[PaleoMammalogy • 2026] Olotimcolobus narokensis & Lemucolobus orkimusei • A Systematic Revision of the Late Miocene Cercopithecid Monkeys From the Lemudong'o Formation, Kenya

 

Olotimcolobus narokensis

Arenson, Gilbert, Yang, Ambrose, Kinyanjui, Manthi, DiMaggio & Hammond, 2026

 DOI: doi.org/10.1002/ajpa.70317 

ABSTRACT

Objectives: The late Miocene Lemudong'o Formation in southwestern Kenya (6.2–6.0 Ma) samples an important period in Africa when early representatives of modern catarrhine clades (Hominidae and Cercopithecidae) appear. Fossils collected from this formation before 2007 include three species of colobine monkey. However, several new fossils have been recovered, including a partial cranium, calling for a reassessment of Lemudong'o Formation cercopithecid taxonomy.

Materials and Methods: This study describes 79 new cercopithecid specimens from collection years 2007–2016 and 2019–2024. These specimens, together with previously published material from the Lemudong'o Formation, are compared to fossil cercopithecids from other contemporaneous sites in Africa as well as a sample of extant cercopithecids.

Results: New material supports the presence of the three previously identified colobine taxa and identifies a novel fourth colobine species. The novel colobine species is described as Olotimcolobus narokensis gen. et sp. nov., and the previously recognized small indeterminate colobine is here named Lemucolobus orkimusei gen. et sp. nov. Existing hypodigms are refined in light of this new taxonomy. A papionin is also identified from this formation for the first time, adding to the scant late Miocene cercopithecine record.

Discussion: The Lemudong'o Formation is remarkable for its high cercopithecid diversity in the Late Miocene. The predominance of colobines here and at other contemporaneous sites (e.g., Lukeino, Lothagam, Nakali, Toros Menalla) suggests that colobines were more diverse than cercopithecines through the early part of their radiation. Possible ecological factors are discussed in the context of early cercopithecid diversity and evolution.

 Olotimcolobus narokensis gen. et sp. nov.

 

Julia L. Arenson, Christopher C. Gilbert, Deming Yang, Stanley H. Ambrose, Rahab N. Kinyanjui, Fredrick K. Manthi, Erin N. DiMaggio and Ashley S. Hammond. 2026. A Systematic Revision of the Late Miocene Cercopithecid Monkeys From the Lemudong'o Formation, Kenya. American Journal of Biological Anthropology. 190(3); e70317. DOI: doi.org/10.1002/ajpa.70317 [27 July 2026]

Descobreixen dos nous gèneres de primats de fa 6 milions d’anys en un jaciment fòssil de Kenya

https://www.icp.cat/index.php/ca/sala-de-premsa-4/noticies-icp/item/3708-descobreixen-dos-nous-generes-de-primats-de-fa-6-milions-d-anys-en-un-jaciment-fossil-de-kenya

[Herpetology • 2026] Calamaria flavicollaris • Integrative Taxonomy reveals A New Species of Calamaria (Squamata: Calamariidae) from the Tenasserim Range, Western Thailand

 

 Calamaria flavicollaris

 Aksornneam, Rujirawan, Yodthong, Pawangkhanant, Sung & Aowphol, 2026

 

Tenasserim Collared Reed Snake | งูพงอ้อบ่วงบาศ ||  DOI: doi.org/10.1163/18759866-bja10102 

 Photographed by Parinya Pawangkhanant

Abstract

A new species of Calamaria is described from the Tenasserim Range, Ratchaburi Province, western Thailand based on morphological and molecular evidence. Calamaria flavicollaris sp. nov. can be distinguished from its congeners by the combination of the following morphological characters: 1) rostral scale wider than high; 2) portion of rostral scale visible from above greater than the half the length of the prefrontal suture; 3) prefrontal length shorter than frontal length, not entering orbit and in contact with first two supralabials; 4) five or six scales surrounding paraparietal scale; 5) preocular and postocular present; 6) four supralabials, the second and third in contact with the eyes; 7) eye diameter smaller than eye-mouth distance; 8) five infralabials, the first pair separating the mental scale from anterior chin shields; 9) seven modified maxillary teeth; 10) 180 ventral scales with three preventral scales; 11) 13 divided subcaudal scales; 12) cloacal plate single; 13) tail relatively short (4.5% of the total length), tapering, gradually to a pointed tail tip; 14) dorsum dark brown with eleven dark longitudinal stripes; 15) ventral surface yellow with a median line of dark flecks on underside of tail; and 16) dark collar followed by yellow ring at nuchal region. Genetically, the new species exhibits a pairwise distance of ≥17.56% from all other congeners based on the mitochondrial cytochrome b gene fragment. Furthermore, we provide cranial osteological description of the new species based on micro-CT scanning. Calamaria flavicollaris sp. nov. is currently known only from the type locality and represents the fourth species of the genus recorded for Thailand. 

