[Botany • 2026] Primulina taishanensis (Gesneriaceae) • A New Species from Guangdong, China

 

Primulina taishanensis F.Wen & Yu Q.Chen, 

in Qian, Xin, Chen, Huang, Bai et Wen, 2026. 

台山报春苣苔  ||  DOI: doi.org/10.1007/s12225-025-10358-w

Summary

Primulina taishanensis F.Wen & Yu Q.Chen, a new species of Gesneriaceae, from Taishan County, Guangdong Province, China, is described. It is morphologically similar to P. dryas (Dunn) Mich.Möller & A.Weber in the cuneate leaf base, bract size, purple pubescent peduncle, pistil size and white glandular hairs on the ovary. It is clearly differentiated from that species by its acuminate leaf blade apex (vs obtuse in P. dryas); pedicels 8 – 25 mm long and purple glandular pilose (vs 30 – 50 mm long, densely white villous) and anthers that are reniform and deeply constricted at the middle, c. 2.5 mm long (vs fusiform, c. 4 mm long). A provisional conservation category of Vulnerable [VU D1], according to the IUCN Red List Categories and Criteria is proposed.

Key Words: IUCN, morphology, new taxon, taxonomy.

   

Primulina taishanensis F.Wen & Yu Q.Chen sp. nov. 

Intraspecific flower colour variation in Primulina taishanensis.
Top row, front views; middle row, lateral view; bottom row, lateral view with corollas opened.

Etymology. “taishanensis” is Latinised from the pinyin and the suffix “-ensis” conveys “from the City of Taishan in Guangdong Province”, where the species was found.

Vernacular name. Tái Shān Bào Chūn Jù Tái (Chinese pronunciation); 台山报春苣苔 (Chinese).

Xiao-Mao Qian, Zi-Bing Xin, Yu-Qiang Chen, Guan-Yun Huang, Xin-Xiang Bai and Fang Wen. 2026. Primulina taishanensis (Gesneriaceae), A New Species from Guangdong, China.  Kew Bulletin. DOI: doi.org/10.1007/s12225-025-10358-w [23 February 2026]

[Herpetology • 2021] Hemidactylus cinganji, H. faustus, ... • Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820 (Squamata, Gekkonidae), with Description of Four New Species

  Hemidactylus cinganji
H. carivoensis
H. faustus 
H. pfindaensis 

 Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Vaz Pinto, 2021 

       

 DOI: 10.3897/vz.71.e64781 

 facebook.com/JavierLobonRovira

 

Abstract

The taxonomy of Angolan Hemidactylus has recently been revised. However, the lack of fresh material for some groups and regions, has led to the misidentification of some taxa and an underestimation of actual diversity in others. To shed light on the evolutionary history and systematics of Angolan Hemidactylus, we generated a new phylogenetic hypothesis for the group, and updated the taxonomy following an integrative approach. This resulted in the description of four new species (Hemidactylus pfindaensis sp. nov., H. faustus sp. nov., H. carivoensis sp. nov. and H. cinganji sp. nov.), the reevaluation of two recently described species (H. vernayi and H. paivae) and the synonymization of a recently described species (H. hannahsabinnae). We estimate divergence times for these lineages, providing a preliminary interpretation of their speciation process. Moreover, we suggest and outline 13 Angolan Main Biogeographic Units (AMBUs) in the area, defining a new biogeographic context for future works on Angolan herpetofauna. We now recognize eleven Hemidactylus species in Angola, and we provide here a new morphological key for Angolan Hemidactylus to help with identifications and species assignments in this group.

Keywords: Angola, biogeography, Gekkota, Hemidactylus, systematics, taxonomy

Hemidactylus bayonii-group 

Hemidactylus vernayi Ceríaco, Agarwal, Marques and Bauer 2020

Hemidactylus pfindaensis sp. nov.

Etymology: The name “pfindaensis” derives from the local word “pfinda” which in Kikongo – the main language used in Uíge Province and northwestern Angola – refers to a “gallery forest” or a “continuous block of thick forest”, the main habitat type associated with the species.

Hemidactylus longicephalus-group

Hemidactylus paivae Ceríaco, Agarwal, Marques and Bauer 2020

A – From top, dorsolateral view of Hemidactylus cinganji sp. nov. (FKH0435), H. benguellensis (FKH0413) and H. carivoensis sp. nov. (MNCN 50543);
B – records of H. benguellensis–group (purple color represents records of H. cinganji sp. nov.; blue H. benguellensis; dark green H. carivoensis sp. nov.; circles represent genetically assigned records, while triangles represent historical or non-genetically assigned records by Ceríaco et al. 2020a; color stars depict type or neotype localities; black star depicts original type locality of H. benguellensis assigned by Bocage (1893);
C – habitat of H. cinganji sp. nov. at Ebanga; D – habitat of H. carivoensis sp. nov between Dombe–Echimina. E – habitat of H. benguellensis at Tchivinguiro.

