptor markgrafi (SNSB-BSPG 1922 X46).
Photograph of the mounted specimen at a point prior to April 1944.

Comparison between Tameryraptor markgrafi [SNSB-BSPG 1922 X 46] and SGM-Din 1.
SNSB-BSPG 1922 X 46: (B) nasalia from left lateral, left maxilla from (D) lateral and (F) medial and endocast in (G) dorsal and (H) left lateral view;  
and SGM-Din1: (A) right nasal in lateral view, right maxilla in (C) lateral and (E) medial view and endocast in (I) dorsal and (J) left lateral view;
Sources: (A, C, E, J) [18], (I) modified from [54], (B, D) UAT 678/20/SNSB-BSPG 1922 X 46, (F) [1], (G, H) MB. R. 2056.

Systematic Paleontology
Theropoda Marsh, 1881
Allosauroidea Marsh, 1878

Carcharodontosauriformes new clade
comprising Carcharodontosaurus saharicus, Sinraptor dongi and all descendants of their most recent common ancestor

Carcharodontosauridae Stromer, 1931

Tameryraptor gen nov. 

Etymology: Tameryraptor meaning “thief from the beloved land” is a combination of one of the more informal ancient Egyptian names for Egypt, (ta-mery) meaning beloved land [Silver, 2021]; and the latin word for thief, raptor
 
  Tameryraptor markgrafi sp. nov.

Etymology: markgrafi in reference to Richard Markgraf, the Austrian fossil collector who discovered most of the dinosaur remains described by Stromer.

Holotype: SNSB-BSPG 1922 X 46, now destroyed. The specimen included parts of the left and right maxilla, both nasals, parts of the braincase and endocast of the braincase, three cervical vertebrae, an anterior to anterior mid-caudal vertebra and chevron, the proximal part of a dorsal rib, a left ischium, right and left pubis, both femora and a left fibula. In the absence of the actual specimen, the descriptions and figures of Stromer [1, 25] and the photograph UAT 678/20 stand as representatives for the holotype, in accordance with ICZN article 73.1.4.

Horizon and Locality: SNSB-BSPG 1922 X 46 was found two kilometers from Ain Gedid on the Western foot of the Gebel Harra (Fig 1) in basal layers of hardened, gypsum-free marl (Layer p of Stromer’s 1914 profile) of the Bahariya Formation, Cenomanian.

Diagnosis: Tameryraptor markgrafi gen. nov sp. nov. is a carcharodontosaurid theropod characterized by the following unique character combination (autapomorphies marked with a *): antorbital fossa visible on nasals in lateral view; small nasal horn on the medial rim of the nasals*; maxilla with well-developed antorbital ridge; maxillary teeth highly symmetrical anteroposteriorly; prefrontal facet on frontal not expanded; frontals vaulted dorsally*; femur with spike like accessory trochanter; proximal fibula more anteriorly than posteriorly expanded.

Skeletal reconstruction of SNSB-BSPG 1922 X 46. Preserved elements shown in white. Each square is 1 sq.m.

Life reconstruction of Tameryraptor markgrafi.
By Joschua Knüppe.


Maximilian Kellermann, Elena Cuesta and Oliver W. M. Rauhut. 2025. Re-evaluation of the Bahariya Formation carcharodontosaurid (Dinosauria: Theropoda) and its implications for allosauroid phylogeny. PLoS ONE. 20(1): e0311096. DOI: doi.org/10.1371/journal.pone.0311096

Friday, May 31, 2024

[Botany • 2018] Phylogeny and Taxonomic Synopsis of Poa subgenus Pseudopoa (including Eremopoa and Lindbergella) (Poaceae: Poeae: Poinae)

 

  Poa subgenus Pseudopoa sect. Pseudopoa

in Gillespie, Soreng, Cabi & Amiri, 2018. 