Keywords: cranial osteology; herpetofauna; Isthmus of Kra; molecular; reed snake; Southeast Asia

Holotype of Calamaria flavicollaris sp. nov. (ZMKU R 01131) in life. A small figure shows the yellow color on the ventral side of this species 

 Photographed by Parinya Pawangkhanant

Head of  Calamaria flavicollaris sp. nov. (ZMKU R 01131).
A–B lateral view C–D dorsal view E–F ventral view

Holotype of  Calamaria flavicollaris sp. nov. (ZMKU R 01131)
in preservative. A dorsal view B ventral view C lateral view at the midbody D dorsal view of tail E ventral view of tail

Calamaria flavicollaris sp. nov.

Diagnosis. Calamaria flavicollaris sp. nov. can be distinguished from all other congeners by the following combination of characters (Tables 3, 4; Suppl. Table 1): 1) rostral scale wider than high; 2) portion of rostral scale visible from above greater than the half the length of the prefrontal suture; 3) prefrontal length shorter than frontal length, not entering orbit, and in contact first two supralabials; 4) five or six scales surrounding the paraparietal scale; 5) preocular and postocular present; 6) four supralabials, the second and third in contact the eyes; 7) eye diameter smaller than eye-mouth distance; 8) five infralabials, the first pair separating the mental scale from anterior chin shields; 9) seven modified maxillary teeth; 10) 180 ventral scales with three preventral scales; 11) 13 divided subcaudal scales; 12) cloacal plate single; 13) tail relatively short (4.5% of the total length), tapering, gradually to a pointed tail tip; 14) dorsum dark brown with eleven dark longitudinal stripes; 15) ventral surface yellow with a median line of dark flecks on underside of tail; and 16) a dark nuchal collar followed by a yellow ring along neck.

Four Calamaria species in Thailand:
A Calamaria flavicollaris sp. nov. from Suan Phueng District, Ratchaburi Province B C. lumbricoidea from Than To District, Yala Province
C C. schlegeli from Betong District, Yala Province D C. pavimentata from Kaeng Hang Maeo District, Chanthaburi Province 

Photos by Parinya Pawangkhanant (A–C) and Attapol Rujirawan (D)

Etymology. The specific epithet flavicollaris is derived from the Latin words “flavo” (yellow), “collare” (collar or neck), and the suffix “-aris”, which refers to the appearance of the yellow nuchal ring of this species. 

We suggested an English common name “Tenasserim Collared Reed Snake” and the Thai common name “งูพงอ้อบ่วงบาศ” (Ngu Phong O Buang Bat) for this species.

 Akrachai Aksornneam, Attapol Rujirawan, Siriporn Yodthong, Parinya Pawangkhanant, Yik-Hei Sung, and Anchalee Aowphol. 2026. Integrative Taxonomy reveals A New Species of Calamaria (Squamata: Calamariidae) from the Tenasserim Range, Western Thailand.  Contributions to Zoology. 95(4); 366–410. DOI: doi.org/10.1163/18759866-bja10102 [22 Sep 2026]

งูพงอ้อหัวแดง Calamaria schlegeli; 

งูพงอ้อหลากลาย Calamaria lumbricoidea; 

งูพงอ้อท้องเหลือง Calamaria pavimentata

[Herpetology • 2026] Ovophis orlovi • An overlooked Mountain Pitviper (Serpentes: Viperidae: Ovophis) from the Montane Regions of Central Vietnam and Southern Laos

 

Ovophis orlovi  

 Idiiatullina, Ustinova, Bragin, Le, Yuhao, Tierui, Pawangkhanant, Ananjeva, David, Vogel, Nguyen & Poyarkov, 2026

 rjh.folium.ru/index.php/rjh/article/view/2315 

facebook.com/tan.sifasv/posts/122268835112144982 

Researchgate.net/publication/415133263

Abstract

An overlooked species of the mountain pitviper genus Ovophis Burger in Hoge and Romano-Hoge, 1981, is described from the highlands of central Vietnam and southern Laos based on an integrative taxonomic approach combining molecular and morphological evidence. Ovophis orlovi sp. nov. has long been confused with O. monticola and, less frequently, O. convictus, but is recovered as a distinct and well-supported evolutionary lineage in mitochondrial phylogenetic analyses, forming the sister taxon to O. anitae. Its distinctiveness is further supported by substantial mitochondrial divergence from its closest relative (cyt b uncorrected p-distance = 8.0%) and by a unique combination of diagnostic morphological characters, including body proportions, scalation, and color pattern. Ovophis orlovi sp. nov. is currently known from montane evergreen forests of the Kon Tum-Gia Lai Plateau in central Vietnam and adjacent montane areas of southern Laos at elevations of approximately 900 – 1,400 m a.s.l. The recognition of Ovophis orlovi sp. nov. resolves a long-standing taxonomic misidentification involving populations previously referred to O. monticola or O. convictus, further demonstrates that species diversity within Ovophis remains underestimated, and highlights the importance of integrative taxonomic studies for uncovering overlooked diversity in the montane regions of mainland Southeast Asia.

Keywords: Ovophis orlovi sp. nov.; integrative taxonomy; molecular phylogeny; morphology; Kon Tum-Gia Lai Plateau; Bolaven Plateau; biogeography

Ovophis orlovi sp. nov.

