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| Drymonia quadrangulata Clavijo & J.L.Clark, in Clavijo et Clark, 2024. |
Abstract
A new species, Drymonia quadrangulata Clavijo & J.L.Clark, sp. nov. (Gesneriaceae, Columneinae), is described from the western Andean slopes of southern Colombia and northern Ecuador. The new species has been historically confused with D. killipii, an endemic species to Colombia, restricted to the lowlands of the Chocó biogeographic region in the departments of Cauca, Chocó, and Valle del Cauca. These two species have large foliaceous calyx lobes that cover at least half of the corolla tube, and tubular-infundibuliform corollas. The new species differs by strigose quadrangulate and sometimes winged stems, leaves reticulate abaxially with obtuse to subcordate bases, midveins green, corolla lobes white to yellow with margins incised to short laciniate, and indehiscent berry fruits. Digital photographs, detailed morphological comparisons with the similar species, and an IUCN preliminary risk extinction assessment are provided for the new species.
Key words: Andes, Chocó biogeographic region, Colombia, Columneinae, Ecuador
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| Drymonia quadrangulata Clavijo & J.L.Clark A lateral view of corolla B front view of flower (yellow form) C lateral view of flower D front view of flower (white form) E quadrangular stem F abaxial surface of leaf G gynoecium H lateral view of immature fruit (A, G from L. Clavijo et al. 1879 B from J.L. Clark 13609 C, E J.L. Clark et al. 10344 D J.L. Clark 17079 F from J.L. Clark 18123 H from J.L. Clark 16320). Photos: A, G by L. Clavijo, B–F, H by J.L. Clark. |
Drymonia quadrangulata Clavijo & J.L.Clark, sp. nov.
Diagnosis: Differs from Drymonia killipii by quadrangular to strongly angulate stems in cross-sections vs. terete; stems strigose apically vs. hirsute; blades 12.0–26.7 cm long vs. 20.5–45.0 cm long; corolla lobes white to yellow vs. royal purple to maroon; and fruits indehiscent globose berries vs. bivalved fleshy capsules.
Etymology: The specific epithet is in reference to the quadrangular to strongly angulate stem cross-section which is occasionally winged (Figs 2E, 3).
Laura Clavijo and John L. Clark. 2024. Resolving the Drymonia killipii (Gesneriaceae) Complex results in A New Species from the northwestern Andes of South America. PhytoKeys. 248: 207-222. DOI: doi.org/10.3897/phytokeys.248.123248
Resumen: Se describe una nueva especie, Drymonia quadrangulata Clavijo & J.L.Clark, sp. nov. (Gesneriaceae, Columneinae), de la vertiente occidental de los Andes del sur de Colombia y el norte de Ecuador. La nueva especie ha sido históricamente confundida con D. killipii, una especie endémica de Colombia, restringida a las tierras bajas del Chocó biogeográfico en los departamentos de Cauca, Chocó y Valle del Cauca. Estas dos especies tienen un cáliz foliáceo que cubre al menos la mitad del tubo de la corola y corolas tubulares-infundibuliformes. La nueva especie se distingue por sus tallos estrigosos, cuadrangulares y a veces alados, hojas reticuladas en el envés con bases obtusas a subcordadas, nervadura central verde, lóbulos de la corola blancos a amarillos con márgenes incisos a cortamente laciniados y frutos en bayas indehiscentes. Se proporcionan para la especie nueva fotografías digitales, comparaciones morfológicas detalladas con las especies similares y una evaluación preliminar de riesgo de extinción de acuerdo con los criterios de la UICN.
