[Botany • 2026] Nymphoides crucioides, N. miniata, N. thailandica, ... • Integration of Cytogenetics and Morphology for the Taxonomic Revision of Nymphoides (Menyanthaceae) in Thailand including Five New Species

 

Nymphoides miniata Noppornch. & Suwanph.,

in Suwanphakdee, Nopporncharoenkul et Hodkinson, 2026. 

DOI: doi.org/10.1016/j.aquabot.2026.104044 

facebook.com: Flora of Thailand

Highlights: 

• A full revision of Thai Nymphoides based on morphology and cytogenetic evidence is provided.

• Cytogenetic and morphological evidence supports Nymphoides species boundaries.

• Ten Nymphoides species are described in Thailand, including five new endemic species

Abstract 

A taxonomic revision of Nymphoides (Menyanthaceae) in Thailand was undertaken using cytogenetics and morphology. Specimens were collected throughout Thailand. Cytogenetic analyses were conducted on living collections. The morphological characters used in the descriptions of all taxa were measured from living specimens, which were subsequently maintained for ex-situ conservation. We provide a new generic description, a key to species, and descriptions of all species. Line drawings are provided for the five new species and color photographic illustrations are provided for all taxa. Herbarium voucher specimens were prepared and deposited in Thai herbaria. Ten species are recognized in Thailand, including five new species described in the present study for the first time, namely Nymphoides chumphonense, N. crucioides, N. miniata, N. thailandica and N. thungyaiense. Cytogenetic evidence supports the proposed species boundaries of Thai Nymphoides.

 

Keywords: Aquatic plants, Terrarium plants, Tropical plants, Wetland

Nymphoides chumphonense Suwanph. 

Nymphoides crucioides Suwanph. & Hodk. 

Nymphoides miniata Noppornch. & Suwanph. 

Nymphoides thungyaiense Suwanph & Noppornch. 

 Nymphoides thailandica Suwanph. 

 

Chalermpol Suwanphakdee, Nattapon Nopporncharoenkul and Trevor R. Hodkinson. 2026. Integration of Cytogenetics and Morphology for the Taxonomic Revision of Nymphoides (Menyanthaceae) in Thailand including Five New Species. Aquatic Botany. 207, 104044. DOI: doi.org/10.1016/j.aquabot.2026.104044 

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🌸 มหาวิทยาลัยเกษตรศาสตร์ พบบัวบาชนิดใหม่ของโลก 5 ชนิด จากแหล่งน้ำของประเทศไทย บ่งชี้ถึงความร่ำรวยทรัพยากรพืชของไทย และความสำคัญของงานวิจัยวิทยาศาสตร์พื้นฐาน ของสถาบันการศึกษา💚

นักวิจัย มก นำโดย รศ.ดร. เฉลิมพล สุวรรณภักดี จากภาควิชาพฤกษศาสตร์ คณะวิทยาศาสตร์ มหาวิทยาลัยเกษตรศาสตร์ ค้นพบบัวบาชนิดใหม่ของโลกจำนวน 5 ชนิด โดยตีพิมพ์เผยแพร่การค้นพบนี้ในวารสาร Aquatic Botany ฉบับปี 2026 การค้นพบครั้งนี้เป็นการบูรณาการวิจัยร่วมระหว่างหน่วยงานซึ่งประกอบด้วย ดร. ณัฐพล นพพรเจริญกุล นักวิจัย จากองค์การพิพิธภัณฑ์วิทยาศาสตร์แห่งชาติ (อพวช) และ Prof. Dr. Trevor R. Hodkinson จากทรินิตี้คอลเลจ มหาวิทยาลัยแห่งกรุงดับลิน ประเทศไอร์แลนด์ โดยเป็นการบูรณาการวิจัยร่วมทางด้านอนุกรมวิธานพืช สัณฐานวิทยาและเซลล์วิทยา 

.