Variation in coloration of Ovophis orlovi sp. nov. in life.
Vietnam: A, Mang Den, Quang Ngai, VRTC NAP-18718, adult female, holotype; B, Kon Chu Rang NR, Gia Lai, ZMMU R-16315, subadult male, paratype; C, Kon Ka Kinh NP, Gia Lai, ZMMU R-15452, subadult female, paratype.
Laos: Bolaven Plateau, Champasak: D, adult individual; E, F, subadult individual.

Photos by A. M. Bragin (A), N. A. Poyarkov (B, C), and A. Teynié (D – F)

Sabira S. Idiiatullina, Darya A. Ustinova, Andrey M. Bragin, Son Xuan Le, Xu Yuhao, Zhang Tierui, Parinya Pawangkhanant, Natalia B. Ananjeva, Patrick David, Gernot Vogel, Tan Van Nguyen and Nikolay A. Poyarkov. 2026. An overlooked Mountain Pitviper from the Montane Regions of Central Vietnam and Southern Laos (Serpentes: Viperidae: Ovophis). Russian Journal of Herpetology. 33(3); 178-196. http://rjh.folium.ru/index.php/rjh/article/view/2315 [September 30, 2026]

 facebook.com/tan.sifasv/posts/122268835112144982

 Researchgate.net/publication/415133263_An_overlooked_Ovophis_from_the_montane_regions_of_Vietnam_Laos

 

 Ovophis orlovi sp. nov., Mang Den Commune, Quang Ngai Province, central Vietnam, June 6, 2016. Photo by Andrey M. Bragin

 

[Arachnida • 2026] Charinus paradoxus• A New whip spider Species (Amblypygi: Charontidae) from Colombia

 

Charinus paradoxus 

Alvarez-Garcia, Suárez-Martínez, & Torres, 2026

DOI: doi.org/10.11646/zootaxa.5886.3.4  

  x.com/LSuarezm_8legs

Abstract

The genus Charinus is currently represented in Colombia by four species: C. miskito, C. mocoa, C. rocamadre, and C. santandereanus. In this study, we describe and illustrate a new species, Charinus paradoxus sp. nov.. This species can be distinguished from its congeners by the absence of median eyes and median ocular tubercle, the presence of three dorsal spines on the pedipalp femur, the first tarsal article of leg I being considerably longer than the second, and the morphology of the glandular openings of the atrium in the female gonopods, which are button-shaped. Additionally, we provide an identification key for the species of Charinus recorded from Colombia and a distribution map of the genus in the country. With the description of this new species, the number of Charinus species known from Colombia increases to five.

Amblypygi, amblypygid, Colombian Caribbean region, morphology, scanning electron microscopy, taxonomy

Charinus paradoxus sp. nov.

Deivys M. ALVAREZ-GARCIA, Leiner A. SUÁREZ-MARTÍNEZ and Richard A. TORRES. 2026. Charinus paradoxus sp. nov. (Amblypygi: Charontidae), A New whip spider Species from Colombia. Zootaxa. 5886(3); 406-422. DOI: doi.org/10.11646/zootaxa.5886.3.4 [2026-10-05]

  x.com/LSuarezm_8legs/status/2107619102122532916

[Arachnida • 2026] Raveniola zhangzongxuni • A New Species of Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from Jinggangshan National Nature Reserve, Jiangxi Province, China

 

Raveniola zhangzongxuni Liu, 

in Li, Liang, Wang et Liu,  2026.

 DOI: doi.org/10.3897/BDJ.14.e211761

Abstract

Background: Prior to this study, only two mygalomorph species from two distinct families had been formally recorded in Jinggangshan National Nature Reserve, Jiangxi Province: Calommata jinggangica Li & Xu, 2022 (Atypidae Thorell, 1870) and Conothele jinggangshan Liu, Xu, Zhang, Liu & Li, 2019 (Halonoproctidae Pocock, 1901), indicating that the overall mygalomorph diversity in this region remains far from fully revealed. The family Nemesiidae Simon, 1889, is known from the neighbouring provinces of Jiangxi: Hunan, Hubei, Fujian and Zhejiang. Nevertheless, no systematic surveys of Nemesiidae have ever been conducted in Jiangxi Province, and to date, no species belonging to this family has been formally reported from the region. During a recent exploratory field survey of local microhabitats in Jinggangshan National Nature Reserve, we collected a series of nemesiid specimens with distinctive morphology. Following morphological examination and preliminary molecular data, these specimens are confirmed to represent a new species of the genus Raveniola Zonstein, 1987. This discovery not only provides the first formal record of Nemesiidae from Jiangxi Province, but also highlights that the mygalomorph diversity in Jiangxi and eastern China remains largely under-investigated, with many potential taxa awaiting discovery.

New information: The spider genus Raveniola Zonstein, 1987 is reported for the first time from Jiangxi Province, China, based on material collected in Jinggangshan National Nature Reserve. A new species, Raveniola zhangzongxuni Liu, sp. nov., is described and illustrated. The discovery provides the first formal record of the family Nemesiidae from Jiangxi Province and bridges the known distributional records of the genus in eastern China, enhancing its geographic continuity on the map. The finding highlights that the mygalomorph diversity in Jiangxi and adjacent regions remains largely under-sampled and contributes to the understanding of the biogeography of the family Nemesiidae in East Asia.

