[Paleontology • 2026] Dianhaia heptacantha • A non-biomineralized trilobitomorph euarthropod (Trilobitomorpha: Helmetiida: Helmetiidae) with appendicular morphology from the early Cambrian Chengjiang Biota, China

 

Dianhaia heptacantha 

Zhu, Zeng, Liu & Zhao, 2026

 DOI: doi.org/10.1017/jpa.2026.10257

Abstract

Helmetiida is a major group of Trilobitomorpha and Artiopoda known from Cambrian Burgess Shale-type Lagerstätten. Phylogenetic studies have suggested that they are closely related to trilobites as their possible sister taxon. With seven genera and eight species, Helmetiida has only been found in the Burgess Shale and the Chengjiang biota. Here we describe a new helmetiid, Dianhaia heptacantha n. gen. n. sp., from the early Cambrian Chengjiang biota (Cambrian Series 2, Stage 3), Yunnan Province, South China. Dianhaia heptacantha n. gen. n. sp. possesses an anterior sclerite that is diagnostic of Helmetiida. The semicircular cephalon bears a pair of short genal spines. The thorax consists of seven tergites, each having a pair of long pleural spines and showing edge-to-edge articulation. The pygidium carries one short terminal spine and three pairs of long lateral spines. Dianhaia heptacantha n. gen. n. sp. is distinguished from other helmetiids by different numbers of thoracic tergites and pygidial spines. Computed microtomography shows that D. heptacantha n. gen. n. sp. has a pair of antennae and 13 pairs of homonomous biramous appendages. Each biramous appendage consists of a gnathobasic protopodite, seven-podomerous endopodite, and a lobate exopodite with lamellae. Phylogenetic analysis of Artiopoda resolved Dianhaia n. gen. as the sister taxon of Haifengella within Helmetiida. The discovery of this new helmetiid enriches the taxonomic diversity and appendicular morphoanatomy of Cambrian trilobitomorphs from the Chengjiang biota.

Systematic paleontology

Euarthropoda Lankester, 1904

Artiopoda Hou and Bergström, 1997

Trilobitomorpha Størmer, 1944

Conciliterga Hou and Bergström, 1997

Helmetiida Novozhilov, 1960

Helmetiidae Simonetta and Delle Cave, 1975

Included genera: Helmetia Walcott, 1918; Kuamaia Hou, 1987; Rhombicalvaria Hou, 1987; Haifengella Zhao et al., 2014.

Dianhaia heptacantha n. gen. n. sp. from the Cambrian Stage 3 Chengjiang biota, holotype NIGP 203984, part and counterpart.
(1, 2, 4–9) Part, NIGP 203984a; (1) whole specimen; (2) interpretative drawing; (4) endopodite; (5) a thoracic biramous appendage; (6) exopodite; (7) setae on the exopodite; (8) spines on the pleural spine; (9) spines on the pygidium.
(3) Counterpart, NIGP 203984b.

Scale bars represent 5 mm (1, 2), 1 mm (3, 7), 2 mm (4–6, 8, 9).

 Abbreviations: as, anterior sclerite; ce, cephalic segment; cs, cephalic shield; cw, claw; ed, endite; en, endopodite; ex, exopodite; gs, genal spine; gn, gnathobase; hy, hypostome; lm, lamella; ls, lateral spine; ps, pleural spine; pt, protopodite; py, pygidium; sp, marginal spine; ts, terminal spine; T1–7, thoracic tergite 1–7; xs, setae on exopodite.

Dianhaia heptacantha n. gen. n. sp. from the Cambrian Stage 3 Chengjiang biota, paratype NIGP 203985.
(1) Whole specimen; (2) interpretative drawing; (3) pygidial segments; (4) cephalic and anterior thoracic segments with appendages; (5) marginal spines on the pygidial pleural spine; (6) marginal spines on the pygidium; (7) whole appendage; (8) endopodite; (9) setae on exopodite. Scale bars represent: 5 mm (1, 2), 2 mm (3), 1 mm (4–9).

Abbreviations: as, anterior sclerite; en, endopodite; ex, exopodite; ls, lateral spine; ps, pleural spine; pt, protopodite; py, pygidium; sp, marginal spine; st, sternite; ts, terminal spine; T1–7, thoracic tergite 1–7; xs, setae on exopodite.

Genus Dianhaia new genus

Etymology: Derived from Dianhai, another name of the Dian Lake, which is close to the fossil locality.