Abstract
Eremopoa is a small genus of annual grasses distributed from Egypt to western China. Phylogenetic analyses of plastid and nuclear ribosomal DNA show that Eremopoa species, together with the monotypic genus Lindbergella and a single species of Poa (P. speluncarum), are nested within the genus Poa, in a clade that we accept as Poa subg. Pseudopoa. Here we accept seven species, four subspecies and four varieties in Poa subg. Pseudopoa. Five new combinations are made: Poa attalica, P. diaphora var. alpina, P. diaphora var. songaricaP. nephelochloides and P. persica subsp. multiradiata; P. millii is proposed as a replacement name for E. capillaris; and Poa sections Lindbergella and Speluncarae are proposed. We provide a diagnosis for Poa subg. Pseudopoa, synonymy for and a key to the taxa. Eight lectotypes are designated: Eragrostis barbeyi Post, Eremopoa nephelochloides Roshev., Glyceria taurica Steud., Nephelochloa tripolitana Boiss. & Blanche, Poa cilicensis Hance, Poa paradoxa Kar. & Kir., Poa persica var. alpina Boiss and Poa persica subsp. cypria Sam. Eremopoa medica is re-identified as a species of Puccinellia.

Keywords: Annuals, classification, DNA, Eremopoa, grasses, Lindbergella, phylogeny, Poa, Poaceae, taxonomy

  Poa subgenus Pseudopoa sect. Pseudopoa.
 A P. diaphora subsp. diaphora var. diaphora, Chu, Kyrgyz Republic (Soreng et al. 7537)
B, C Ppersica subsp. persica, Adiyaman, Turkey (Soreng et al. 9215) B habit C closeup of base of plant showing keeled leaf sheaths and caniculate blades
 D, E P. persica subsp. multiradiata, Mardin, Turkey (Soreng et al. 9240) D habit E spikelet showing glabrous lemmas. Photos by R.J. Soreng.


 Lynn J. Gillespie, Robert John Soreng, Evren Cabi and Neda Amiri. 2018. Phylogeny and Taxonomic Synopsis of Poa subgenus Pseudopoa (including Eremopoa and Lindbergella) (Poaceae, Poeae, Poinae). PhytoKeys. 111: 69-101.  DOI: 10.3897/phytokeys.111.28081

Friday, August 11, 2023

[PaleoMammalogy • 2023] Tutcetus rayanensis • A diminutive New basilosaurid Whale (Pelagiceti: Basilosauridae) reveals the Trajectory of the Cetacean Life Histories during the Eocene


Tutcetus rayanensis
Antar, Gohar, El-Desouky, Seiffert, El-Sayed, Claxton & Sallam, 2023

Artwork: Ahmed Morsi

Abstract
Soon after whales originated from small terrestrial artiodactyl ancestors, basal stem forms (archaeocetes) came to inhabit more specialized aquatic ecologies and underwent a tremendous adaptive radiation that culminated in the adoption of a fully aquatic lifestyle. This adaptive strategy is first documented by the geographically widespread extinct family Basilosauridae. Here we report a new basilosaurid genus and species, Tutcetus rayanensis, from the middle Eocene of Fayum, Egypt. This new whale is not only the smallest known basilosaurid, but it is also one of the oldest records of this family from Africa. Tutcetus allows us to further test hypotheses regarding basilosaurids’ early success in the aquatic ecosystem, which lasted into the latest Eocene, and their ability to outcompete amphibious stem whales and opportunistically adapt to new niches after they completely severed their ties to the land. Tutcetus also significantly expands the size range of the basilosaurids and reveals new details about their life histories, phylogeny, and paleobiogeography.




Systematic paleontology
Mammalia Linnaeus, 1758
Cetacea Brisson, 1762
Pelagiceti Uhen, 2008
Basilosauridae Cope, 1868

Tutcetus, new genus

Etymology: A combination of Tut, for the ancient Egyptian Pharoah Tutankhamun, commonly known as King Tut, who unexpectedly died in his 18th year, and cetus, Greek for a whale. Genus name is used in reference to the subadult status and the diminutive size of the type specimen.

Type species: Tutcetus rayanensis, new genus and species.

Generic diagnosis: The specimen was assigned to the family Basilosauridae based on the presence of multiple accessory cusps on the cheek teeth and well-developed pterygoid sinuses around the auditory region. Tutcetus rayanensis differs from other basilosaurids by its diminutive size (possibly the smallest known basilosaurid); it further differs in having a maxilla that abuts most of the lateral sides of the nasal, leaving only a small anterior portion of the nasal to articulate with the ascending process of the premaxilla; the number of mesial and distal accessory cusps on the upper and lower premolars (mainly two mesially and three distally). Furthermore, the premolars of Tutcetus rayanensis are more gracile than those of any other known basilosaurid and have extremely smooth enamel, and the fourth premolar (P44) is the largest tooth in both the upper and lower jaws. Tutcetus rayanensis lacks replacement of the first premolar. The Supplementary Information provides a more detailed diagnosis (Supplementary Note 1).