Keywords: East Asia, mygalomorph diversity, distribution, new record

Raveniola zhangzongxuni sp. nov., photos of living specimen.
 A−C male; D−F female.

Raveniola zhangzongxuni Liu, sp. nov.

Diagnosis: Differs from Raveniola chayi Li & Zonstein, 2015 by having the dorsal abdominal pattern consisting of numerous darker spots on a lighter background (vs. numerous lighter spots on a darker background), the maxillae with 3 or 4 cuspules (vs. 12–15), the tibial length/cymbial length ca. 2.5 on male palp (vs. > 3.4), the copulatory bulb is relatively broader ca. 1.2–1.5 times as long as wide (vs. 2.5–4 times) and the lateral diverticulum arising from about 1/3 of spermathecal trunk (vs. about the base of spermathecal trunk). The new species also differs from R. gracilis Li & Zonstein, 2015 by the ventral abdominal pattern consisting of some darker spots on a lighter background (vs. pale yellow, without spots), the male PMS absent (vs. present), the maxillae with 3 or 4 cuspules (vs. 4–6), the tibial length/cymbial length ca. 2.5 on male palp (vs. > 2.6), the copulatory bulb is relatively broader ca. 1.2–1.5 times as long as wide (vs. nearly as long as wide) and the lateral diverticulum approximately 0.6 times as long as spermathecal trunk (vs. approximately 0.3 times).

Etymology: The species is named in honour of Zong-xun Zhang, one of the famous figures of the extremely hard and bitter struggle in Jinggangshan; noun (name) in genitive case.

 

 Congzheng Li, Qinlong Liang, Zhongjing Wang and Ke-Ke Liu. 2026. A New Species of Raveniola Zonstein, 1987 (Araneae, Nemesiidae) from Jinggangshan National Nature Reserve, Jiangxi Province, China. Biodiversity Data Journal. 14: e211761.  DOI: doi.org/10.3897/BDJ.14.e211761 [28-09-2026]

[Herpetology • 2026] Leptobrachella quangninhensis • A New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Quang Ninh Province, Vietnam

Leptobrachella quangninhensis 

Luong, Hoang, Pham, Ziegler & Nguyen, 2026

 

 http://rjh.folium.ru/index.php/rjh/article/view/2250 

 facebook.com/HoangChung2019

Abstract 

A new species of the genus Leptobrachella is described from Quang Ninh Province, Vietnam based on genetic divergence and morphological differences. Leptobrachella quangninhensis sp. nov. is characterized by a combination of the following characters: size medium (SVL 28.1 – 28.7 mm (n = 4) in adult males and 36.0 mm in one adult female); head longer than wide; tympanum distinct; belly creamy white; toe webbing rudimentary and without lateral fringes; supratympanic fold edged by a distinct black line; iris bicolored, copper in upper part, fading to golden in lower part; interorbital region with distinct black triangle; dorsal skin smooth with fine scattered tubercles. In terms of genetic distance, the new species differs from other congeners for which comparable sequences are available by at least 6.15% (16S rRNA gene). This species description brings the total number of known species in the genus Leptobrachella from Vietnam to 38.

Keywords: Leptobrachella quangninhensis sp. nov.; morphology; molecular phylogeny; taxonomy

Leptobrachella quangninhensis sp. nov.  

Anh Mai Luong, Chung Van Hoang, Cuong The Pham, Thomas Ziegler and Truong Quang Nguyen. 2026. A New Species of Leptobrachella Smith 1925 (Amphibia: Anura: Megophryidae) from Quang Ninh Province, Vietnam. Russian Journal of Herpetology. 33(3); 197-216. [September 30, 2026] 

 http://rjh.folium.ru/index.php/rjh/article/view/2250 

 facebook.com/HoangChung2019/posts/28710851798545157

[PaleoMammalogy • 2026] Pampaluctor calibar • A New sabre-toothed thylacosmilid (Metatheria: Thylacosmilidae) from the Pliocene–Pleistocene of southern Buenos Aires Province, Argentina

 

 Pampaluctor calibar

del Campo, Agnolín, Chimento, Boh, Magnussen & De Cianni, 2026

 

DOI: doi.org/10.1080/03115518.2026.2727509 

Illustration by Sebastián Rozadilla

 

Abstract

Thylacosmilids are a clade of metatherian predators known from lower Miocene to upper Pliocene strata of South America. They had highly specialized skulls with enlarged canines. Here, we describe remains of a new thylacosmilid taxon from the Argentine Pampas. This specimen comprises a partial postcranial skeleton from the upper Pliocene–Lower Pleistocene Vorohué Formation near Miramar in Buenos Aires Province. It shares several features supporting referral to Thylacosmilidae including a strong neck, robust forelimbs, and a shortened tibia. However, it shows notable differences from more derived thylacosmilids, especially Thylacosmilus atrox, the only thylacosmilid represented by relatively complete postcranium. The Vorohué Formation taxon differs by having a robust atlas, simpler ventral crest in the fifth cervical vertebra, enlarged medial epicondyle of the humerus, less distally projected capitulum, a more developed process on top of the lateral epicondylar crest, an enlarged radial fossa, deep coronoid fossa bordered by a marked ridge, smaller medial epicondyle on the femur, and a straight tibial shaft. These character states support establishment of a new genus and species. Phylogenetic analysis places the Vorohué Formation taxon as the sister of Thylacosmilus atrox. Functional comparisons suggest powerful neck musculature and forelimbs, coupled with low cursoriality—traits typical of ambush predators. The Vorohué Formation taxon is one of the geologically youngest sparassodonts, and shows that thylacosmilids coexisted not only with early North American placental carnivores (e.g., procyonids), but also probably mustelids and foxes.