Remarks: Dianhaia n. gen. has an oval anterior sclerite and tergites with edge-to-edge articulation, supporting its assignment to Helmetiida. Its well-developed segmental boundaries and several large pygidial spines indicate that it was a member of Helmetiidae (Losso et al., 2025). Different from Helmetia and Kuamaia, Dianhaia n. gen. has long pleural spines and pygidial spines that are similar to the counterparts in Rhombicalvaria and Haifengella (Hou and Bergström, 1997; Zhao et al., 2014; Losso et al., 2025). But compared with Rhombicalvaria and Haifengella, Dianhaia n. gen. has different numbers of thoracic tergites and pygidial spines, suggesting it is a separate genus.

Dianhaia heptacantha new species

Holotype: NIGP 203984a, b, part and counterpart.

Materials: Holotype (NIGP 203984a, b); paratype (NIGP 203985).

Diagnosis: The cephalic shield possesses a pair of short genal spines. The thorax comprises seven thoracic tergites with long pleural spines. The semicircular pygidium bears one very short terminal spine and three pairs of long lateral spines. Biramous appendages have an undivided long-ovate flap with marginal setae and imbricated lamellae.

Occurrence: Maotianshan Shale Member, Yu’anshan Formation, Wutingaspis–Eoredlichia trilobite Zone, Cambrian Series 2, Stage 3, Jianshan section, Kunming City, Yunnan Province, China.

Etymology: Hepta means seven in Greek. Acantha means spine in Greek. The name derives from the seven pairs of thoracic spines and seven pygidial spines in this species.

Reconstruction of the helmetiid euarthropod Dianhaia heptacantha n. gen. n. sp. from the Cambrian Stage 3 Chengjiang biota.
 (1) Dorsal view; (2) ventral view. 
 Morphology of the articular membrane and position of antennae are inferred. The number of antennal podomeres was referred from Kuamaia lata.

 

Yuyan Zhu, Han Zeng, Yao Liu and Fangchen Zhao. 2026.  A non-biomineralized trilobitomorph euarthropod (Helmetiida) with appendicular morphology from the early Cambrian Chengjiang Biota, China. Journal of Paleontology.  First View, pp. 1 - 10. DOI: doi.org/10.1017/jpa.2026.10257 [07 October 2026]

[PaleoMammalogy • 2024] Dianoconodon youngi • Fossils document Evolutionary Changes of jaw joint to mammalian middle ear

 

Dianoconodon youngi 

Mao, C. Zhang, Ren, T. Wang, G. Wang, F. Zhang, Rich, Vickers-Rich & Meng, 2024

DOI: doi.org/10.1038/s41586-024-07235-0  

 Researchgate.net/publication/379540254

Reconstruction by Chuang Zhao

Abstract

The dual jaw joint of Morganucodon consists of the dentary–squamosal joint laterally and the articular–quadrate one medially. The articular–quadrate joint and its associated post-dentary bones constitute the precursor of the mammalian middle ear. Fossils documenting the transition from such a precursor to the mammalian middle ear are poor, resulting in inconsistent interpretations of this hallmark apparatus in the earliest stage of mammaliaform evolution. Here we report mandibular middle ears from two Jurassic mammaliaforms: a new morganucodontan-like species and a pseudotribosphenic shuotheriid species. The morganucodontan-like species shows many previously unknown post-dentary bone morphologies and exhibits features that suggest a loss of load-bearing function in its articular–quadrate joint. The middle ear of the shuotheriid approaches the mammalian condition in that it has features that are suitable for an exclusively auditory function, although the post-dentary bones are still attached to the dentary. With size reduction of the jaw-joint bones, the quadrate shifts medially at different degrees in relation to the articular in the two mammaliaforms. These changes provide evidence of a gradual loss of load-bearing function in the articular–quadrate jaw joint—a prerequisite for the detachment of the post-dentary bones from the dentary and the eventual breakdown of the Meckel’s cartilage during the evolution of mammaliaforms.

Reconstruction of the newly described species Dianoconodon youngi.

illustration by Chuang Zhao 

Mammaliaformes Rowe, 1988

Dianoconodon youngi gen. et. sp. nov.

Holotype. A complete skull with associated dentitions and partialpostcranial skeleton (IVPP V4257, Institute of Vertebrate Paleontologyand Paleoanthropology, Chinese Academy of Sciences, Beijing) (Figs. 1and 2; Methods and Extended Data Figs. 1–6).

Etymology. Dian, the short form (Chinese phonetic Pinyin) of Yunnan Province, where the Lufeng fossil site is located; konos, Greek ‘cone’ or ‘cusp’; odon, Greek ‘tooth’. The specific name honours C. C. Young, who initiated the study of mammaliamorphs in Lufeng.  