Tutcetus rayanensis, new species

Etymology: A combination of “Rayan,” in reference to the Wadi El-Rayan Area, the locality of the holotype, and “-ensis” (Gr., N.L. masc. adj.).

  


 
Mohammed S. Antar, Abdullah S. Gohar, Heba El-Desouky, Erik R. Seiffert, Sanaa El-Sayed, Alexander G. Claxton and Hesham M. Sallam. 2023. A diminutive New basilosaurid Whale reveals the Trajectory of the Cetacean Life Histories during the Eocene. Communications Biology. 6: 707. DOI: 10.1038/s42003-023-04986-w

Wednesday, July 26, 2023

[Paleontology • 2023] Khargachelys caironensis • The First Side-Necked Turtle (Pleurodira: Bothremydidae) from the Campanian (Late Cretaceous) of Egypt


Khargachelys caironensis 
AbdelGawad, Pérez-García, Hirayama, Mohesn, Tantawy & El-Kheir, 2023


Abstract
The Quseir Formation is an Upper Cretaceous (Campanian) deposit in the Kharga oasis of the Southwestern Desert (Egypt). This formation comprises a clastic sequence of bioturbated mudstone and sandstone intercalations, including rare scattered and fragmented vertebrate remains such as shark teeth, dinosaur remains, and turtle plates. These deposits indicate a supratidal marsh environment. A complete shell of a turtle discovered from the Quseir Formation, at the Kharga oasis, is attributed here to Bothremydini (Pleurodira, Bothremydidae), and determined as a new taxon: Khargachelys caironensis gen. et sp. nov. This form represents the only Bothremydini member currently identified in the Campanian record not only of Egypt but also of North Africa. Therefore, it helps fill the missing evolutionary gap from the Late Cretaceous pleurodires in Egypt and in North Africa.

Keywords: Campanian; Quseir Formation; Kharga; Bothremydini; Khargachelys caironensis gen. et sp. nov


  NVP005, the shell of the bothremydid turtle Khargachelys caironensis gen. et sp. nov., from the Campanian age of Kharga Oasis, South Western Desert, Egypt, in dorsal (A,A’) and ventral (B,B’) views.

Systematic Paleontology
Testudines Batsch, 1788 
Pleurodira Cope, 1864 
Pelomedusoides Cope, 1868 

Bothremydidae Baur, 1891 
Bothremydini Gaffney, Tong and Meylan, 2006  

Khargachelys caironensis gen. et sp. nov
 
Holotype: NVP005, complete carapace and plastron.

Type locality and horizon: Qarn Gennah (Kharga oasis); Quseir Formation; Campanian age.

Diagnosis: Member of Bothemydini that differs from the other representatives of this clade by the following character combination: long shell close to 60 cm in length; oval-shaped carapace, reaching its widest point at the fifth peripherals; narrow and shallow nuchal emargination; seven neurals; almost as long and wide as the first pair of peripherals; first vertebral scute being the widest of the vertebral series; second to the fourth vertebrals noticeably longer than wide; first pair of marginals overlapping half the lateral nuchal margins; anterior margin of the plastron reaching the anterior carapace edge; anterior plastral lobe wider than the posterior one; twice as wide as long entoplastron; entoplastron width equivalent to almost half the anterior plastral lobe width; extragulars overlapping the entoplastron; humero-pectoral sulcus located behind the epiplastra, showing a relatively long distance between them; pectoral scutes overlapping the anterior region of the mesoplastra.


Description: The carapace is almost complete but partially broken (Figure 2). The damaged and crushed parts are due to the pressure from the overlying sediments. The carapace is estimated to be about 580 mm long and 430 mm wide. The height of the carapace cannot ...

Etymology:Kharga” honors Kharga Oasis, where the specimen was collected; “chelys” is Greek for, turtle; “caironensis” honors Cairo, the capital city of Egypt, and honors Cairo University.