 Photos Javier Lobón–Rovira (A–D) and William R. Branch (E).

Hemidactylus benguellensis-group

Hemidactylus cinganji sp. nov.

Etymology: The name “cinganji” is a widespread traditional word used in Angolan local languages that represents an ancestral spiritual entity that reincarnates assuming different physical forms in different places and occasions. This name is suitable as the new species corresponds to a taxon that was first described under a different name, then became lost and now resurfaces after its original name had been hijacked by a surrogate sister-species. The species epithet is used as a neuter singular noun in opposition to the generic name.

Hemidactylus carivoensis sp. nov.

Etymology: The species epithet “carivoensis” refers to the Farm Carivo, an old estate situated along the banks of the mid-lower Coporolo River on the coastal plain of Benguela Province, and where most of the type series was collected. The species proved to be common in the area, and by recognizing the farm, we also acknowledge the ongoing support from the owners to researchers, similar to the Chapmans nearly a century ago.

Species not assignable to specific species groups

Hemidactylus faustus sp. nov.   

  

Etymology: The name “faustus” applies to a Latin word that designate ‘good luck’, evoking the serendipitous discovery of this species. The species epithet is used as a masculine adjective singular. The first specimen was found by Beatriz Vaz Pinto, daughter of PVP, under a small stone which was removed while preparing a campsite. This unexpected find led to further collecting of this new and previously unrecorded form, albeit from a locality that had been regularly surveyed since the mid-19th century.  

 

 Javier Lobón-Rovira, Werner Conradie, David Buckley Iglesias, Raffael Ernst, Luis Veríssimo, Ninda Baptista and Pedro Vaz Pinto. 2021. Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820, with Description of Four New Species. Vertebrate Zoology. 71: 465-501. DOI: 10.3897/vz.71.e64781

  facebook.com/JavierLobonRovira/posts/3969307526513233

[Ichthyology • 2026] Oreichthys elianae & Glyptothorax sentimereni • New Species (Teleostei: Sisoridae, Cyprinidae) from Nagaland, Northeast India

 

Oreichthys elianae  &

Glyptothorax sentimereni

 Praveenraj & Vijayakrishnan, 2026

DOI: doi.org/10.11646/zootaxa.5763.2.5

Abstract

Two new species of fishes are described from the tributaries of the Brahmaputra River in Nagaland, Northeast India. Glyptothorax sentimereni, described from the Dikhu River, is distinguished from congeners in having the following combination of characters: ventral surfaces of pectoral spine and first pelvic-fin ray with plicae, serrated posterior edge of dorsal-fin spine, thoracic adhesive apparatus completely encinctured by striae, length of dorsal-fin spine 13.7–16.1% SL, body depth at anus 17.1–19.6% SL, length of dorsal-fin base 12.0–13.1% SL, post-adipose distance 20.8–22.2% SL, head depth 16.5–17.0% SL and dorsal-to-adipose distance 21.8–23.5% SL. Oreichthys elianae, described from the Tsurang River, is distinguished from its congeners by a combination of characters that include 19+1 scales (with 5 pored scales) in longitudinal series, ½3/1/2½ scales in transverse row, 9 predorsal scales, 14–15 pores on cheek, a black spot on the caudal-fin base, and reddish fins.

Pisces, Brahmaputra River, Cypriniformes, Diversity, endemic fish, Siluriformes

 

Glyptothorax sentimereni, a rheophilic torrent catfish from the Dikhu River, Nagaland

Oreichthys elianae, from the Tsurang River in Mokokchung, loses its coloration immediately under stress

Jayasimhan PRAVEENRAJ and Balaji VIJAYAKRISHNAN. 2026. New Species of the Genera Glyptothorax (Teleostei: Sisoridae) and Oreichthys (Teleostei: Cyprinidae) from Nagaland, Northeast India.  Zootaxa. 5763(2); 255-267. DOI: doi.org/10.11646/zootaxa.5763.2.5 [2026-03-03]

[Botany • 2026] Sadiria kachinensis (Primulaceae: Myrsinoideae) • A New Species from Myanmar

Sadiria kachinensis Utteridge & Nob.Tanaka,

in Utteridge, Armstrong, Yoneda et Tanaka. 2026.  

DOI: doi.org/10.11646/phytotaxa.744.3.1

 

Abstract

Sadiria kachinensis from Kachin State, Myanmar is described and illustrated as a species new to science. It is morphologically similar to S. boweri and S. solanifolia but differs in several characters especially the smaller leaves on the vegetative shoots and the flowering branches, larger inflorescences and petals, and smaller sepals. In addition, this new species is compared to S. eugeniifolia var. burmanica, the only other member of Sadiria in Myanmar.

 Burma, Ericales, Indo-China, South-East Asia, taxonomy, Eudicots

Sadiria kachinensis Utteridge & Nob.Tanaka.
 A habit; B detail of leaf margin; C inflorescences; D flower; E single calyx lobe; F detail of overlapping corolla lobes; G flower opened to show stamens opposite the corolla lobes; H ovary and style (flower with corolla removed); I stamen abaxial view; J stamen adaxial view; K ovary; L fruit with persistent style.