Mandibular middle ears and transformation of the articular-quadrate joints in mammaliaforms. 

Fangyuan Mao, Chi Zhang, Jicheng Ren, Tao Wang, Guofu Wang, Fakui Zhang, Thomas Rich, Patricia Vickers-Rich and Jin Meng. 2024. Fossils document Evolutionary Changes of jaw joint to mammalian middle ear. Nature. DOI: doi.org/10.1038/s41586-024-07235-0 [03 April 2024]

 Researchgate.net/publication/379540254_Fossils_document_evolutionary_changes_of_jaw_joint_to_mammalian_middle_ear

 

[PaleoMammalogy • 2024] Feredocodon chowi • Jurassic shuotheriids show earliest dental diversification of mammaliaforms

 Feredocodon chowi

Mao, Li, Wang, Zhang, Rich, Vickers-Rich & Meng, 2024

 DOI: doi.org/10.1038/s41586-024-07235-0 

  Researchgate.net/publication/379540347

illustration by Chuang Zhao

Abstract

Shuotheriids are Jurassic mammaliaforms that possess pseudotribosphenic teeth in which a pseudotalonid is anterior to the trigonid in the lower molar, contrasting with the tribosphenic pattern of therian mammals (placentals, marsupials and kin) in which the talonid is posterior to the trigonid. The origin of the pseudotribosphenic teeth remains unclear, obscuring our perception of shuotheriid affinities and the early evolution of mammaliaforms. Here we report a new Jurassic shuotheriid represented by two skeletal specimens. Their complete pseudotribosphenic dentitions allow reidentification of dental structures using serial homology and the tooth occlusal relationship. Contrary to the conventional view, our findings show that dental structures of shuotheriids can be homologized to those of docodontans and partly support homologous statements for some dental structures between docodontans and other mammaliaforms. The phylogenetic analysis based on new evidence removes shuotheriids from the tribosphenic ausktribosphenids (including monotremes) and clusters them with docodontans to form a new clade, Docodontiformes, that is characterized by pseudotribosphenic features. In the phylogeny, docodontiforms and ‘holotherians’ (Kuehneotherium, monotremes and therians) evolve independently from a Morganucodon-like ancestor with triconodont molars by labio-lingual widening their posterior teeth for more efficient food processing. The pseudotribosphenic pattern passed a cusp semitriangulation stage, whereas the tribosphenic pattern and its precursor went through a stage of cusp triangulation. The two different processes resulted in complex tooth structures and occlusal patterns that elucidate the earliest diversification of mammaliaforms.

 Feredocodon chowi 

illustration by Chuang Zhao

Mammaliaformes Rowe, 1988

Shuotheridia Chow and Rich, 1982

 Feredocodon chowi

 

Fangyuan Mao, Zhiyu Li, Zhili Wang, Chi Zhang, Thomas Rich, Patricia Vickers-Rich and Jin Meng. 2024. Jurassic shuotheriids show earliest dental diversification of mammaliaforms. Nature. 628; 569–575. DOI: doi.org/10.1038/s41586-024-07235-0 [03 April 2024] 

  Researchgate.net/publication/379540347_Jurassic_shuotheriids_show_earliest_dental_diversification_of_mammaliaforms

 

[Herpetology • 2026] Ovophis orlovi • An overlooked Mountain Pitviper (Serpentes: Viperidae: Ovophis) from the Montane Regions of Central Vietnam and Southern Laos

 

Ovophis orlovi  

 Idiiatullina, Ustinova, Bragin, Le, Yuhao, Tierui, Pawangkhanant, Ananjeva, David, Vogel, Nguyen & Poyarkov, 2026

 rjh.folium.ru/index.php/rjh/article/view/2315 

facebook.com/tan.sifasv/posts/122268835112144982 

Researchgate.net/publication/415133263

Abstract

An overlooked species of the mountain pitviper genus Ovophis Burger in Hoge and Romano-Hoge, 1981, is described from the highlands of central Vietnam and southern Laos based on an integrative taxonomic approach combining molecular and morphological evidence. Ovophis orlovi sp. nov. has long been confused with O. monticola and, less frequently, O. convictus, but is recovered as a distinct and well-supported evolutionary lineage in mitochondrial phylogenetic analyses, forming the sister taxon to O. anitae. Its distinctiveness is further supported by substantial mitochondrial divergence from its closest relative (cyt b uncorrected p-distance = 8.0%) and by a unique combination of diagnostic morphological characters, including body proportions, scalation, and color pattern. Ovophis orlovi sp. nov. is currently known from montane evergreen forests of the Kon Tum-Gia Lai Plateau in central Vietnam and adjacent montane areas of southern Laos at elevations of approximately 900 – 1,400 m a.s.l. The recognition of Ovophis orlovi sp. nov. resolves a long-standing taxonomic misidentification involving populations previously referred to O. monticola or O. convictus, further demonstrates that species diversity within Ovophis remains underestimated, and highlights the importance of integrative taxonomic studies for uncovering overlooked diversity in the montane regions of mainland Southeast Asia.