รศ.ดร. เฉลิมพล สุวรรณภักดี  กล่าวว่า พืชสกุลบัวบา (Nymphoides) วงศ์ Menyanthaceae เป็นพืชน้ำที่มีการใช้ประโยชน์ทางด้านการใช้เป็นพืชประดับ และผักพื้นบ้าน ในประเทศไทยเคยมีรายงาน 5 ชนิด จากการศึกษาวิจัยพบว่า บัวบาของประเทศไทยมีทั้งหมด 10 ชนิด และเป็นพืชชนิดใหม่ของโลก 5 ชนิด แต่ละชนิดมีศักยภาพในการนำไปใช้ประโยชน์ทางด้านการใช้เป็นพืชประดับสวนขวด สวนกระจก และตู้ไม้น้ำ อีกทั้งยังเน้นให้ตระหนักถึงการอนุรักษ์แหล่งน้ำจืด ซึ่งเป็นที่อยู่ของพืชชนิดเหล่านี้ บ่งชี้ถึงความร่ำรวยทรัพยากรพืชของประเทศไทย และความสำคัญของงานวิจัยวิทยาศาสตร์พื้นฐาน 

บัวบา 5 ชนิดนี้ประกอบด้วย 

.

1. สายติ้ง หรือ สายติ่ง Nymphoides chumphonense Suwanph. 

 พบในแหล่งน้ำจืดตื้นๆ บริเวณ จ. ชุมพร และสุราษฎร์ธานี  โดยตั้งชื่อตาม จ. ชุมพร สถานที่พบตัวอย่างเป็นที่แรก พืชชนิดนี้มีการใช้ประโยชน์เป็นผักพื้นบ้าน โดยกินสดกับน้ำพริก และมีขายในตลาด ถูกจัดสถานะใกล้สูญพันธุ์ (Endangered: EN) เนื่องจากการสูญเสียแหล่งที่อยู่ และการรุกรานของน้ำเค็ม อีกทั้งติดผลและเมล็ดได้น้อย

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2. บัวบาหัวลูกศร Nymphoides crucioides Suwanph. & Hodk. 

 พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี และยะลา โดยตั้งชื่อตามลักษณะดอกที่คล้ายกากบาท  พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

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3. บาจิ๋ว Nymphoides miniata Noppornch. & Suwanph. 

 เป็นบัวบาที่มีขนาดเล็กที่สุดของโลก พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี โดยตั้งชื่อตามลักษณะดอกที่มีขนาดเล็กที่สุดของโลก พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

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4. บาสยาม Nymphoides thailandica Suwanph. 

บัวบาที่มีขนาดเล็ก พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี กระบี่ และยะลา โดยตั้งชื่อตามประเทศไทย พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

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5. ดาวรดา Nymphoides thungyaiense Suwanph & Noppornch. 

เป็นบัวบาที่มีขนาดค่อนข้างใหญ่ พบในแหล่งน้ำจืดตื้นๆ บริเวณ จ. ตาก โดยตั้งชื่อตามแหล่งที่พบ ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากพบประชากรเพียงแหล่งเดียว  

[Cnidaria • 2026] Hemicorallium osmanthogemmum • A New Species of Hemicorallium (Anthozoa: Octocorallia: Coralliidae), along with a revised identification key to species of the genus

Hemicorallium osmanthogemmum

Koido, 2026 

 

DOI: doi.org/10.3897/zookeys.1277.184620 

 

Abstract

A new precious coral species, Hemicorallium osmanthogemmum sp. nov. (Scleralcyonacea, Coralliidae), retrieved from a depth of 550 m in Kochi Prefecture, Japan, is described, and an updated identification key for the genus Hemicorallium is provided. The new species is confirmed as belonging to the genus Hemicorallium based on the presence of nonretractile but contractile autozooids with an ovate-cylindrical shape and sclerites comprising elongated rods, 6–8-radiates and double clubs, with elongated rods present exclusively in the tentacles. Moreover, the species is distinct from all its congeners in that it has orange coenenchyme and a pink axis, with autozooids distributed only on the front side of the branch, forming clusters at the branch tips. These morphological features differ from those of all previously described species of Hemicorallium, and molecular analysis of mitochondrial sequences (16S, ND1, ND2, ND3, ND6, COI, MSH, and IGR1) further corroborates the validity of this new species. This is the first report of Hemicorallium off the coast of Kochi Prefecture, Japan.