Drawn from Armstrong et al. 4307 (TNS). Scale bars: A = 3 cm, C, D, G, H, L = 5 mm, E & F = 1 mm, I, J & K = 3 mm. Illustration by Kaoru Yoneda.

Sadiria kachinensis Utteridge & Nob.Tanaka.
A. habit; B. plagiotropic (flowering) branch with inflorescences and flowers pre-anthesis; C. plagiotropic branch with fruits; D. inflorescence/flower detail, note exserted styles in the flowers on the right hand side; E. fruit detail, note persistent style (dry and black).

 Photographed by Kate Armstrong.

Sadiria kachinensis Utteridge & Nob.Tanaka, sp. nov. 

 Recognised in the genus Sadiria by the leaves (ortho- and plagiotropic) drying greenish and the shorter plagiotropic reproductive branches, and unique amongst those species in the combination of the following characters: 3–4 leaves in pseudowhorls along the erect (orthotropic) stems, these orthotropic leaves, elliptic and 9.5–13 × 3.5–4.5 cm; the lateral plagiotropic (reproductive) flower-bearing branches (4–)6.5–11 cm long, with 3(–5) leaves along the length of the branch; orthotropic and reproductive shoots leaves drying green; pendulous inflorescences sessile or with a very short rachis to 2 mm long; the corolla fused for approximately three-quarters of its length; stamens with anthers 2 mm long; and the style conspicuously exserted at anthesis and extending ca. 3 mm beyond the apex of the corolla lobes.

Timothy UTTERIDGE, Kate E. ARMSTRONG, Kaoru YONEDA and Nobuyuki TANAKA. 2026. Sadiria kachinensis (Myrsinoideae, Primulaceae), A New Species from Myanmar.  Phytotaxa. 744(3); 205-211. DOI: doi.org/10.11646/phytotaxa.744.3.1 [2026-03-10]

[Mollusca • 2025] Chamalycaeus aduncus • The First Record of an Operculate Micro Land Snail from the Alycaeinae (Caenogastropoda: Cyclophoridae) in Cambodia

 

Chamalycaeus aduncus Jirapatrasilp,   

in Jirapatrasilp, Thi, Chantha, Chourn, Sophea, Jeratthitikul, Pholyotha, Siriwut, Srisonchai, Panha et Sutcharit, 2025. 

www.ruthenica.net/node/6016

 Researchgate.net/publication/390396808 

 facebook.com/ASRUChula

 

This study reports the first record of the operculate micro land snail from the Alycaeinae, Chamalycaeus aduncus Jirapatrasilp sp. nov., also the first new species of the genus in Cambodia, based on a recent survey in Battambang. This new species is similar to Chamalycaeus excisus (Möllendorff, 1887) and C. excisus sublimus Páll-Gergely et Auffenberg, 2019 from the Philippines, in shell size and shape, and the strongly downwards coiling of R3, but differs in having a shorter R2 without any blunt keel, a round aperture without any incision, and a beak- or hook-like basal edge of the outer peristome. 

Keywords: Battambang; Cyclophoroidea; Gastropoda; Mollusca; taxonomy

Chamalycaeus aduncus sp. nov.
A paratype CUMZ 14462/1, and B paratype CUMZ 14462/2, showing a scraping hole at the umbilical side.

. Representative SEM images of Chamalycaeus aduncus sp. nov., holotype CUMZ 14461
 A whole shell in different positions, and B Close-up images of the aperture; white arrows pointing to the beak- or hook-like basal edge of the outer peristome.

Chamalycaeus aduncus Jirapatrasilp sp. nov.

 

Parin JIRAPATRASILP, Sothearen THI, Nasak CHANTHA, Phyroum CHOURN, Chhin SOPHEA, Ekgachai JERATTHITIKUL, Arthit PHOLYOTHA, Warut SIRIWUT, Ruttapon SRISONCHAI, Somsak PANHA and Chirasak SUTCHARIT. 2025. The First Record of an Operculate Micro Land Snail from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Cambodia, with Description of A New Species of Chamalycaeus von Möllendorff, 1897. Ruthenica Russian Malacological Journal. 35(2); 65-72. https://www.ruthenica.net/node/6016

 Researchgate.net/publication/390396808_The_first_record_of_an_operculate_micro_land_snail_from_the_Alycaeinae_Cyclophoridae_in_Cambodia

 facebook.com/ASRUChula/posts/1081408774003851

[Entomology • 2026] Methocha haaksek, M. hongkongensis, ... • The Methocha (Hymenoptera: Thynnidae: Methochinae) of Hong Kong (China), A Preliminary Faunal List with Descriptions of Five New Species and Sexual Associations rendered through Molecular Analysis

 

A. Methocha haaksek Barthélémy & Terayama sp. nov. 
B & C. M. hongkongensis Barthélémy & Terayama sp. nov. 
 D. M. leleji Barthélémy & Terayama sp. nov. 
 E. M. wilsoni Barthélémy & Terayama sp. nov. 
 F. M. zetetes Barthélémy & Terayama sp. nov. 

in Barthélémy, Terayama, Poon et Sin. 2026. 