Keywords: Ovophis orlovi sp. nov.; integrative taxonomy; molecular phylogeny; morphology; Kon Tum-Gia Lai Plateau; Bolaven Plateau; biogeography

Ovophis orlovi sp. nov.

Variation in coloration of Ovophis orlovi sp. nov. in life.
Vietnam: A, Mang Den, Quang Ngai, VRTC NAP-18718, adult female, holotype; B, Kon Chu Rang NR, Gia Lai, ZMMU R-16315, subadult male, paratype; C, Kon Ka Kinh NP, Gia Lai, ZMMU R-15452, subadult female, paratype.
Laos: Bolaven Plateau, Champasak: D, adult individual; E, F, subadult individual.

Photos by A. M. Bragin (A), N. A. Poyarkov (B, C), and A. Teynié (D – F)

Sabira S. Idiiatullina, Darya A. Ustinova, Andrey M. Bragin, Son Xuan Le, Xu Yuhao, Zhang Tierui, Parinya Pawangkhanant, Natalia B. Ananjeva, Patrick David, Gernot Vogel, Tan Van Nguyen and Nikolay A. Poyarkov. 2026. An overlooked Mountain Pitviper from the Montane Regions of Central Vietnam and Southern Laos (Serpentes: Viperidae: Ovophis). Russian Journal of Herpetology. 33(3); 178-196. http://rjh.folium.ru/index.php/rjh/article/view/2315 [September 30, 2026]

 facebook.com/tan.sifasv/posts/122268835112144982

 Researchgate.net/publication/415133263_An_overlooked_Ovophis_from_the_montane_regions_of_Vietnam_Laos

 

 Ovophis orlovi sp. nov., Mang Den Commune, Quang Ngai Province, central Vietnam, June 6, 2016. Photo by Andrey M. Bragin

 

[Paleontology • 2026] Infernogallus jamesi • Fruit-eating Adaptations in a Late Cretaceous oviraptorosaur (Oviraptorosauria: Caenagnathidae)

 Infernogallus jamesi 

Funston, Dudgeon, Simon, Dufault, Boyd, Fowler, Currie & Evans, 2026

 

DOI: doi.org/10.1038/s41467-026-78343-w

Abstract

The diversification of herbivorous dinosaurs and the rise of angiosperms in the Late Cretaceous are each well established, but the link between them is debated. Broad-scale diversity patterns in the two groups appear not to closely align, and there is no direct evidence of dinosaur adaptation to angiosperm-based diets. Here we describe Infernogallus jamesi gen. et. sp. nov., a diminutive caenagnathid oviraptorosaur, and show that these theropods were specialised herbivores that originated alongside the rise of fleshy angiosperm fruits. The nearly complete, articulated skull of Infernogallus jamesi—the first for the clade—reveals drastic changes to the adductor musculature organisation, unique in non-avian theropods but converged upon by extant frugivorous parrots. These modifications enabled extensive oral processing of soft food using anteroposterior movement of the mandible and occlusion of ridges and grooves on opposed elements. Altogether, this attests to a highly specialised herbivorous diet in caenagnathids, which likely consisted in large part of fleshy angiosperm fruits, which increased in size and diversity in concert with the radiation of oviraptorosaurs. Caenagnathids thus provide one of the strongest candidates for angiosperm-based diet adaptations in dinosaurs, but their role in the Late Cretaceous proliferation of fruit—which shaped early Cenozoic flora—remains unclear.

Systematic palaeontology

Theropoda

Oviraptorosauria

Caenagnathidae

Eucaenagnathinae Clade nov.

Definition. The most inclusive monophyletic group including Anzu wyliei and Elmisaurus rarus, but not Anomalipes zhaoi.

Diagnosis. Eucaenagnathines (meaning ‘true caenagnathids’) are distinguished from other caenagnathids by the morphology of the fused dentaries, with an upturned tomial edge marked with lateral and anterior occlusal ridges, and a midline sulcus bordered by longitudinal lingual ridges separated from the tomial edge by a lingual groove.

