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| Nannotanyderus granieri Azar, Maksoud, Huang, Mounir Maalouf & Cai, 2024 |
Abstract
A new species, †Nannotanyderus granieri sp. nov., belonging to the Tanyderidae (Diptera, Nematocera), from Lower Barremian amber of Bqaatouta (Lebanon), is characterized, described, figured, and its taxononomic position discussed. The possible confusion regarding the attribution of new fossils to either the Tanyderidae or the Psychodidae, in addition to the results of recent molecular phylogenies, have led us to re-evaluate the taxonomic position of the Tanyderidae and to consider it as a subfamily within the Psychodidae. The fossil described herein is a tiny tanyderine sensu nov. species, and the second one belonging to the genus Nannotanyderus from Lebanese amber. A tentative molecular phylogeny of recent Psychodidae and an exhaustive catalogue of fossil Tanyderinae stat. nov. are provided.
Keywords: • Diptera; • Tanyderidae; • Psychodidae; • Tanyderinae stat. nov.; • Tanyderini; • Nannotanyderini; • Lower Cretaceous; • Lebanon
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| Nannotanyderus granieri sp. nov., holotype, female, specimen number BKT-11A. A) Habitus, right lateral side. Scale bar: 500 µm. B) Habitus, left lateral side. Scale bar: 500 µm. C) Head photomicrograph with confocal microscope. Scale bar: 100 µm. D) Wing photomicrograph with confocal microscope. Scale bar: 500 µm. E) Female terminalia, photomicrograph with confocal microscope. Scale bar: 50 µm. F) Female terminalia, photomicrograph with compound microscope. Scale bar: 50 µm. |
Order DIPTERA LINNAEUS, 1758
Infraorder PSYCHODOMORPHA HENNIG, 1968
Family PSYCHODIDAE NEWMAN, 1834
former family Tanyderidae OSTEN-SACKEN, 1880
Subfamily TANYDERINAE OSTEN-SACKEN, 1880, stat. nov.
former family Tanyderidae OSTEN-SACKEN, 1880
Tribe NANNOTANYDERINI SKIBIŃSKA, 2016, stat. nov.
former subfamily Nannotanyderinae SKIBIŃSKA, 2016
Genus NANNOTANYDERUS ANSORGE, 1994
Type species. Nannotanyderus krzeminskii ANSORGE, 1994,
Lower Jurassic (Toarcian), Grimmen, Germany.
Nannotanyderus granieri sp. nov.
Material. Holotype (specimen BKT-11A), female, part of the MAALOUF collection, Lebanese Lower Cretaceous amber found at the Bqaatouta outcrop, in the Caza (District) of Kesserouan, Central Lebanon. Syninclusions include a spider and a male chirbnomid dipteran. This material is housed in the collections of Natural History Museum of the Lebanese University, Faculty of Sciences II, located in Fanar, Lebanon.
Diagnosis. Female only. Head lacking an elongated neck; mouthparts well developed, longer than head, with well-developed sclerotized maxillae. R2 very short, slightly longer than half of R3, with R2+3 about 3 times longer than R2, hooklike at its basal part. Presence of cross-veins mm
and mcu, as well as cell d.
...
Etymology. The species is named after Prof. Bruno GRANIER, whose research has significantly advanced the dating of amber outcrops in Lebanon (e.g., GRANIER et al., 2015, 2016).
Dany AZAR, Sibelle MAKSOUD, Di-Ying HUANG, Mounir MAALOUF and Chen-Yang CAI. 2024. A New fossil psychodomorphan Fly from Lower Barremian Lebanese Amber elucidates the relationship of the Tanyderinae stat. nov. within the Psychodidae. Carnets Geol. 23(6); 113-125. DOI: 10.2110/carnets.2024. 2406