Key words: Cnidaria, deep-sea corals, precious coral, Scleralcyonacea, Western Pacific

 

Hemicorallium osmanthogemmum sp. nov. (Holotype: KBF-OA-00436).
 A. ‘‘Front’’ and ‘‘back’’ of colony; B. Branch, showing autozooid (AZ); C. Exposed axis with the coenenchyme removed; D, E. Surface detail of branch, showing autozooid (AZ), siphonozooid (SZ), burrows (Bu), cluster of autozooids (CL), warts on coenenchyme surface (W), and commensal polychaete (Po).

 Scale bars: 50 mm (A); 10 mm (B–E).

Class Octocorallia Haeckel, 1866

Order Scleralcyonacea McFadden, van Ofwegen & Quattrini, 2022

Family Coralliidae Lamouroux, 1812

Genus Hemicorallium Gray, 1867

Hemicorallium osmanthogemmum sp. nov. 

 New Japanese name. Kinmokusei-mizo-sango.

Diagnosis. The colony branches in one plane, and anastomoses are present only where broken branches come into contact with other branches. Autozooids are contracted but not retracted, and they are distributed only on one side of the colony. Spacing is almost absent on the twigs, and clusters are formed at the tips. The axis has no pits beneath the autozooids and is pale pink to pink in colour. Sclerites are predominantly radiate, and double clubs are rare. Warty rods are present only in the tentacles.

 

Tatsuki Koido. 2026. A New Species of Hemicorallium (Anthozoa, Octocorallia, Coralliidae), along with a revised identification key to species of the genus. ZooKeys. 1277: 339-354.  DOI: doi.org/10.3897/zookeys.1277.184620 [20 Apr 2026]

[Arachnida • 2026] Orchestina nojimai & O. insulana • Taxonomic Revision of the Jumping Goblin Spider Genus Orchestina Simon, 1882 (Araneae: Oonopidae) in Japan with Descriptions of Two New Species

A–B. Orchestina flava Ono, 2005, females. 
C. Orchestina insulana sp. nov., female; D. O. sanguinea Oi, 1955;
Orchestina saltitans Banks, 1894; O. okitsui Oi, 1958, male. 

Suzuki, 2026

 DOI: doi.org/10.5852/ejt.2026.1068.3293 

Abstract 

The goblin spider genus Orchestina Simon, 1882 (Araneae: Oonopidae) in Japan has been revised, recognizing nine species including two new species and four species newly recorded from Japan: O. okitsui Oi, 1958 (♂♀), O. flava Ono, 2005 (♂♀), O. sanguinea Oi, 1955 (♀), O. saaristoi Henrard & Jocqué, 2012 (♂♀, new record), O. saltitans Banks, 1894 (♂♀, new record), O. colubrina Liu, Henrard & Xu, 2019 (♂, new record), O. pavesii (Simon, 1873) (♂♀, new record), Orchestina nojimai sp. nov. (♀) and O. insulana sp. nov. (♀). A literature survey revealed errors in the correspondences between figures and captions in a pictorial book, which likely led to the misidentification of the synanthropically introduced species O. saltitans as the forest-dwelling native species O. okitsui. The misrepresentation also gave rise to the erroneous assumption that the male of O. sanguinea had been described, although no male specimen has actually been reported. In addition, Orchestina flagella Saaristo & van Harten, 2006, is here synonymized with O. saltitans (O. flagella syn. nov.). This study provides photographs and illustrations of nine Orchestina species, including the type specimens of O. okitsui, O. flava and O. sanguinea, notes on their habitats, distribution maps, supplemental molecular data, and a graphical character matrix of Orchestina species in Japan.