 

Keywords: Sparassodontapost, cranium, Vorohué Formation, Metatheria, systematic palaeontology, South America

Systematic palaeontology

Class MAMMALIA Linnaeus, 1758

Infraclass METATHERIA Huxley, 1880

Order SPARASSODONTA Ameghino, 1894

Superfamily BORHYAENOIDEA Simpson, 1930

Clade THYLACOSMILIFORMES Suarez, Goin, Montalvo, Acosta, Cadena & Tomassini, 2025

Family THYLACOSMILIDAE Riggs, 1933

Pampaluctor gen. nov.

Diagnosis: Large thylacosmilid differing from Anachlysictis in possessing a more robust and anteroposteriorly elongate atlas with a prominent dorsal spine, anterior articular facets not protruding beyond the anterior margin of the dorsal arch, smaller atlantal foramen groove extending anterolaterally instead of posterolaterally, concave posterior articular facets, and an anteroposteriorly extended ventral tubercle on the intercentrum. Differs from Thylacosmilus in having a more robust and compact atlas, smaller atlantal foramen groove, more robust humerus with an expanded medial epicondyle extending over more than a fourth of the total width of the distal epiphysis, transversely broad lateral epicondyle, enlarged radial fossa in contact with the coronoid fossa, femur with much smaller medial epicondyle and distal extremity, and tibia with a straight diaphysis. Differs from Dimartinia and Patagosmilus in larger size and stratigraphic age; no overlapping skeletal elements are known for these taxa.

Etymology: Refers to the Argentine Pampas, and luctor (Latin), to wrestle.

Pampaluctor calibar sp. nov.

Etymology: Refers to ‘Calibar’, a famous nineteenth-century gaucho who tracked thieves and rescued lost people in Central Argentina (described by the author and politician Domingo F. Sarmiento in 1845).

Holotype: MPH P0730, incomplete postcranial skeleton including atlas, axis, third, fourth, fifth and sixth cervical vertebrae, scapula, humerus, radius, femur, tibia, two caudal vertebrae and unidentified bone fragments.

Type locality, unit and age: Locality ..., near Miramar city, Buenos Aires Province, Argentina; Vorohué Formation, upper Pliocene–lower Pleistocene.

Artistic construction of Pampaluctor calibar. 

Illustration by Sebastián Rozadilla, courtesy of E. del Campo

Eric N. del Campo, Federico L. Agnolín, Nicolás R. Chimento, Daniel Boh, Mariano Magnussen and Francisco De Cianni. 2026. A New sabre-toothed thylacosmilid (Thylacosmilidae, Metatheria) from the Pliocene–Pleistocene of southern Buenos Aires Province, Argentina. Alcheringa: An Australasian Journal of Palaeontology. DOI: doi.org/10.1080/03115518.2026.2727509  [22 Sep 2026]

[Botany • 2026] Beaumontia laotica (Apocynaceae) • A New Species from central Laos

 

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen, 

in Ma, Xi, Lorphengsy et Shen, 2026.

DOI: doi.org/10.5735/085.063.0130 

Abstract

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen (Apocynaceae) is described as a new species from Bolikhamxai Province, Laos. It closely resembles B. grandiflora, B. murtonii and B. jerdoniana, but differs from them by several features of the leaves, sepals and corolla, and by flowering in November. A full taxonomic description, diagnostic illustration and formal IUCN conservation assessment are presented for the new species. An identification key to all species of Beaumontia is provided.

Beaumontia laotica (from the holotype).
 — A: Branch. — B: Longitudinal section of flower. — C: Style-head and anther, side view. — D: Anther, adaxial view. — E: Anther, abaxial view. — F: Ovary, side view.

Beaumontia laotica (A and B photographed at type locality, 11 November 2025; C–P from the holotype).
— A: Natural habitat. — B: Habit. — C: Branch. — D: Young stem. — E: Leaf, adaxial view. — F: Leaf, abaxial view. — G: Adaxial leaf surface. — H: Abaxial leaf surface. — I: Inflorescence. — J: Bracts, adaxial views. — K: Sepals, adaxial views. — L: Flower, front view. — M: Longitudinal section of corolla showing inner surface and stamens. — N: Pedicel and pistil, side view. — O: Disc and ovary, front view. — P: Ovary, side view.

