Monday, October 13, 2025

[Botany • 2025] Sideritis carpetana (Lamiaceae) • A New high-mountain Mediterranean Species from the marble outcrops of the Sierra de Guadarrama (Central System, Madrid, Segovia, Spain)


Sideritis carpetana  Izquierdo & Gavilán,

in Izquierdo, Ugarte, Gutiérrez-Girón, Castro, Núñez et Gavilán, 2025. 

Abstract
A new species of Sideritis (Sideritis carpetana) is described from the calcareous, high-mountain Spanish flora in the central part of the Iberian Peninsula. It is found in a Mediterranean climate at high-elevation, perennial, calcareous grasslands, as well as in marble screes of anthropogenic origin in the Sierra de Guadarrama, Central System (Spain), in a reserve area within the Sierra de Guadarrama National Park, at 1996 m asl. Taxonomic morphological measurements were performed on collected specimens from Sierra de Guadarrama as well as on geographically-adjacent Sideritis (i.e., S. glacialis, S. pungens, S. hyssopifolia). The relationships among them were then explored with multivariate analysis. Sideritis carpetana is a dwarf shrub with an erect or decumbent habit, growing up to 15 cm; non-woody twigs with long hairs of 3–4 cells, leaves are entire, linear-oblanceolate, sparsely covered with trichomes; inflorescence is spiciform or slightly verticillated, flowers are yellow and nutlets ovoid. A key is supplied to help distinguish it from other high-mountain Iberian species included in section Sideritis. The species is unique in its combination of morphological and autoecological characters. S. carpetana shares similarities with S. glacialis, a species from Sierra Nevada, and its northern Mediterranean variant, S. glacialis subsp. fontqueriana from Sierra de Gúdar. They share morphological characters that are absent in other high-mountain Sideritis, reinforcing their Mediterranean character, as opposed to a more temperate or submediterranean character, such as those of the hyssopifolia group.

Key words: Calcareous high-mountain grasslands, Carpetan Mountains, high mountain species, marble outcrops, Mediterranean area, Sideritis sect. SideritisSideritis subsect. Fruticulosa

Sideritis carpetana natural habit in the marble anthropogenic outcrops of La Flecha Pass (Sierra de Guadarrama National Park).

Sideritis carpetana, Sierra de Guadarrama, La Flecha Pass (1996 m asl)
a habit b Lower branches c inflorescence detail showing lower verticillaster bract and calyx d inflorescence detail showing upper verticillaster bract.

 Sideritis carpetana Izquierdo & Gavilán, sp. nov.


Jose Luis Izquierdo, Rosina Magaña Ugarte, Alba Gutiérrez-Girón, Concepción Obón de Castro, Diego Rivera Núñez and Rosario G. Gavilán. 2025. Sideritis carpetana (Labiatae), A New high-mountain Mediterranean Species from the marble outcrops of the Sierra de Guadarrama (Central System, Madrid, Segovia, Spain). PhytoKeys. 251: 143-159. DOI: doi.org/10.3897/phytokeys.251.129982