Infernogallus jamesi (ROM VP 65050). Photograph (a) and line drawing (b) of ROM VP 65050. c Skeletal reconstruction of Infernogallus jamesi based on ROM VP 65050 and other eucaenagnathines (shaded areas are preserved in ROM 65050). d Palaeohistological thin section of the femur of ROM VP 65050, showing lines of arrested growth (arrows) and an external fundamental system indicative of advanced age and skeletal maturity. cv cervical vertebrae, dv dorsal vertebrae, efs external fundamental system, fe femur, ga gastralia, hu humerus, il ilium, ma manus, pb pubis, pp plane-polarised light, sc scapulacoracoid, sk skull, tb tibia, xp cross-polarised light. Scale bars as indicated.

Cranial anatomy of Infernogallus jamesi gen. et sp. nov.
Photograph (a) and line drawing (b) of the skull of ROM VP 65050 in left lateral view. ...

 Infernogallus jamesi gen et. sp. nov. (Figs. 1–4) 

Etymology. The name comes from the Latin “Inferno” (Hell) and “gallus” (chicken), referring to the Hell Creek Formation and the small, bird-like appearance of the species; “jamesi”, a patronym for James William Boyd, the paternal grandfather of author CAB, who sparked his interest in the field of palaeontology. 

Holotype. ROM VP 65050, articulated partial skeleton including the skull, cervical and dorsal vertebrae, right pectoral girdle and forelimb, partial pelvic girdle, and partial right hindlimb. Paratypes. ROM VP 64193, dentaries; ROM VP 64194, articularsurangular-coronoid complex; ROM VP 88690, left and right pterygoids; NDGS 4351, dentaries. 

Locality and horizon. Hell Creek Formation (late Maastrichtian), Garfield County, Montana; Meade County, South Dakota; and Bowman County, North Dakota, USA. 

Diagnosis. Small (femur length 185 mm; est. 12 ± 3 kg body mass) caenagnathid oviraptorosaur diagnosed by the following autapomorphies: broad jugal with large anterior ascending process and which contacts the squamosal posteriorly; infratemporal fenestra extremely reduced, about 25% the diameter of the orbit; quadratojugal with greatly enlarged, circular posterior process; squamosal with dorsoventrally broad body lacking a depression and ridge for adductor musculature; pterygoids concave dorsally, unfused along the midline, and with a lateral buttress; ectopterygoid–pterygoid bar greatly expanded and extending far ventrally; long axis of vomer oriented ...

Gregory F. Funston, Thomas W. Dudgeon, D. Jade Simon, Danielle M. Dufault, Clint Boyd, Denver Fowler, Philip J. Currie and David C. Evans. 2026. Fruit-eating Adaptations in a Late Cretaceous oviraptorosaur (Theropoda). Nature Communications. DOI: doi.org/10.1038/s41467-026-78343-w [10 October 2026]

[PaleoMammalogy • 2026] Megacauda sungei • A Jurassic mammaliaform swimmer and transformation of the mammalian pharynx

 Megacauda sungei 

Y. Li, P. Li, Neander, Zhang, Zhou, Martin & Luo, 2026 

 

DOI: doi.org/10.1038/s41586-026-11107-0 

Abstract

The discovery of a well-preserved mammaliaform fossil from the Middle Jurassic period of China reveals evidence on the evolutionary transformation of the feeding apparatus and the ecological diversity of Mesozoic era mammals. Mammaliaforms are evolutionary predecessors to modern Mammalia, and their morphologies provide primary insights into the evolutionary origin of mammals. The rare preservation of a therian-like pharyngeal vault (throat passage) and hyoid skeletal anatomy (throat bones) in this mammaliaform suggests that critical soft-tissue anatomy for therian-like swallowing, drinking and suckling behaviours had evolved in the common ancestors of Docodonta and Mammalia, long before the rise of modern mammals. This fossil exhibits a wealth of features specialized for swimming, filling in a major gap of anatomical knowledge of pelvis and hindlimb for early mammals. It is characterized by highly distinctive canines and pseudo-carnassial sectorial teeth for a faunivorous diet. We interpret that this mammaliaform was semiaquatic, swimming and foraging in water, with a lifestyle analogous to the extant platypus. This species is the largest of Jurassic mammaliaforms. It expands the body size range of docodontans, and adds to a growing body of evidence that multiple docodontans exploited diverse niches with semiaquatic and other ecomorphotypes. Docodontans are an early example of replicated evolution of the semiaquatic and other ecomorphotypes in the Mesozoic ecosystem dominated by dinosaurs, long before the diversification of extant mammals.

Holotype main-part of Megacauda sungei (Docodonta, Mammaliaformes) (PMOL–AM00026A).