Beaumontia laotica X.D. Ma, H.C. Xi, S. Lorphengsy & J.Y. Shen, sp. nova

Diagnosis. Beaumontia laotica differs from B. grandiflora, B. murtonii and B. jerdoniana by its lanceolate or narrowly obovate (vs. elliptic or obovate) leaf blade; persistent (vs. deciduous) bracts; glabrous (vs. pubescent) sepals; and glabrous (vs. pubescent or puberulent) corolla tube, lower portion ca. 7.4 cm long (vs. 1.5–2.5 cm long), upper portion abruptly expanding to trumpet-shaped, ca. 2 cm long (vs. obconical or campanulate, 6.5–11 cm long). 

Xing-Da Ma, Hou-Cheng Xi, Sengvilay Lorphengsy and Jian-Yong Shen. 2026. Beaumontia laotica (Apocynaceae), A New Species from central Laos. Annales Botanici Fennici. 63(1), 233-237. DOI: doi.org/10.5735/085.063.0130  (11 September 2026) 

[Paleontology • 2026] Cranial Anatomy of the Middle Eocene Egyptian crocodylian Herugavialis cairensis provides Nnew Insights into gavialoid diversity in the Mediterranean Region

 

Life reconstruction of Herugavialis cairensis depicted with The Great Sphinx and pyramids of Giza in the background. 

Herugavialis gen. nov.
Herugavialis cairensis n. comb. (Müller, 1927)

in Burke, Spiekman, Salem, Saber, Sallam et Mannion, 2026. 

DOI: doi.org/10.1098/rsos.261173

Artwork by Karim Zanaty.

Abstract

Gavialoidea is presently composed of only two extant crocodylian species, the Indian gharial Gavialis gangeticus, and the false gharial Tomistoma schlegelii, both from southeast Asia. However, Gavialoidea displays a rich and near-cosmopolitan diversity in the fossil record, with over 40 species currently recognized. Several of these gavialoids are recovered from Cenozoic deposits in the Mediterranean region, with many assigned historically to the genus Tomistoma. However, recent phylogenetic analyses do not typically recover these species as closely related to the false gharial, with several fossil species in need of taxonomic revision. Here, we present a detailed re-evaluation of one of these species, Tomistoma cairense from the Middle Eocene Mokattam Formation of Egypt, based on the holotype skull and several referred specimens. Supplemented by endocranial anatomy revealed via computed tomography (CT), we identify numerous anatomical differences with T. schlegelii and other gavialoids, including the approximately contemporaneous Egyptian species, Eogavialis africanum and Paratomistoma courti. These differences are reinforced through phylogenetic analysis under maximum parsimony, in which T. cairense is recovered as a gavialoid that does not cluster with T. schlegelii or Mediterranean taxa. We erect a new genus name for T. cairense, with the resultant new combination of Herugavialis cairensis diagnosed by several characteristics, including telescoped orbits. An apparent decline in Mediterranean gavialoid diversity during the Late Eocene–Oligocene is potentially driven by a reduction in habitat, as a result of global cooling and declining eustatic sea level. Warming in the Miocene might have facilitated a subsequent rebound in gavialoid diversity, prior to the extirpation of crocodylians in the Mediterranean region in the Late Miocene, which might have been caused by the Messinian Salinity Crisis. The taxonomic revision of additional Mediterranean gavialoid species should facilitate greater insight into their evolutionary history and demise.

Keywords: Gavialoidea, Mediterranean, Crocodylia, Tomistoma, neuroanatomy, phylogeny

Eusuchia Huxley, 1875

Crocodylia Gmelin, 1788

Gavialoidea Hay, 1930

Herugavialis gen. nov.,

 

Herugavialis cairensis n. comb. Müller, 1927 

Tomistoma cairense (Müller, 1927).

Etymology: Heru, after the Egyptian god (also known as Horus), is often associated with, and possibly the inspiration for, the Great Sphinx of Giza. The latter was sculpted from the Mokattam Formation, from which the type specimen of H. cairensis was recovered, with both sharing the absence of the anterior tip of their nose/snout. The genus suffix, -gavialis, reflects the relationship of H. cairensis to Gavialis gangeticus within Gavialoidea. Accordingly, the original species epithet, cairense, is modified to cairensis to agree with the masculine gender of Herugavialis.

  

Locality and horizon: The holotype and referred specimens were collected from Gebel Mokattam, east of Cairo, in northeastern Egypt. They were recovered from the Mokattam Formation, which is a nummulitic marine limestone dated to the Lutetian, Middle Eocene.

 Diagnosis: Within Gavialoidea, H. cairensis is characterized by a unique combination of characters: (i) lacrimal with an anteroposterior length more than twice that of the prefrontal; (ii) telescoped orbit formed by raised margins of the lacrimal, prefrontal and frontal; (iii) a squamosal wherein the dorsal and ventral margins do not diverge anteriorly; (iv) the distance between the distal tip of the paraoccipital process and the distal end of the quadrate condyle is greater than the maximum mediolateral width of the latter; and (v) a blunt dural peak on the encephalic endocast.

Life reconstruction of Herugavialis cairensis in left lateral view. Herugavialis is depicted with The Great Sphinx and pyramids of Giza in the background.

 Artwork by Karim Zanaty.