DOI: doi.org/10.5852/ejt.2026.1043.3213 

Abstract

Five new species of Methocha Latreille, 1804 are described from Hong Kong: two species based on females only, Methocha haaksek Barthélémy & Terayama sp. nov. and Methocha leleji Barthélémy & Terayama sp. nov.; two on males only, M. wilsoni Barthélémy & Terayama sp. nov. and M. zetetes Barthélémy & Terayama sp. nov.; and one with females and males associated through molecular analysis, M. hongkongensis Barthélémy & Terayama sp. nov. A key to species and a phylogenetic tree of Methocha occurring in the Hong Kong SAR are provided.

Keywords: Thynnoidea, morphometrics, Cicindelinae, DNA barcoding, phylogenetic

Holotypes and paratypes, lateral habitus.
A. Methocha haaksek Barthélémy & Terayama sp. nov., holotype, ♀, M459.D.Hy.1, CASC. B. M. hongkongensis Barthélémy & Terayama sp. nov., holotype, ♀, M364.C.Hy.2, CASC. C. M. hongkongensis, paratype, ♂, M561CHY3, CBC. D. ♀, M. leleji Barthélémy & Terayama sp. nov., holotype, ♂, M099.C.Hy.2, CASC.
E. M. wilsoni Barthélémy & Terayama sp. nov., holotype, ♂, M060.C.Hy.4, CASC. F. M. zetetes Barthélémy & Terayama sp. nov., paratype, ♂, M321.E.Hy.4B, CBC.

Methocha haaksek Barthélémy & Terayama sp. nov. 

M. hongkongensis Barthélémy & Terayama sp. nov. 

M. leleji Barthélémy & Terayama sp. nov.; 

M. wilsoni Barthélémy & Terayama sp. nov. 

M. zetetes Barthélémy & Terayama sp. nov.

 Christophe Barthélémy, Mamoru Terayama, Emily Shui Kei Poon and Simon Yung Wa Sin. 2026. The Methocha (Hymenoptera: Thynnidae: Methochinae) of Hong Kong (China), A Preliminary Faunal List with Descriptions of Five New Species and Sexual Associations rendered through Molecular Analysis. European Journal of Taxonomy. 1043(1); 118–151. DOI: doi.org/10.5852/ejt.2026.1043.3213 

[Invertebrate • 2026] Dalhousiella yabukii & Leocratides watanabeae • Single Origin and Convergent Host Use of hexactinellid Sponge Symbiosis in Hesionidae (Annelida: Polychaeta) with Descriptions of Two New Deep-sea Species

Dalhousiella yabukii and Leocratides watanabeae 

 Jimi, Downey, Mella & Hookabe, 2026

 DOI: doi.org/10.1093/zoolinnean/zlag028  

 x.com/nu_mbl

Abstract

Symbiotic associations between annelids and sponges are widespread and have evolved repeatedly across diverse families. However, their evolutionary history within Hesionidae remains poorly understood. Using the manned submersible Shinkai 6500, we conducted in situ observations and collected hesionid specimens from a hexactinellid sponge on the Getsuyo Seamount, north-west Pacific Ocean. An integrative taxonomic approach combining morphology and molecular phylogenetics revealed that they belong to two new species, Dalhousiella yabukii sp. nov. and Leocratides watanabeae sp. nov. Phylogenetic analyses suggest that sponge symbiosis in Hesionidae originated once, in the common ancestor of Dalhousiella and Leocratides. Nevertheless, the two new species are not each other’s closest relatives; instead, they form independent sister-clades, each including different congeners. Specimens of the two new species inhabited the same sponge host and probably shared similar ecological niches, suggesting convergent ecological specialization toward the limited resource of hexactinellid sponges in the deep sea. Such niche overlap and convergent evolution between closely related symbiotic species have not been previously reported in Hesionidae. Our findings shed new light on the origin and diversification of sponge symbiosis in Hesionidae and highlight the importance of integrative taxonomy and direct in situ observation for understanding deep-sea biodiversity.

Hesionini, taxonomy, symbiosis, Polychaeta, seamount

Dalhousiella yabukii sp. nov.  

Leocratides watanabeae sp. nov. 