Megacauda sungei

Y. Li, P. Li, A. I. Neander, H. Zhang, C.-F. Zhou, T. Martin and Z.-X. Luo. 2026. A Jurassic mammaliaform swimmer and transformation of the mammalian pharynx. Nature. DOI: doi.org/10.1038/s41586-026-11107-0 [07 October 2026]

https://news.uchicago.edu/story/uchicago-scientists-reveal-jurassic-otter-could-chew-and-swallow-modern-mammals 

  

 

[Botany • 2026] Beaumontia laotica (Apocynaceae) • A New Species from central Laos

 

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen, 

in Ma, Xi, Lorphengsy et Shen, 2026.

DOI: doi.org/10.5735/085.063.0130 

Abstract

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen (Apocynaceae) is described as a new species from Bolikhamxai Province, Laos. It closely resembles B. grandiflora, B. murtonii and B. jerdoniana, but differs from them by several features of the leaves, sepals and corolla, and by flowering in November. A full taxonomic description, diagnostic illustration and formal IUCN conservation assessment are presented for the new species. An identification key to all species of Beaumontia is provided.

Beaumontia laotica (from the holotype).
 — A: Branch. — B: Longitudinal section of flower. — C: Style-head and anther, side view. — D: Anther, adaxial view. — E: Anther, abaxial view. — F: Ovary, side view.

Beaumontia laotica (A and B photographed at type locality, 11 November 2025; C–P from the holotype).
— A: Natural habitat. — B: Habit. — C: Branch. — D: Young stem. — E: Leaf, adaxial view. — F: Leaf, abaxial view. — G: Adaxial leaf surface. — H: Abaxial leaf surface. — I: Inflorescence. — J: Bracts, adaxial views. — K: Sepals, adaxial views. — L: Flower, front view. — M: Longitudinal section of corolla showing inner surface and stamens. — N: Pedicel and pistil, side view. — O: Disc and ovary, front view. — P: Ovary, side view.

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen, sp. nova

Diagnosis. Beaumontia laotica differs from B. grandiflora, B. murtonii and B. jerdoniana by its lanceolate or narrowly obovate (vs. elliptic or obovate) leaf blade; persistent (vs. deciduous) bracts; glabrous (vs. pubescent) sepals; and glabrous (vs. pubescent or puberulent) corolla tube, lower portion ca. 7.4 cm long (vs. 1.5–2.5 cm long), upper portion abruptly expanding to trumpet-shaped, ca. 2 cm long (vs. obconical or campanulate, 6.5–11 cm long). 

Xing-Da Ma, Hou-Cheng Xi, Sengvilay Lorphengsy and Jian-Yong Shen. 2026. Beaumontia laotica (Apocynaceae), A New Species from central Laos. Annales Botanici Fennici. 63(1), 233-237. DOI: doi.org/10.5735/085.063.0130  (11 September 2026) 

[Entomology • 2026] Anchiale mabiensis & A. robusta • A Revision of Australian Anchiale Stål, 1875 (Phasmida: Phasmatidae), with the Description of Two New Species

  Anchiale mabiensis Coupland & Emmott, sp. nov. female, in situ

in Jones, Brock, Coupland, Emmott et Lo, 2026.

DOI: doi.org/10.1111/aen.70096

 (photo by Angus J. Emmott).

ABSTRACT 

The Australian species of the Phasmida genus Anchiale Stål, 1875 (Phasmatinae: Phasmatini) are updated to include five species: Anchiale albopunctata (Kirby, 1896) stat. rev. (removed from synonymy with A. briareus (Gray, 1834)); A. briareus Gray, 1834 (= Ctenomorpha tessulata Gray, 1835 syn. nov.; = A. austrotessulata Brock & Hasenpush, 2007, replacement name for C. tessulata Gray, 1835), A. mabiensis Coupland & Emmott, sp. nov.; A. robusta Brock & Jones, sp. nov.; and A. spinicollis Gray, 1833. Lectotypes are designated for Ctenomorpha albopunctatum Kirby, 1896, C. nigrovaria Stål, 1875 (= A. briareus) and C. tessulata Gray, 1835. These taxonomic decisions are supported by Bayesian and Maximum Likelihood phylogenetic analyses based on the mitochondrial COXI gene fragment of all species except A. spinicollis. Anchiale albopunctata is currently known from mideastern Queensland, A. briareus from north-eastern New South Wales and south-eastern Queensland, A. mabiensis sp. nov. and A. robusta sp. nov. from tropical eastern Queensland, and A. spinicollis from the Northern Territory. A dichotomous key to all Australian species of Anchiale and their eggs is provided.