Keywords: Arachnida, misidentification, type material, synanthropic species, Synspermiata

Live specimens of species of Orchestina Simon, 1882.
A. Orchestina okitsui Oi, 1958, male (YSPC-Op-1). B. Orchestina saltitans Banks, 1894, male (YSPC-Op-2).

Live specimens of species of Orchestina Simon, 1882.
 A–B. Orchestina flava Ono, 2005, females (TKPM-AR, voucher specimen numbers not specified). C. Orchestina insulana sp. nov., female (TKPM-AR 3489). D. Orchestina sanguinea Oi, 1955, female (TKPM-AR 3491).

 

Yuya SUZUKI. 2026. Taxonomic Revision of the Jumping Goblin Spider Genus Orchestina Simon, 1882 (Araneae: Oonopidae) in Japan with Descriptions of Two New Species. European Journal of Taxonomy.  1068: 1–49. DOI: doi.org/10.5852/ejt.2026.1068.3293 [2026-06-12]

[Entomology • 2026] Kongobatha spinosistyla, K. serpens & K. rufilinea • Striking, Slender, and Secretly Spinose: A Revision of the Snake Mantises of the Genus Kongobatha (Mantodea: Nanomantidae: Fulciniinae: Neomantini)

Kongobatha spinosistyla Connors, 

in Connors, Yeeles, Lach et  Rentz, 2026

DOI: doi.org/10.11646/zootaxa.5807.1.2

 

Abstract

Kongobatha is one of the most commonly encountered of all Australian mantis genera, and yet despite this, very little is known about the taxonomy or biology of the genus. Described to include a single Australian species, K. diademata, the only subsequent work on the genus has been the description of a second species from New Guinea, K. papua. We here describe three additional species, K. spinosistyla Connors sp. nov., K. serpens Connors sp. nov., and K. rufilinea Connors sp. nov., and redescribe K. diademata and K. papua in detail, the latter of which is recorded from Australia for the first time. We also describe for the first time the unusual, heavily spinose styli of male Kongobatha. These are apparently unique among Mantodea as a whole, but their function remains unknown. 

Mantodea, mantis, snake mantis, Australia, New Guinea, taxonomy, styli, citizen science

Male and female Kongobatha spinosistyla (Rainforest Snake Mantis), Kuranda, North Queensland.

photo: Maurice Allan

MATTHEW G. CONNORS, PETER YEELES, LORI LACH and DAVID C.F. RENTZ. 2026. Striking, Slender, and Secretly Spinose: A Revision of the Snake Mantises of the Genus Kongobatha (Mantodea: Nanomantidae: Fulciniinae: Neomantini).  Zootaxa. 5807(1); 45-84. DOI: doi.org/10.11646/zootaxa.5807.1.2 [2026-05-08]

[Entomology • 2026] Stygioides hecate • A Golden Shadow in the Mountains: A New Stygioides Bruand, 1853 (Lepidoptera: Cossidae) from Turkey

 Stygioides hecate Japaridze & Junnilainen, 

in Japaridze, Junnilainen et Sihvonen, 2026.  

 DOI: doi.org/10.11646/zootaxa.5821.2.8

 

Abstract

A new species of the genus Stygioides Bruand, 1853 (Lepidoptera: Cossidae) is described from central Turkey. Stygioides hecate Japaridze & Junnilainen sp. nov. is based on material collected in the Sultan Daðlarý Mountains (Konya Province). The new species is characterised by its distinct wing coloration in females. Adults are diurnal, as in other members of the genus, and were collected by daytime netting. DNA barcode data (COI) are provided and compared with available sequences in the Barcode of Life Data Systems (BOLD Systems), supporting the distinctiveness of the new taxon. The discovery of this species increases the number of Stygioides species to 10 and those from Turkey to three, thus contributing to a better understanding of the diversity and distribution of the genus in the Palaearctic realm.