Conclusions:
The described turtle shell here, from the Quseir Formation of Kharga Oasis, in the South Western Desert (Egypt), represents the first record of Bothremydidae (Pleurodira) from the Campanian age in Egypt and in Africa. The Campanian deposits in the Kharga area indicate a supratidal marsh environment, concordant with the occurrence of such aquatic turtles. The specimen is attributed to a new genus and species of Bothremydini, Khargachelys caironensis gen. et sp. nov. This specimen brings new data to the poorly known diversity of turtles from the Late Cretaceous of Egypt.
The restricted brackish water environments and the gradual shallow shelf together with fossils such as freshwater gastropods and palynofloras, support indications of the ability of Bothremydidae to reside in different environments, such as freshwater and coastlines. The diversity of the paleoenvironments of the Bothremydidae enabled them to reach a wide distribution in both the northern and southern continents during the Late Cretaceous, along the coastlines of the Atlantic Ocean, breaking the barriers of the vicariance pattern of evolution.


 Mohamed AbdelGawad, Adán Pérez-García, Ren Hirayama, Sara Mohesn, Abdel-Aziz Tantawy and Gebely Abu El-Kheir. 2023. The First Side-Necked Turtle (Pleurodira, Bothremydidae) from the Campanian (Late Cretaceous) of Egypt. Diversity. 15(2); 284. DOI: 10.3390/d15020284
 (This article belongs to the Special Issue Fossil Reptiles and Associated Faunal Record)

Friday, July 21, 2023

[Paleontology • 2023] Igai semkhu • A New titanosaurian (Dinosauria: Sauropoda) from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Egypt


 Igai semkhu
Gorscak, Lamanna, Schwarz, Díez Díaz, Salem, Sallam & Wiechmann, 2023


ABSTRACT
Dinosaur fossils from the latest Cretaceous (Campanian–Maastrichtian) of Africa and the Arabian Peninsula are rare. Most discoveries to date have consisted of limited fossils that have precluded detailed phylogenetic and paleobiogeographic interpretations. Fortunately, recent discoveries such as the informative Egyptian titanosaurian sauropod dinosaur Mansourasaurus shahinae are beginning to address these long-standing issues. Here we describe an associated partial postcranial skeleton of a new titanosaurian taxon from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Western Desert of Egypt. Consisting of five dorsal vertebrae and 12 appendicular elements, Igai semkhu gen. et sp. nov. constitutes one of the most informative dinosaurs yet recovered from the latest Cretaceous of Afro-Arabia. The relatively gracile limb bones and differences in the coracoid and metatarsal I preclude referral of the new specimen to Mansourasaurus. Both model-based Bayesian tip-dating and parsimony-based phylogenetic analyses support the affinities of Igai semkhu with other Late Cretaceous Afro-Eurasian titanosaurs (e.g., Mansourasaurus, Lirainosaurus astibiae, Opisthocoelicaudia skarzynskii), a conclusion supported by posterior dorsal vertebrae that lack a postzygodiapophyseal lamina, for example. Igai semkhu strengthens the hypothesis that northern Africa and Eurasia shared closely related terrestrial tetrapod faunas at the end of the Cretaceous and further differentiates this fauna from penecontemporaneous assemblages elsewhere in Africa, such as the Galula Formation in Tanzania, that exhibit more traditional Gondwanan assemblages. At present, the specific paleobiogeographic signal appears to vary between different dinosaur groups, suggesting that Afro-Arabian Cretaceous biotas may have experienced evolutionary and paleobiogeographic histories that were more complex than previously appreciated.


DINOSAURIA Owen, 1842

SAUROPODA Marsh, 1878
TITANOSAURIA Bonaparte and Coria, 1993
LITHOSTROTIA Upchurch et al., 2004

Location of discovery and quarry map of Igai semkhu (Vb-621–640).
A, map of Egypt showing the location of the town of Baris in the Kharga Oasis region, denoted by orange star (modified from Sallam et al., Citation2018:fig. 1a); B, satellite image from Google Earth Pro of the research area south of Baris with approximate quarry location indicated by orange star; C, quarry map showing disposition of skeletal elements in situ with currently missing and/or obliterated elements in gray, modified from Wiechmann (Citation1999b:17); and D, skeletal silhouette (reversed and modified from Sallam et al., Citation2018:fig. 1c) with elements described in the current study shown in orange.
Abbreviations: cor, coracoid; dv, dorsal vertebra; fib, fibula; mtc I, metacarpal I; mtc IV, metacarpal IV; mtc V, metacarpal V; mtt I, metatarsal I; mtt II, metatarsal II; pub, pubis; tib, tibia; ul, ulna.
 