Keywords: Acrophylla, distribution, DNA, genus, insects, molecular, Phasmida, phylogenetic, specimens, taxonomy

Anchiale albopunctata (Kirby, 1896) stat. rev. 

(removed from synonymy with A. briareus (Gray, 1834)); 

A. briareus Gray, 1834 

(= Ctenomorpha tessulata Gray, 1835 syn. nov.; 

= A. austrotessulata Brock & Hasenpush, 2007,
replacement name for C. tessulata Gray, 1835), 

A. mabiensis Coupland & Emmott, sp. nov.; 

A. robusta Brock & Jones, sp. nov.; 

A. spinicollis Gray, 1833. 

  Anchiale mabiensis sp. nov. female, in situ

 (photo by Angus J. Emmott).

Anchiale mabiensis sp. nov. in situ.
(a) Female holotype (photo by Ross M. Coupland).
(b) paratype male (photo by Ross M. Coupland).
(c) Montane rainforest habitat (photo by Angus J. Emmott).

Anchiale mabiensis Coupland & Emmott, sp. nov. 

Anchiale robusta sp. nov. in situ.
 (a) Male. (b) Female. (c) Female defence display, inset: wings of preserved paratype female. (d) Coastal habitat, Bramston Beach, Qld. (e) Garden habitat, Kuranda, Qld

(all photos by Paul D. Brock).

Anchiale robusta Brock & Jones, sp. nov.

Braxton R. Jones, Paul D. Brock, Ross M. Coupland, Angus Emmott and Nathan Lo. 2026. A Revision of Australian Anchiale Stål, 1875 (Phasmida: Phasmatidae), with the Description of Two New Species. Austral Entomology. 65(4); e70096. DOI: doi.org/10.1111/aen.70096 [21 September 2026]

[Mollusca • 2026] Sitala chorchati • A New Species of the Tiny Tree Snail Genus Sitala H. Adams, 1865 (Eupulmonata: Helicarionidae) from Northeastern Thailand

 

Sitala chorchati Tanmuangpak & S. Tumpeesuwan, 

in Tanmuangpak, Nahok, Chanlabut, Taesuk, S. Tumpeesuwan et C. Tumpeesuwan, 2026. 

หอยยอดเจดีย์ช่อฉัตร | DOI: doi.org/10.3390/taxonomy6040061 

 

Abstract

A new species of the tiny tree stylommatophoran snails in the genus Sitala H. Adams, 1865, is described based on specimens recently discovered in dry evergreen and dry dipterocarp forests on limestone hills and in agricultural plantations near limestone hills in Loei Province, northeastern Thailand. The new species is characterized by a high, pyramid-shaped, thin, and translucent yellowish-brown shell with a sharp peripheral keel. Although its external shell morphology resembles that of Sitala liricincta (Stoliczka, 1871), the new species can be distinguished by its greater number of whorls (9¼). Comparative anatomical analysis further revealed significant interspecific differences in genital organs, particularly in the length and shape of the vagina, and in the proportions of the penis, epiphallus, and penial caecum. This discovery is the first verified record of the genus Sitala in Thailand.

Keywords: shell morphology; radula; epiphallic retractor caecum; leaf inhabitant; limestone hill

Taxonomy

Superfamily Helicarionoidea Bourguignat, 1877

Family Helicarionidae Bourguignat, 1877

Subfamily Durgellinae, Godwin-Austen, 1888

Tribe Durgellini Godwin-Austen, 1888

Genus Sitala H. Adams, 1865

Type species. Helix infula Benson, 1848

Diagnosis of genus. Shell turbinoid, thin and translucent, conic or pyramid in shape, outlines, spire conical, consisting of 5.5–7 convex striated whorls. Protoconch and subsequent whorls covered with spiral sculpture, base convex. Umbilicus minute, partially covered. Aperture lack teeth. Peristome thin, simple and not expanded. Penis, vagina, and spermatheca (gametolytic sac) connect to atrium; Penis long, subcylindrical. Epiphallus short, with a simple epiphallic caecum produce into penial retractor. Vas deferens entering epiphallus laterally. Vagina very short (or absent), depending on the position at which the spermatheca enter the genital tract near the atrium. Free oviduct relatively long. Spermathecal stalk enlarges basally; reservoir elongated and embedded in spermoviduct. Amatorial organ (Sarcobellum) present or absent.

 Live snail, Sitala chorchati sp. nov., paratype NHLRU080.

 Photograph: Kitti Tanmuangpak.

