|Figure 4. Skull and mandible of Gavialis cf. bengawanicus from the Early Pleistocene of Khok Sung (Nakhon Ratchasima Province, Thailand). |
A, right lateral view (mirrored for comparison) of DMR-KS-03-25-23. B, line drawing from the left lateral view of the posterior portion of the skull of DMR-KS-201202-1. C, left lateral view of skull of DMR-KS-201202-1. D, E, G, mandible of DMR-KS-201202-1 left lateral (D), occlusal (E), and ventral (G) views. F, incomplete dentaries (DMR-KS-05-06-22-1) in occlusal view; H, detail of the dentition in lateral view as seen on the maxillary fragment DMR-KS-05-03-08-37.
Abbreviations: asan, anterior tip of surangular; asp, anterior tip of splenial; ec, ectopterygoid; exo, exoccipital; fr, frontal; j, jugal; l, lacrimal; ltf, lower temporal fenestra; mx, maxilla; on, otic notch; or, orbit; pfr, prefrontal; po, postorbital; q, quadrate; qj, quadratojugal; sq, squamosal. || DOI: 10.1371/journal.pone.0044541
The genus Gavialis comprises a single living but endangered species, G. gangeticus, as well as fossil species recorded in the Miocene to Pleistocene deposits of the Indian subcontinent. The genus is also represented in the Pleistocene deposits of Java by the species G. bengawanicus, which was recently recognized to be valid. Surprisingly, no detailed report of the genus exists between these two provinces and the recent evolutionary history of Gavialis is not understood.
We report new material consisting of skull and mandibular remains of Gavialis from the Early Pleistocene of Khok Sung, Nakhon Ratchasima Province, northeastern Thailand. The Gavialis material described herein is attributed to Gavialis cf. bengawanicus and sheds new light on the occurrence of the genus in mainland SE Asia.
Comparison of this new material with other species referred to the genus Gavialis led us to preliminary restrict the content of the genus to three species, namely G. gangeticus Gmelin, G. bengawanicus Dubois and G. lewisi Lull. The occurrence of G. cf. bengawanicus in Thailand allows us to propose a scenario for the dispersal of Gavialis from Indo-Pakistan to Indonesia, thus bridging a geographical gap between these two provinces. Dispersal by sea appears a less likely possibility than dispersal through fluvial drainages.
Jeremy E. Martin, Eric Buffetaut, Wilailuck Naksri, Komsorn Lauprasert and Julien Claude. 2012. Gavialis from the Pleistocene of Thailand and Its Relevance for Drainage Connections from India to Java. PLoS ONE. 7(9): e44541. DOI: 10.1371/journal.pone.0044541