 IGAI SEMKHU gen. et sp. nov.

Diagnosis— Characters supporting Igai semkhu as a titanosaurian sauropod dinosaur: dorsal vertebrae lacking hyposphene-hypantrum articulations; and ulna with prominent olecranon process. Autapomorphic characters of Igai semkhu: metacarpal V with proximomedial and distomedial tubercles; reduced cnemial crest of the tibia (does not exceed anterior margin of distal end of the tibia); and distal groove along dorsal margins of metatarsals I and II.

Etymology— Igai” is the name of the enigmatic “lord of the oasis” deity that was venerated by inhabitants of the Dakhla and Kharga oases (and surrounding regions) in Egypt from roughly the Old Kingdom to the Late Period. The species epithet “semkhu” is the perfect passive singular participle, “the forgotten,” of “semekh,” the ancient Egyptian verb “to forget.” Collectively, “the forgotten lord of the oasis” alludes to both the relatively recent emergence of latest Cretaceous non-marine vertebrate fossils from continental Africa (particularly Egypt) and the lengthy and complicated history of the holotypic specimen (see above).


Eric Gorscak, Matthew C. Lamanna, Daniela Schwarz, Verónica Díez Díaz, Belal S. Salem, Hesham M. Sallam and Marc Filip Wiechmann. 2023. A New titanosaurian (Dinosauria: Sauropoda) from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Egypt. Journal of Vertebrate Paleontology.  e2199810. DOI: 10.1080/02724634.2023.2199810  

Thursday, June 9, 2022

[Paleontology • 2022] First Definitive Record of Abelisauridae (Theropoda: Ceratosauria) from the Cretaceous Bahariya Formation, Bahariya Oasis, Western Desert of Egypt



 Reconstruction of the palaeoecosystem of the Upper Cretaceous (Cenomanian) Bahariya Formation of the Bahariya Oasis, Western Desert of Egypt, showing diversity of large-bodied theropod dinosaurs.

in Salem, Lamanna, O'Connor, ... et Sallam, 2022. 
Artwork by Andrew McAfee, Carnegie Museum of Natural History.

Abstract
Numerous non-avian theropod dinosaur fossils have been reported from the Upper Cretaceous (Cenomanian) Bahariya Formation, Bahariya Oasis, Western Desert of Egypt, but unambiguous materials of Abelisauridae have yet to be documented. Here we report Mansoura University Vertebrate Paleontology Center (MUVP) specimen 477, an isolated, well-preserved tenth cervical vertebra of a medium-sized abelisaurid from the Bahariya Formation. The new vertebra shows affinities with those of other Upper Cretaceous abelisaurids from Madagascar and South America, such as Majungasaurus crenatissimus, Carnotaurus sastrei, Viavenator exxoni and a generically indeterminate Patagonian specimen (Museo Padre Molina specimen 99). Phylogenetic analysis recovers the Bahariya form within Abelisauridae, either in a polytomy of all included abelisaurids (strict consensus tree) or as an early branching member of the otherwise South American clade Brachyrostra (50% majority rule consensus tree). MUVP 477, therefore, represents the first confirmed abelisaurid fossil from the Bahariya Formation and the oldest definitive record of the clade from Egypt and northeastern Africa more generally. The new vertebra demonstrates the wide geographical distribution of Abelisauridae across North Africa during the middle Cretaceous and augments the already extraordinarily diverse large-bodied theropod assemblage of the Bahariya Formation, a record that also includes representatives of Spinosauridae, Carcharodontosauridae and Bahariasauridae.

Keywords: Abelisauridae, Egypt, Africa, Cretaceous, Bahariya Formation


Tenth cervical vertebra of Abelisauridae indet. (MUVP 477)
in cranial (a), caudal (b), left lateral (c), right dorsolateral (d), ventral (e) and dorsal (f) views.