 Shell morphology of Sitala chorchati sp. nov., holotype NHLRU079.
(A), Apertural view; (B), Lateral view; (C), Ventral view; (D), Dorsal view.

Photograph: Kitti Tanmuangpak.

Sitala chorchati Tanmuangpak & S. Tumpeesuwan, sp. nov.

Diagnosis. Sitala chorchati sp. nov. is similar in shell morphology to Sitala attegia (Benson, 1859) and Sitala liricincta (Stoliczka, 1871), particularly in having shell conical, high spire shell, thin and translucent. Differing from both species in smaller shell and a greater number of whorls, 4.12–5.28 mm and 9¼ whorls, whereas 5.8 mm and 6½–7 whorls in S. liricincta, and 7–12 mm and 6½–8 whorls in S. attegia (Table 4).

Etymology. The specific epithet “chorchati” is given in honour of Admiral Chorchat Gra-tes of the Royal Thai navy, former Deputy Chief of Defence, an independent conchologist and owner of a private shell collection in Bangkok, Thailand.

 Kitti Tanmuangpak, Benchawan Nahok, Utain Chanlabut, Nonthiwat Taesuk, Sakboworn Tumpeesuwan and Chanidaporn Tumpeesuwan. 2026. A New Species of the Tiny Tree Snail Genus Sitala H. Adams, 1865 (Gastropoda, Eupulmonata, Helicarionidae) from Northeastern Thailand. Taxonomy. 6(4), 61. DOI: doi.org/10.3390/taxonomy6040061 [7 October 2026]

[Botany • 2026] Primula octopoda (Primulaceae) • A stoloniferous New Species from western Sichuan, China

 

Primula octopoda W.B.Ju & Q.Huang,  

in Q. Huang, X. Wang, W.-J. Huang, Yang, Pan, Ru, J.-P. Wang et Ju, 2026.

巴塘粉报春 | DOI: doi.org/10.1002/ece3.74088 

 

ABSTRACT

Primula octopoda W.B.Ju & Q.Huang, a new species of Primula (Primulaceae) in sect. Aleuritia, is herein described and illustrated from Batang County, Sichuan Province, southwestern China. Morphologically, P. octopoda is most similar to P. stenocalyx, but it can be readily distinguished by its dwarf habit, the presence of stolons 2–7 cm in length ending in small rosettes occasionally bearing a single flower, elliptic to broadly obovate leaf blades, a very short or absent scape bearing one or two flowers, and a smaller, annulate corolla with a prominent yellow eye. The complete chloroplast genome of P. octopoda spans 154,164 bp with a GC content of 36.97% and contains 125 genes. Phylogenetic analyses using plastid and ITS data both place P. octopoda within sect. Aleuritia while supporting its distinct species status. Given the current lack of adequate information on its population size and geographic range, the species is provisionally assigned a status of Data Deficient (DD) according to the IUCN Red List criteria.

Keywords: molecular phylogeny, morphology, new species, Primula, taxonomy

Habitat of the Primula octopoda.
 (A) Habitat; (B–D) Plants.

Morphology of Primula octopoda.
 (A) Plant; (B) roots; (C) leaves; (D) inflorescence; (E) corolla; (F) bracts; (G) calyx; (H) thrum flower and pin flower; (I) capsule.

Primula octopoda W.B.Ju & Q.Huang, sp. nov. 

Diagnosis. The new species most closely resembles P. stenocalyx, sharing narrowly lanceolate bracts, a tubular calyx with similarly divided oblong or lanceolate lobes, stamens and style included within the corolla tube, and an oblong capsule. However, it differs in its shorter stature, the presence of 2–7 cm long stolons each ending in a small rosette and occasionally bearing a solitary flower, elliptic to broadly obovate leaves, a scape up to 2 cm long or absent with 1–2 flowers, and an annulate corolla with a yellow eye and smaller flowers (Figures 2 and 3).

Etymology. The specific epithet of the new species refers to its distinctive feature of producing multiple stout, fleshy stolons that resemble the tentacles of an octopus; hence the name octopoda, which vividly reflects the plant's numerous stolons extending like “eight legs.”

Vernacular Name. Chinese mandarin: “bā táng fĕn bào chūn” (巴塘粉报春), meaning “Batang Farinose Primrose” in English.

 

Qin Huang, Xin Wang, Wen-Jun Huang, Jie-Yi Yang, Hong-Li Pan, Jian Ru, Jiang-Ping Wang and Wen-Bin Ju. 2026. Primula octopoda, A stoloniferous New Species from Western Sichuan, China. Ecology and Evolution. DOI: doi.org/10.1002/ece3.74088 [27 August 2026]