Lepidoptera, carpenter moths, DNA barcoding, Heteroneura, Middle East

Adults of Stygioides hecate Japaridze & Junnilainen sp. nov.
 A–B. Paratype male, dorsal and ventral view and associated labels.
C–D. Holotype female, dorsal and ventral view and associated labels.

Scale bar: 10 mm

Stygioides hecate Japaridze & Junnilainen sp. nov. 

Diagnosis: Males of S. hecate Japaridze & Junnilainen sp. nov. differ from S. nupponenorum by the presence of a distinct white tip of the male abdomen and white scales on the antennae, the absence of brown piliform scales on the abdomen, a narrower apex of the valvae, and a shorter saccus (Figs 1A–1B, 2 C; cf. Lingenhöle et al. 2016: fig. 6; Yakovlev 2011: fig. 35). Females are readily distinguished from all congeners by the presence of golden-yellow scales on the antennae, head, thorax and wings (Figs 1C–1D; cf. Lingenhöle et al. 2017: figs 1–3, 9; Japaridze et al. 2025b: fig. 22).

Etymology: The species name is derived from Hecate, the ancient Greek goddess associated with the night and commonly depicted carrying yellow or golden torches. This name alludes to the coloration of the holotype female, which is predominantly black with yellow-golden scales on the wings, thorax, head, and antennae.

LASHA-GIORGI JAPARIDZE, JARI JUNNILAINEN and PASI SIHVONEN. 2026. A Golden Shadow in the Mountains: A New Stygioides Bruand, 1853 (Lepidoptera: Cossidae) from Turkey.  Zootaxa. 5821(2); 274-280. DOI: doi.org/10.11646/zootaxa.5821.2.8 [2026-05-29]

[Arachnida • 2026] Kambiwa itacarambi, K. maracas, Sertana capivara, S. sagarana, ... • Ninetine Spiders in Brazilian Caatinga and Cerrado: Revision of Kambiwa and Description of Sertana gen. nov. (Araneae: Pholcidae), with analyses of predicted range shifts due to climate change

A–B. Kambiwa itacarambi Huber sp. nov., ♂♀ with egg sac from NW of Itacarambi.
E–F. K. coribe Huber sp. nov., ♂♀ with egg sac from E of São Felix do Coribe.
G–H. K. maracas Huber sp. nov., ♂♀ with egg sac from S of Contendas do Sincorá.
K–L. K. mucuge Huber sp. nov., ♂♀ from NE of Mucugê.

in Huber, Meng, Král, Herrera et Carvalho., 2026. 

DOI: doi.org/10.5852/ejt.2026.1054.3276 

Abstract

Among daddy long-legs spiders (Pholcidae), Ninetinae is a distinctive subfamily that comprises short-legged, fast-running spiders. Most species are small or tiny, lead reclusive lives, and are largely restricted to semiarid regions, which together has made them poorly collected and poorly known. Here, we build on focused recent collections in the Brazilian Cerrado and Caatinga biomes, two of the World’s richest tropical savanna, xeric shrubland and thorn forest regions. Our focus is on the taxonomy of the genus Kambiwa Huber, 2000 that previously contained only two nominal species, each known from a single locality. Combining morphological and molecular (CO1 barcode) data, we describe six new species in Kambiwa (K. brumado Huber sp. nov.; K. coribe Huber sp. nov.; K. ibo Huber sp. nov.; K. itacarambi Huber sp. nov.; K. maracas Huber sp. nov.; K. mucuge Huber sp. nov.), redescribe the type species K. neotropica (Kraus, 1957), and synonymize the monotypic genus Pemona Huber, 2019 with Kambiwa, resulting in the new combination K. sapo (Huber, 2019) comb. nov. In addition, we describe a new genus of superficially Kambiwa-like spiders from the same geographic region: Sertana Huber gen. nov., with five new species (S. bumba Huber gen. et sp. nov.; S. capivara Huber gen. et sp. nov.; S. igapora Huber gen. et sp. nov.; S. lapa Huber gen. et sp. nov.; S. sagarana Huber gen. et sp. nov.). In line with previous efforts to explore the processes underlying the geographical distribution of Ninetinae, we also evaluate the potential effects of future climate change on the environmental niche occupied by three selected species of Kambiwa. Our results corroborate previous findings that demonstrate an altitude-mediated response to climate change. For a highland species, areas of high habitat suitability almost disappear under more severe climate change scenarios. For two species with lowland records, the areas with high habitat suitability increase significantly. Finally, we analyze the male karyotype of K. ibo which consists of 28 chromosomes including a X1X2X3Y system. All chromosomes are biarmed except for the Y chromosome. This contribution concludes a series of publications on the subfamily Ninetinae. We use this opportunity to summarize current knowledge about the subfamily, to discuss open questions and knowledge gaps, and to suggest further research topics focusing on these tiny but exceptional pholcids.

