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| Fig. 3. A–D, Shell and operculum of Pterocyclos rupestris: A, lectotype UMZC 2359.1; B, paralectotype UMZC 2359.2; C, syntype of var. 3 UMZC 2359.3; D, operculum from same lot of the paralectotype (showing top, side and bottom views). E, F, Specimens of P. blandi from Langkawi Island, Perlis, Malaysia CUMZ 4582. G, Holotype UMZC 1032 of P. subalatus. H–J, Pterocyclos spaleotes: H, lectotype NMW 1955.158.01107, I, paralectotype NMW 1981.118.02705; J, topotype specimen CUMZ 4585. K, Holotype ZMA Moll. 135622 of P. umbraticus. L–P, Pterocyclos diluvium Sutcharit & Panha, new species: L, holotype CUMZ 4595, M, paratype CUMZ 4588, N, specimen from Gua Cenderawasih, Perlis, Malaysia CUMZ 4592; and O, P, specimens from Tam Tone-din, Kuan-Don, Satun, Thailand CUMZ 4590 showing a: O, uniform whitish shell; and P, dark brown colour patterns. Q–S, Pterocyclos frednaggsi Sutcharit & Panha, new species: Q, holotype CUMZ 4594; R, paratype CUMZ 4581; S, specimen from Gua Pulai, Gua Musang, Kelantan, Malaysia CUMZ 4597 |
Abstract
Specimens of the operculated land snail genus Pterocyclos Benson, 1832, from Thailand and Peninsular Malaysia were investigated based on their shell characteristics. Type specimens and topotypic material were studied and compared with newly collected specimens. Two new species are described, viz. Pterocyclos diluvium Sutcharit & Panha, new species, from Tam Sumano, Patthalung, Thailand, adjacent areas in Thailand, and Malaysia, and Pterocyclos frednaggsi Sutcharit & Panha, new species, collected from Bukit Chintamani, Pahang, Malaysia, and adjacent areas in Peninsular Malaysia. Two species that hitherto were included in Pterocyclos, viz. P. blandi Benson, 1851 and P. subulatus Sykes, 1903, were retained in this genus, while two other former Pterocyclos species were re-assigned to other genera, viz. Pearsonia regelspergeri (Morgan, 1885) and Ptychopoma perrieri (Morlet, 1889). Conversely, two previously described species in other genera (Rhiostoma spaleotes Tomlin, 1932, and Cyclotus umbraticus Benthem Jutting, 1949) are here transferred to Pterocyclos, viz. P. spaleotes (Tomlin, 1932) and P. umbraticus (Benthem Jutting, 1949). To stabilise the nomenclatural status, the lectotype of Pterocyclos rupestris
Benson, 1832, and Pterocyclos spaleotes are designated herein.
Key words: Taxonomy, systematics, limestone, biodiversity, Gastropoda
Chirasak Sutcharit, Piyoros Tongkerd & Somsak Panha. 2014. The Land Snail Genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with Descriptions of Two New Species. Raffles. Bull. Zool. 62: 330–338.