Naoto Jimi, Rachel Downey, María Belén Arias Mella and Natsumi Hookabe. 2026. Single Origin and Convergent Host Use of hexactinellid Sponge Symbiosis in Hesionidae (Annelida: Polychaeta) with Descriptions of Two New Deep-sea Species. Zoological Journal of the Linnean Society. 206(3); zlag028. DOI: doi.org/10.1093/zoolinnean/zlag028 [09 March 2026] 

 x.com/nu_mbl/status/2030905993211810299

[Herpetology • 2026] Ololygon paracatu • A New Species of the Ololygon catharinae group (Anura: Hylidae: Scinaxini) from the Brazilian Cerrado

Ololygon paracatu 

 Carvalho, Valencia-Zuleta, Araujo-Vieira, Faivovich, Maciel & Brandão,  2026

DOI: doi.org/10.11646/zootaxa.5757.6.2

 

Abstract

We describe a new species of the Ololygon catharinae group from the Cerrado biome, in the state of Minas Gerais, Brazil. Ololygon paracatu sp. nov. is morphologically similar to O. goya and O. skaios, although in phylogenetic analyses, it is recovered as the poorly supported sister taxon of O. pombali. It is distinguished from other species of the O. catharinae group by having a canthus rostralis marked and curved; subovoid snout in dorsal view and protruding in profile; inverted triangle shaped interocular blotch, exceeding the posterior margin of the eyes; inguinal region and hidden areas of thighs with dark brown irregular blotches on a pale yellow background in life; and advertisement call composed of 3–5 pulsed notes and dominant frequency of 2.5–3.5 kHz. Ololygon paracatu sp. nov. inhabits gallery forests associated with streams in the Cerrado biome.

Amphibia, Hylinae, taxonomy, phylogeny, calls, gallery forest

 

 

Ololygon paracatu sp. nov.

Daniele CARVALHO, Alejandro VALENCIA-ZULETA, Katyuscia ARAUJO-VIEIRA, Julián FAIVOVICH, Natan M. MACIEL and Reuber A. BRANDÃO. 2026. A New Species of the Ololygon catharinae group from the Brazilian Cerrado (Anura, Hylidae, Scinaxini). Zootaxa. 5757(6); 522-542. DOI: doi.org/10.11646/zootaxa.5757.6.2 [2026-02-11]

[Mollusca • 2026] Operculate Land Snails (Gastropoda: Caenogastropoda: Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the Description of A New Species, Chamalycaeus dayangmerindu

 

c. Chamalycaeus dayangmerindu Aulia & Nurinsiyah, sp. nov.;
d. C. korintjiensis (Páll-Gergely, 2020), holotype (ZMA.MOLL 135672).
a. Stomacosmethis cf. jagori (E. von Martens, 1860); b. S. jagori (E. von Martens, 1860), syntype (SMF 109304); 

in Aulia, Nurinsiyah, Mujiono, Páll-Gergely et Ambarwati, 2026.  

 DOI: doi.org/10.3897/zookeys.1272.179378

Abstract

The study on Cyclophoroidea from Padang Bindu Karst, South Sumatra, was conducted to document the species diversity of the superfamily in the area. The samples, including leaf litter and soil samples, were collected in May–June 2021 and followed by the determination and examination on 2023 to 2024 in the Museum Zoologicum Bogoriense. In total 3,780 specimens from the superfamily Cyclophoroidea were examined. Measurements of the shell and operculum were performed using L.A.S V4. 13.0 and IMAGE J. The research revealed 11 species from three families (Cyclophoridae, Diplomatinidae, Pupinidae) and four subfamilies. Plectostoma kitteli is the most abundant species followed by Stomacosmethis cf. jagori (19.84%) and Diplommatina liwaensis (6.67%). A new species, Chamalycaeus dayangmerindu Aulia & Nurinsiyah, sp. nov. is described. The study also discovered four species endemics to Sumatra with one species among them so far only recorded from Padang Bindu karst area. These findings emphasize the region’s unique biodiversity.

Key words: Biodiversity, conservation, limestone, molluscs, operculum

Shells of Alycaeinae. a. Stomacosmethis cf. jagori (E. von Martens, 1860); b. Stomacosmethis jagori (E. von Martens, 1860), syntype (SMF 109304);
c. Chamalycaeus dayangmerindu sp. nov.; d. Chamalycaeus korintjiensis (Páll-Gergely, 2020), holotype (ZMA.MOLL 135672).

Chamalycaeus dayangmerindu Aulia & Nurinsiyah, sp. nov.

Habitat. Limestone rocks.

Distribution. The species is so far only recorded from Padang Bindu Karst, South Sumatra.

Etymology. The species name Dayang Merindu was inspired by Princess Dayang Merindu, a figure from local folklore. According to the legend, Princess Dayang Merindu ignored the greeting of a man, as she was already married. Her action was then perceived as arrogance, and the man cursed her into stone. The stone associated with this tale is located inside Gua Putri, a karst cave formation in the Padang Bindu area.