Keywords: CO1 barcodes, climate change, karyotype, niche modeling, taxonomy

Kambiwa Huber, 2000; live specimens from Brazil; all at same scale.
A–B. Kambiwa itacarambi Huber sp. nov., ♂♀ with egg sac from NW of Itacarambi. C–D. K. ibo Huber sp. nov., ♂♀ with egg sac from NE of Lagoa Grande. E–F. K. coribe Huber sp. nov., ♂♀ with egg sac from E of São Felix do Coribe.
G–H. K. maracas Huber sp. nov., ♂♀ with egg sac from S of Contendas do Sincorá. I–J. K. brumado Huber sp. nov., ♂♀ with egg sac from W of Marcolino Moura. K–L. K. mucuge Huber sp. nov., ♂♀ from NE of Mucugê.

Kambiwa Huber, 2000 spp., typical habitats; all localities are in Brazil.
A. Bahia, between Ibó and Curaçá; Kambiwa sp. aff. neotropica. B. Bahia, E of São Felix do Coribe; type locality of Kambiwa coribe Huber sp. nov. 
C. Bahia, NE of Brumado; type locality of K. brumado Huber sp. nov. D. Bahia, SW of Maracas; type locality of K. maracas Huber sp. nov.
E. Bahia, NE of Mucugê; type locality of K. mucuge Huber sp. nov. F. Minas Gerais, NW of Itacarambi; type locality of K. itacarambi Huber sp. nov.

Kambiwa brumado Huber sp. nov.

 K. coribe Huber sp. nov.

K. ibo Huber sp. nov.

K. itacarambi Huber sp. nov.

 K. maracas Huber sp. nov.

 K. mucuge Huber sp. nov.

K. sapo (Huber, 2019) comb. nov.  

 Sertana Huber gen. nov. 

S. bumba Huber gen. et sp. nov. 

 S. capivara Huber gen. et sp. nov. 

 S. igapora Huber gen. et sp. nov.

S. lapa Huber gen. et sp. nov.

 S. sagarana Huber gen. et sp. nov. 

 

Bernhard A. Huber, Guanliang Meng, Jiří Král, Ivalú M. Ávila Herrera, Leonardo S. Carvalho. 2026. Ninetine Spiders in Brazilian Caatinga and Cerrado: Revision of Kambiwa and Description of Sertana gen. nov. (Araneae, Pholcidae), with analyses of predicted range shifts due to climate change. European Journal of Taxonomy. 1054(1), 1–126. DOI: doi.org/10.5852/ejt.2026.1054.3276 [2026-05-14]

[Crustacea • 2026] Frodromia elegans, F. granulosa, ... • Revision of Frodromia McLay, 1993 (Brachyura: Dromiidae) and the Status of Frodromiinae Števčić, 2005

A, Frodromia atypica (Sakai, 1936),; B, F. reticulata (Sakai, 1974);
C, F. elegans, new species, holotype male, Taiwan; D, F. granulosa, new species, holotype male, Vanuatu;
 E, F, F. iners, new species, holotype male, Philippines. 