 

 Conclusions: 

The redescription of the holotype and referred specimens of the Middle Eocene crocodylian species ‘Tomistoma’ cairense demonstrates numerous morphological differences from extant T. schlegelii. Coupled with the recognition of several autapomorphies, as well as its phylogenetic separation from T. schlegelii, we erect the new genus, Herugavialis cairensis. Herugavialis can be differentiated from approximately contemporaneous Egyptian gavialoids by numerous anatomical features, exemplifying the diversity of gavialoids in the Mediterranean region during this period. Global cooling and a drop in eustatic sea level during the Late Eocene–Oligocene potentially limited suitable habitat for gavialoids in the Mediterranean region, with subsequent increases in eustatic sea level and temperature potentially facilitating the return to high gavialoid diversity observed in the Miocene. The taxonomic classification of some of these European species awaits re-evaluation, which will facilitate greater insight into the evolutionary history of the Mediterranean gavialoids.

Paul M. J. Burke; Stephan N. F. Spiekman; Belal S. Salem; Sara Saber; Hesham M. Sallam and Philip D. Mannion. 2026. Cranial Anatomy of the Middle Eocene Egyptian crocodylian Herugavialis cairensis provides Nnew Insights into gavialoid diversity in the Mediterranean Region. R. Soc. Open Sci 13 (10): 261173. DOI: doi.org/10.1098/rsos.261173 [07 Oct 2026]

[Botany • 2026] Primula octopoda (Primulaceae) • A stoloniferous New Species from western Sichuan, China

 

Primula octopoda W.B.Ju & Q.Huang,  

in Q. Huang, X. Wang, W.-J. Huang, Yang, Pan, Ru, J.-P. Wang et Ju, 2026.

巴塘粉报春 | DOI: doi.org/10.1002/ece3.74088 

 

ABSTRACT

Primula octopoda W.B.Ju & Q.Huang, a new species of Primula (Primulaceae) in sect. Aleuritia, is herein described and illustrated from Batang County, Sichuan Province, southwestern China. Morphologically, P. octopoda is most similar to P. stenocalyx, but it can be readily distinguished by its dwarf habit, the presence of stolons 2–7 cm in length ending in small rosettes occasionally bearing a single flower, elliptic to broadly obovate leaf blades, a very short or absent scape bearing one or two flowers, and a smaller, annulate corolla with a prominent yellow eye. The complete chloroplast genome of P. octopoda spans 154,164 bp with a GC content of 36.97% and contains 125 genes. Phylogenetic analyses using plastid and ITS data both place P. octopoda within sect. Aleuritia while supporting its distinct species status. Given the current lack of adequate information on its population size and geographic range, the species is provisionally assigned a status of Data Deficient (DD) according to the IUCN Red List criteria.

Keywords: molecular phylogeny, morphology, new species, Primula, taxonomy

Habitat of the Primula octopoda.
 (A) Habitat; (B–D) Plants.

Morphology of Primula octopoda.
 (A) Plant; (B) roots; (C) leaves; (D) inflorescence; (E) corolla; (F) bracts; (G) calyx; (H) thrum flower and pin flower; (I) capsule.

Primula octopoda W.B.Ju & Q.Huang, sp. nov. 

Diagnosis. The new species most closely resembles P. stenocalyx, sharing narrowly lanceolate bracts, a tubular calyx with similarly divided oblong or lanceolate lobes, stamens and style included within the corolla tube, and an oblong capsule. However, it differs in its shorter stature, the presence of 2–7 cm long stolons each ending in a small rosette and occasionally bearing a solitary flower, elliptic to broadly obovate leaves, a scape up to 2 cm long or absent with 1–2 flowers, and an annulate corolla with a yellow eye and smaller flowers (Figures 2 and 3).

Etymology. The specific epithet of the new species refers to its distinctive feature of producing multiple stout, fleshy stolons that resemble the tentacles of an octopus; hence the name octopoda, which vividly reflects the plant's numerous stolons extending like “eight legs.”

Vernacular Name. Chinese mandarin: “bā táng fĕn bào chūn” (巴塘粉报春), meaning “Batang Farinose Primrose” in English.

 

Qin Huang, Xin Wang, Wen-Jun Huang, Jie-Yi Yang, Hong-Li Pan, Jian Ru, Jiang-Ping Wang and Wen-Bin Ju. 2026. Primula octopoda, A stoloniferous New Species from Western Sichuan, China. Ecology and Evolution. DOI: doi.org/10.1002/ece3.74088 [27 August 2026]

[Herpetology • 2025] Leptobrachella batxatensis & L. duyenae • Hidden Biodiversity in the Tropical Rain Forests: Two New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Vietnam

Leptobrachella duyenae 

Hoang, Pham, Phan, Do, Bin Wang, Jiang & Nguyen, 2025 

Cóc mày duyên  ||  DOI: doi.org/10.3724/ahr.2095-0357.2024.0039 

 facebook.com/HoangChung2019

Abstract

Two new species of Leptobrachella are described from Vietnam based on morphological differences and genetic divergences in 16S rRNA mitochondrial gene sequences. The new taxa are distinguished from each other and from other species of the genus Leptobrachella in body size, head width/length ratio, tympanum morphology, dorsal skin texture, the presence/absence of fringes on toes, color of dorsal and ventral body, and iris color. The two new species are also divergent from each other and from other congeners by a 4.14% or greater uncorrected genetic distance. Leptobrachella batxatensis sp. nov. is genetically closest to L. shiwandashanensis and L. wuhuangmontis from China. Leptobrachella duyenae sp. nov. is genetically closest to L. bidoupensis from Vietnam with strong nodal support from both BI and ML analyses (1.0/99%).