 Latifah Nurul Aulia, Ayu Savitri Nurinsiyah, Nova Mujiono, Barna Páll-Gergely and Reni Ambarwati. 2026. Operculate Land Snails (Gastropoda, Caenogastropoda, Cyclophoroidea) from Padang Bindu Karst, South Sumatra, Indonesia with the Description of A New Species, Chamalycaeus dayangmerindu. ZooKeys. 1272: 1-31. DOI: doi.org/10.3897/zookeys.1272.179378 [02 Mar 2026]

[Paleontology • 2026] Sonselasuchus cedrus • Osteology and Relationships of A New shuvosaurid (Pseudosuchia: Poposauroidea: Shuvosauridae) from the Upper Triassic Chinle Formation of Petrified Forest National Park, Arizona, U.S.A

Sonselasuchus cedrus

Smith & Sidor, 2026

DOI: doi.org/10.1080/02724634.2025.2604859

Artwork by Gabriel Ugueto

ABSTRACT

Shuvosauridae is a clade of pseudosuchian archosaurs currently represented by three named species characterized by a body plan strikingly convergent with that of ornithomimid theropod dinosaurs. This paper documents a new genus and species of shuvosaurid from the Upper Triassic Chinle Formation of Petrified Forest National Park, Arizona. Sonselasuchus cedrus, gen. et sp. nov., is largely diagnosed by features of the maxilla, including a reduced body and anterior process, an enlarged subnarial foramen, and an expanded facet on the posterior process, although other unique cranial features are also noted herein. By contrast, its postcranial anatomy is largely similar to what has been described for other shuvosaurids, although subtle differences are noted. A phylogenetic analysis finds S. cedrus in an unresolved clade with Effigia okeeffeae and Shuvosaurus inexpectatus, likely as a result of non-overlapping missing data, particularly for the skull of Shuvosaurus. A review of the shuvosaurid fossil record indicates that shuvosaurids were persistent components of Late Triassic terrestrial vertebrate faunas in North America. S. cedrus is represented by a minimum number of 36 individuals, mostly skeletally immature, that occur within a multitaxic bonebed of almost exclusively disarticulated elements. An analysis of the relative size change of limb dimensions indicates the forelimb was growing on a negative allometric trajectory relative to the hindlimb, which is consistent with a transition in locomotory mode from quadrupedal to bipedal in S. cedrus during ontogeny.

Life restoration of Sonselasuchus cedrus as depicted by Gabriel Ugueto. Note the occurrence of individuals in both bipedal and quadrupedal stances.

SYSTEMATIC PALEONTOLOGY

ARCHOSAURIA Cope, Citation1869

PSEUDOSUCHIA Zittel, Citation1887–Citation1890

PARACROCODYLOMORPHA Parrish, Citation1993

POPOSAUROIDEA Nopcsa, Citation1923

SHUVOSAURIDAE Chatterjee, Citation1993

SONSELASUCHUS CEDRUS gen. et sp. nov.

Etymology—For the genus, ‘Sonsela’ refers to the Sonsela Member of the Chinle Formation, the type section is located at the Sonsela Buttes on the Navajo Nation; ‘suchus’ is a Latinization of the Greek word soukhos, for the Egyptian crocodile deity Sobek, reflecting the pseudosuchian affinities of the taxon. The specific epithet, ‘cedrus’ is the genus name for the common cedar (actually the juniper, Juniperus spp.) and refers to Cedar Tank, the geographic feature in Petrified Forest National Park near the type locality.

Holotype—PEFO 47305/UWBM 119436, left maxilla.

Locality and Horizon—The Kaye Quarry (PFV 410/UWBM C2226) is located west of Cedar Tank at Petrified Forest National Park, Apache County, Arizona, U.S.A. (Fig. 1). The site is within the Jim Camp Wash beds of the Sonsela Member of the Chinle Formation and is Norian in age (Marsh et al., 2024). ...

Elliott Armour Smith and Christian A. Sidor. 2026. Osteology and Relationships of A New shuvosaurid (Pseudosuchia, Poposauroidea) from the Upper Triassic Chinle Formation of Petrified Forest National Park, Arizona, U.S.A. Journal of Vertebrate Paleontology. e2604859. DOI: doi.org/10.1080/02724634.2025.2604859 [08 Mar 2026]

[Diplopoda • 2026] Rhopalomeris dulcia, R. lentiginosa, R. verhoeffi, ... • Five New Species of the Pill Millipede Genus Rhopalomeris Verhoeff, 1906 (Glomerida: Glomeridae) from Thailand

Rhopalomeris dulcia, 
R. verhoeffi  

Sapparojpattana & Likhitrakarn,

in Sapparojpattana, Jeratthitikul, Siriwut, Srisonchai, Wesener, Sutcharit et Likhitrakarn, 2026.  