 Reconstruction of the palaeoecosystem of the Upper Cretaceous (Cenomanian) Bahariya Formation of the Bahariya Oasis, Western Desert of Egypt, showing diversity of large-bodied theropod dinosaurs.
In the foreground, the unidentified abelisaurid described herein (right) confronts the spinosaurid Spinosaurus aegyptiacus (left centre, with dipnoan (lungfish) Retodus tuberculatus in jaws) and the carcharodontosaurid Carcharodontosaurus saharicus (right centre) while two individuals of the stomatosuchid crocodyliform Stomatosuchus inermis (left) look on. In the background, a herd of the titanosaurian sauropod Paralititan stromeri (left) warily regards these theropods and two individuals of the bahariasaurid Bahariasaurus ingens (far right) while a small flock of an undescribed pterosaur soars above. The vegetation is dominated by the mangrove-like tree fern Weichselia reticulata.
Artwork by Andrew McAfee, Carnegie Museum of Natural History.

Conclusion: 
The cervical vertebra (MUVP 477) described herein represents the first confirmed fossil of Abelisauridae from the Bahariya Formation, establishing it as the oldest definitive record of this theropod clade from Egypt and northeastern Africa more generally. The new vertebra demonstrates the wide geographical distribution of abelisaurids across North Africa during the middle Cretaceous and augments the already extraordinarily diverse large-bodied non-avian theropod record of the Bahariya Formation, a unit that also preserves representatives of Spinosauridae, Carcharodontosauridae and Bahariasauridae. This abelisaurid/spinosaurid/carcharodontosaurid/bahariasaurid faunal assemblage appears to have extended across most or all of northern Africa during the Cenomanian, suggesting that the Trans-Saharan Seaway did not represent a significant barrier to large-bodied theropod dispersal at this time. The Bahariya Formation holds unrealized potential to improve understanding of this northern African Cenomanian fauna due to the relative commonality of phylogenetically informative associated partial skeletons in this stratigraphic unit.

 
Belal S. Salem, Matthew C. Lamanna, Patrick M. O'Connor, Gamal M. El-Qot, Fatma Shaker, Wael A. Thabet, Sanaa El-Sayed and Hesham M. Sallam. 2022. First Definitive Record of Abelisauridae (Theropoda: Ceratosauria) from the Cretaceous Bahariya Formation, Bahariya Oasis, Western Desert of Egypt. R. Soc. open sci. 9220106. DOI: 10.1098/rsos.220106 

Hesham Sallam Leads Egyptian Team’s Discovery of Bizarre Meat-Eating Dinosaur in Egypt’s Bahariya Oasis 

Thursday, August 26, 2021

[PaleoMammalogy • 2021] Phiomicetus anubis • A New Protocetid Whale offers clues to Biogeography and Feeding Ecology in early Cetacean Evolution


Phiomicetus anubis 
Gohar, Antar, Boessenecker, Sabry, El-Sayed, Seiffert, Zalmout & Sallam, 2021

 
Abstract
Over about 10 million years, the ancestors of whales transformed from herbivorous, deer-like, terrestrial mammals into carnivorous and fully aquatic cetaceans. Protocetids are Eocene whales that represent a unique semiaquatic stage in that dramatic evolutionary transformation. Here, we report on a new medium-sized protocetid, Phiomicetus anubis gen. et sp. nov., consisting of a partial skeleton from the middle Eocene (Lutetian) of the Fayum Depression in Egypt. The new species differs from other protocetids in having large, elongated temporal fossae, anteriorly placed pterygoids, elongated parietals, an unfused mandibular symphysis that terminates at the level of P3, and a relatively enlarged I3. Unique features of the skull and mandible suggest a capacity for more efficient oral mechanical processing than the typical protocetid condition, thereby allowing for a strong raptorial feeding style. Phylogenetic analysis nests Phiomicetus within the paraphyletic Protocetidae, as the most basal protocetid known from Africa. Recovery of Phiomicetus from the same bed that yielded the remingtonocetid Rayanistes afer provides the first clear evidence for the co-occurrence of the basal cetacean families Remingtonocetidae and Protocetidae in Africa. The discovery of Phiomicetus further augments our understanding of the biogeography and feeding ecology of early whales.

Keywords: biogeography, whales, phylogeny, evolution, Fayum 




  Phiomicetus anubis gen. et sp. nov.

 




Abdullah S. Gohar, Mohammed S. Antar, Robert W. Boessenecker, Dalia A. Sabry, Sanaa El-Sayed, Erik R. Seiffert, Iyad S. Zalmout and Hesham M. Sallam. 2021. A New Protocetid Whale offers clues to Biogeography and Feeding Ecology in early Cetacean Evolution. Proc. R. Soc. B.. 288. 20211368. DOI: 10.1098/rspb.2021.1368