Keywords: 16S rRNA mitochondrial gene, Leptobrachella batxatensis sp. nov., Leptobrachella duyenae sp. nov., morphology, taxonomy

Variation of Leptobrachella batxatensis sp. nov. in life.
Dorsolateral view of A: Paratype (IEBR A.5224), B: Paratype (IEBR A.5225), C: Paratype (IEBR A.5226), D: Paratype (IEBR A.5227), E: Paratype (IEBR A.5228); and F: Ventral view of Paratype (IEBR A.5228).

Leptobrachella batxatensis sp. nov. 

Etymology: Specific epithet is an adjective in reference to the type locality, the Bat Xat Nature Reserve of Lao Cai Province. 

We recommend “Bat Xat Litter Frog” as the common English name and “Cóc mày bát xát” as the Vietnamese name.

Holotype of Leptobrachella duyenae sp. nov. in life (IEBR A.5229). A: Dorsolateral view; B: Ventral view; C: Left hand; D: Left food; and E: Cloacal area. Scale bars = 10 mm.
Variation of  Leptobrachella duyenae sp. nov. in life. A: Dorsolateral view; B: Ventral view of Paratype female (IEBR A.5233); C: Dorsolateral view; and D: Ventral view of Paratype male (IEBR A.5230).

Leptobrachella duyenae sp. nov.

Etymolog: The species is epithet, a noun that recognizes Mrs. Duyen Thi NGUYEN, for her understanding and strong support of C. V. HOANG’s research. She also assisted with fieldwork and collected the holotype. 

We recommend “Duyen’s Litter Frog” as the common English name and “Cóc mày duyên” as the Vietnamese name.  

Van Chung HOANG, The Cuong PHAM, Quang Tien PHAN, Trong Dang DO, Bin WANG, Jianping JIANG and Quang Truong NGUYEN. 2025. Hidden Biodiversity in the Tropical Rain Forests: Two New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Vietnam. Asian Herpetological Research. 16(2); 169 - 191. DOI: doi.org/10.3724/ahr.2095-0357.2024.0039 [Mar 6, 2025]

 facebook.com/HoangChung2019/posts/25595701346726900

[Mollusca • 2023] Siamochlamys panhai • A New Limestone Karst‐Restricted Terrestrial Snail (Eupulmonata: Helicarionidae) from Thailand

Siamochlamys panhai Pholyotha, 

Pholyotha, Tongkerd et Sutcharit, 2023. 

DOI: doi.org/10.58837/tnh.23.7.258814 

Abstract

A new genus and new species of helicarionid snail, Siamochlamys panhai, is described from limestone outcrops in Thailand. This new genus differs from all helicarionid genera based on morphological characters of shell, radula, and genitalia. The diagnostic characters of Siamochlamys panhai gen. et sp. nov. are depressed shell, well-polished surface, umbilicus opened, and aperture less covered by reflection of columellar margin. The genitalia exhibit a straight epiphallic caecum and small flagellum but lack a dart apparatus; radula has tricuspid central and lateral teeth and bicuspid marginal teeth. The discovery of this karst-associated land snail increases the knowledge of endemic malacofauna in Thailand, and also suggests that many taxa remain undiscovered.

Keywords: endemic, Indochina, land snail, limestones, taxonomy

Superfamily Helicarionoidea Bourguignat, 1877

Family Helicarionidae Bourguignat, 1877

Subfamily Durgellinae Godwin-Austen, 1888

Living snails and mantle extensions of Siamochlamys panhai gen. et sp. nov. paratype CUMZ14211.
A. living snail during relaxed position showing the expansion of mantles B. alcohol-preserved snailshowingshell lobes and dorsal lobes and posterior body C. dorsal view of mantle extensions (shell lobes and dorsal lobes).

Genus Siamochlamys Pholyotha gen. nov.

Etymology.− The name combines ‘Siam’,which is a former name of Thailand,chosen because this snail is restricted to the country as far as we know, and the Greek word ‘chlamys’,meaning “mantle or cloak”. Therefore, the generic name means the land snail with well-developed mantle extensions from Thailand. 

Diagnosis.− Depressed, glossy shellwith open umbilicus, straight epiphallic caecum, short flagellum, no dart apparatus, radula with tricuspidcentral and lateral teeth and bicuspid marginal teeth, andfive mantle extensions well-developed.

Siamochlamys panhai Pholyotha, sp. nov.

Arthit Pholyotha, Piyoros Tongkerd, Chirasak Sutcharit. 2023. Siamochlamys panhai, A New Limestone Karst‐Restricted Terrestrial Snail from Thailand (Eupulmonata: Helicarionidae). Tropical Natural History. Supplement 7;41-50. DOI: doi.org/10.58837/tnh.23.7.258814