DOI: doi.org/10.1163/18759866-bja10090 

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Abstract

An integrative approach based on morphological characteristics and DNA data from the mitochondrial COI gene has revealed five new species of the pill millipede genus Rhopalomeris Verhoeff, 1906 from Thailand: Rhopalomeris muka Sapparojpattana & Likhitrakarn, sp. nov., R. lentiginosa Sapparojpattana & Likhitrakarn, sp. nov., R. dulcia Sapparojpattana & Likhitrakarn, sp. nov., R. punctata Sapparojpattana & Likhitrakarn, sp. nov., and R. verhoeffi Sapparojpattana & Likhitrakarn, sp. nov. The five new species are distinguishable from their congeners by distinctive color patterns and telopod structure. The interspecific genetic divergences between the new species and other Rhopalomeris species ranged from 7.86 to 13.71%. Intraspecific genetic divergences within these five species ranged from 0 to 4.53%, with the highest divergences found in R. carnifex and R. verhoeffi sp. nov. This is consistent with both the wide distribution range and the significant intraspecific morphological variations observed in these latter two species. Furthermore, the taxonomic scope of R. carnifex in Thailand is reviewed, and intraspecific morphological variations within R. carnifex and R. verhoeffi sp. nov. are discussed. A distribution map, morphological illustrations from SEM, and an updated key to all known Rhopalomeris species are also provided.

Keywords: DNA barcoding; Indochina; integrative taxonomy; phylogeny; systematics

Rhopalomeris dulcia sp. nov., (A, B) paratype (MUMNH-GLO188), habitus, live coloration (C–E) ♀ paratype (MUMNH-GLO188), alcohol preserved, in dorsal, ventral, and lateral views (A) unrolled in sublateral view (B) rolled in sublateral view.

(A, B) image not to scale (C–E) scale bars: 2 mm

Rhopalomeris verhoeffi sp. nov., (A, B) paratype (MUMNH-GLO201), habitus, live coloration of color morph 2 (C–E) ♀ paratype (MUMNH-GLO201), alcohol preserved specimen of color morph 2 in dorsal, ventral, and lateral views (A) unrolled in sublateral view (B) rolled in sublateral view.

(A, B) image not to scale (C–E) scale bars: 2 mm

 Rhopalomeris muka Sapparojpattana & Likhitrakarn, sp. nov.

R. lentiginosa Sapparojpattana & Likhitrakarn, sp. nov. 

 R. dulcia Sapparojpattana & Likhitrakarn, sp. nov. 

 R. punctata Sapparojpattana & Likhitrakarn, sp. nov. 

 R. verhoeffi Sapparojpattana & Likhitrakarn, sp. nov. 

Distributions of currently known Rhopalomeris species in Thailand and Myanmar

(A) Bayesian inference tree focusing on the detailed topology of Rhopalomeris carnifex (Pocock, 1889) and R. verhoeffi sp. nov., with colored drawings illustrating the color variations within each species. All millipede drawings are not to scale.
(B, C) Maps of southern Thailand showing the distributions of color morphs for (B) R. carnifex and (C) R. verhoeffi sp. nov. The different shaded colors in each pie chart represent the proportion of the corresponding color morphs in that locality.

 Pichsinee Sapparojpattana, Ekgachai Jeratthitikul, Warut Siriwut, Ruttapon Srisonchai, Thomas Wesener, Chirasak Sutcharit, and Natdanai Likhitrakarn. 2026. Five New Species of the Pill Millipede Genus Rhopalomeris Verhoeff, 1906 (Diplopoda, Glomerida) from Thailand. Contributions to Zoology. 95(1); 52–102. DOI: doi.org/10.1163/18759866-bja10090 [20 Feb 2026]

 facebook.com/ASRUChula/posts/1351095590368500

***งานวิจัยใหม่: กิ้งกือกระสุนลูกกวาด 5 ชนิดใหม่ของโลกจากประเทศไทย
และการมีรูปแบบสีที่หลากหลายซึ่งพบในบางชนิด***

...

รายชื่อชนิดใหม่ดังนี้

.

1. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙢𝙪𝙠𝙖 Sapparojpattana & Likhitrakarn, 2026
กิ้งกือกระสุนลูกกวาดมหิดลกาญจน์
 ค้นพบที่ มหาวิทยาลัยมหิดล วิทยาเขตกาญจนบุรี อำเภอไทรโยค จังหวัดกาญจนบุรี

2. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙡𝙚𝙣𝙩𝙞𝙜𝙞𝙣𝙤𝙨𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลท่าขนุน อำเภอทองผาภูมิ จังหวัดกาญจนบุรี

3. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙙𝙪𝙡𝙘𝙞𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลบ้านนา อำเภอเมืองชุมพร จังหวัดชุมพร

4. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙥𝙪𝙣𝙘𝙩𝙖𝙩𝙖 Sapparojpattana & Likhitrakarn, 2026
ค้นพบที่ ตำบลไร่เก่า อำเภอสามร้อยยอด จังหวัดประจวบคีรีขันธ์

