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| Pristimantis etsa Figueroa-Coronel, Cisneros-Heredia, Brito-Zapata, Carrión-Olmedo & Reyes-Puig, 2026 |
Abstract
A new species of Pristimantis is described from the Cordillera del Cóndor, Zamora Chinchipe Province, southeastern Ecuador, based on morphological and molecular data, Pristimantis etsa sp. nov. The new species is diagnosed from its congeners by the following combination of characters: female SVL 32.5 mm and male SVL 18.3 mm, dorsolateral folds formed by rows of subconical tubercles, strongly areolate ventral skin, two distinct rows of forearm tubercles, one along the ventrolateral margin and a second along the externolateral margin of the forearm, and a prominent yellow groin blotch in the female holotype. The species belongs to the Pristimantis cryptomelas group, part of the Huicundomantis subgenus, and is closely related to P. nangaritza, P. verrucosus, and P. plateado. Currently, the species is known only from its type locality, where it inhabits low montane evergreen forests at elevations of 1,655–1,830 m. Additionally, we discuss the use of the term “ulnar tubercles” in Pristimantis, noting that it may refer to tubercles occupying different positions on the forearm.
Key words: Amphibia, forearm tubercles, Huicundomantis, sub-Andean cordilleras
Pristimantis etsa sp. nov.
Proposed standard English name. Etsa Rain Frog.
Proposed standard Spanish name. Cutín de Etsa.
Generic placement. The new species is assigned to Pristimantis based on the presence of a differentiated tympanic membrane, S-shaped adductor muscles, and expanded terminal discs on digits bearing well-defined circumferential grooves (Hedges et al. 2008).
Diagnosis. The diagnosis is summarized in Table 1, and a visual comparison of P. etsa sp. nov. and P. nangaritza is shown in Fig. 8. The species included in the comparative diagnosis were selected because they are phylogenetically close to Pristimantis etsa sp. nov. within the P. cryptomelas group or because they share one or more externally similar characters, especially colored groin, ulnar tuberculation, and occurrence in the Cordillera del Cóndor, southern Ecuador, or adjacent northern Peru. Pristimantis etsa sp. nov. is distinguished from these congeners by the following combination of characters: snout rounded in dorsal view and profile; dorsolateral folds conspicuous, composed of subconical ...
Etymology. The specific epithet etsa is a noun in apposition derived from the Shuar language. Among the Shuar people, an indigenous nationality inhabiting eastern Ecuador and northern Peru, including parts of the Cordillera del Cóndor, Etsa is a powerful anthropomorphic being whose primary manifestation is the sun. In Shuar cosmology, Etsa acts as a cultural transmitter who endows animals and people with essential skills, such as hunting techniques, restores life to forest birds, and upholds moral order (Pellizaro 1984; Barrueco 1985).
Elías Figueroa-Coronel, Diego F. Cisneros-Heredia, David Brito-Zapata, Julio C. Carrión-Olmedo and Carolina Reyes-Puig. 2026. A New rain frog of the Genus Pristimantis (Anura, Craugastoridae) from the Cordillera del Cóndor, southeastern Ecuador. ZooKeys. 1282: 205-228. DOI: doi.org/10.3897/zookeys.1282.187506 [15 Jun 2026]











