5. 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨 𝙫𝙚𝙧𝙝𝙤𝙚𝙛𝙛𝙞 Sapparojpattana & Likhitrakarn, 2026
ตั้งชื่อเป็นเกียรติแก่ Karl Wilhelm Verhoeff ผู้ตั้งสกุลกื้งกือกระสุนลูกกวาด 𝙍𝙝𝙤𝙥𝙖𝙡𝙤𝙢𝙚𝙧𝙞𝙨
ค้นพบครั้งแรกที่ น้ำตกกะทู้ อำเภอกะทู้ จังหวัดภูเก็ต และกระจายตัวหลายพื้นที่ในภาคใต้ของประเทศไทย 

[Entomology • 2026] Staphylus neideae, S. ricardoi, ... • Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera: Hesperiidae: Pyrginae), with Descriptions of Four New Species

[M–P] Staphylus (C.) nicoleae Lemes, sp. nov.; [Q, R] S. (C.) neivae sp. nov.; 
[S, T] S. (C.) ricardoi Lemes, sp. nov.; [U–X] S. (C.) neideae Lemes, sp. nov.: 

in Lemes, Siewert, Mielke, Casagrande et Warren, 2026. 

DOI: doi.org/10.3897/asp.84.e179328 

Abstract

The taxonomy of the subgenus Staphylus (Capilla) is reviewed, including redescriptions of known species, identification keys, and detailed distribution maps. The female genitalia of Staphylus (Capilla) azteca (Scudder, 1872), S. (C.) caribbea (Williams & Bell, 1940), S. (C.) corumba (Williams & Bell, 1940), S. (C.) eryx Evans, 1953, S. (C.) tucumanus (Plötz, 1884) and S. (C.) tyro (Mabille, 1878) are described and illustrated for the first time. Four new species are described: Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, S. (C.) ricardoi Lemes, sp. nov. from Peru, S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil. Pholisora imperspicua Hayward, 1940 is a syn. nov. of Staphylus (C.) lizeri lizeri (Hayward, 1938) and Hesperia melangon epicaste Mabille, 1903 is a syn. nov. of Staphylus (C.) melangon melangon (Mabille, 1883). Neotypes are designated for Staphylus epicaste Mabille, 1903, Nisoniades tucumanus Plötz, 1884 and Staphylus fascia Hayward, 1933. Lectotypes are designated for Pholisora azteca Scudder, 1872, Bolla machuca Schaus, 1913, Helias tyro Mabille, 1878, Staphylus anginus Schaus, 1902, Hesperia melangon 1883, Hesperia musculus Burmeister, 1875 and Helias aurocapilla Staudinger, 1876.

Keywords: Butterflies, Cytochrome oxidase subunit 1, Neotropical, Skippers

Species of Staphylus (Capilla) in dorsal and ventral views.  
A–D S. (C.) ascalon: A, B male (Brazil, São Paulo, Teodoro Sampaio, Morro do Diabo, OM 17.996); C, D female (Brazil, Minas Gerais, Passa Quatro, DZ 46.813).
E, F S. (C.) buena: male (Bolivia, Santa Cruz, Buenavista, holotype, CMNH 7188).
G–J S. (C.) caribbea: G, H male (Costa Rica, Limón Province, La Bomba, MGCL 1112082); I, J female (Costa Rica, Limón Province, Germania, MGCL 1112078).
K–N S. (C.) eryx: K, L male (Brazil, Pará, Santarém, DZ 46.566); M, N female (Brazil, Pará, Santarém, MGCL 1112109).
O–R S. (C.) corumba: O, P male (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.033); Q, R female (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.005).
S, T S. (C.) lizeri album: male (Brazil, Minas Gerais, Carmo do Rio Claro, OM 2.788). U–X S. (C.) lizeri lizeri: U, V male (Peru, Madre de Dios, Tambopata Reserve, DZ 46.176); W, X female (Colombia, Meta, Villavencio, Bosque Bavaria, DZ 45.526).

 Scale bar = 1 cm.

Species of Staphylus (Capilla) in dorsal and ventral views.
A–D S. (C.) tucumanus: A, B male (Argentina, Salta, Pichanal, DZ 45.125); C, D female (Paraguay, Presidente Hayes, Estancia Victoria, Rio Norte Lindo, OM 37.963).
E–H S. (C.) azteca: E, F male (El Salvador, La Libertad, Santa Tecla, MGCL 1112133); G, H female (Mexico, Chiapas, San Jerónimo, DZ 45.085).
I–L S. (C.) tyro: I, J male (Venezuela, Aragua, Ocumare de la Costa, DZ 22.893); K, L female (Venezuela, Aragua, Ocumare de la Costa, DZ 46.608).

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944).
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944). 
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, 

S. (C.) ricardoi Lemes, sp. nov. from Peru, 

S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil.  

 José Ricardo Assmann Lemes, Ricardo Russo Siewert, Olaf Hermann Hendrik Mielke, Mirna Martins Casagrande and Andrew David Warren. 2026. Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera, Hesperiidae, Pyrginae, Carcharodini), with Descriptions of Four New Species. Arthropod Systematics & Phylogeny. 84: 123-173. DOI: doi.org/10.3897/asp.84.e179328 [24 Feb 2026]