Ng, 2026 

RAFFLES BULLETIN OF ZOOLOGY. 74: 

Photographs: C, T.-Y. Chan; D, J.C.E. Mendoza; E, F, P.K.L. Ng.

Abstract

 The taxonomy of the West Pacific dromiid genus, Frodromia McLay, 1993, is revised using a suite of characters pertaining to the carapace form and armature, proportions of the ambulatory dactylus, and structures of the pereopods, thoracic sternum, male pleon, and gonopods. The identity of the poorly known type species, F. atypica (Sakai, 1936), is clarified and the species is rediagnosed and illustrated based on the single non-type dried specimen available. Frodromia reticulata (Sakai, 1974) is also rediagnosed on the basis of a series of Japanese specimens. The material identified as F. atypica by McLay (1993) from New Caledonia and the Indonesian Moluccas is here shown to belong to two new species, F. granulosa and F. caileani, respectively; with F. granulosa also reported from Vanuatu. Two other new species, F. elegans and F. iners, are described from the South China Sea and the Philippines, respectively. As a result, Frodromia is hereby classified in the subfamily Frodromiinae Števčić, 2005, which was originally established as a tribe in Sphaerodromiinae Guinot & Tavares, 2003, and which is here redefined and rediagnosed.

Key words. sponge crab, Dromioidea, systematics, subfamily definition, new species, key to taxa, Indo-West Pacific

Colour in life. A, Frodromia atypica (Sakai, 1936), after Sakai (1936: pl. 2 fig. 1); B, F. reticulata (Sakai, 1974), after Sakai (1976: pl. 5 fig. 1);
C, F. elegans, new species, holotype male (6.6 × 7.5 mm) (ZRC 2024.0144), Taiwan; D, F. granulosa, new species, holotype male (6.5 × 7.1 mm) (ZRC 2024.0517), Vanuatu;
E, F, F. iners, new species, holotype male (6.3 × 7.0 mm) (ZRC 2024.0518), Philippines [partly covered with sand particles and debris].

Photographs: C, T.-Y. Chan; D, J.C.E. Mendoza; E, F, P.K.L. Ng.

Frodromia granulosa 

F. caileani 

F. elegans 

 F. iners

PETER K. L. NG. 2026. Revision of Frodromia McLay, 1993 (Crustacea: Brachyura: Dromiidae) and the Status of Frodromiinae Števčić, 2005. RAFFLES BULLETIN OF ZOOLOGY. 74: 332–358. DOI: 10.26107/RBZ-2026-0023  [4 May 2026]

[Cnidaria • 2026] Rhytisma acoronatum, R. calyaceum, R. karibu, ... • Phylogenomics-guided Revision of the Genus Rhytisma Alderslade, 2000 (Octocorallia: Malacalcyonacea: Lemnaliidae), with Descriptions of Six New Sspecies

 

(a, b) Rhytisma karibu sp. nov., 

(c, d) R. acoronatum sp. nov. 

(e, f) R. oblongum sp. nov.

 (c) R. inaequale sp. nov., with expanded polyps 

(d) R. calyaceum sp. nov., with partially expanded polyps 

(e) R. sperkolae sp. nov., polyps partly expanded

Samimi-Namin, Benayahu, Durkin, Ekins, Quattrini & McFadden, 2026

DOI: doi.org/10.1071/IS25068 

Photographs by Y. Benayahu, and M. Ekins.

 

The genus Rhytisma Alderslade, 2000 (Octocorallia: Malacalcyonacea: Lemnaliidae), formerly comprising four nominal species (R. fulvum, R. fuscum, R. monticulum and R. rubiginosum), is revised using an integrative approach. We combine morphological and phylogenomic data for newly collected and historical specimens. A neotype is designated for R. fulvum and a lectotype for R. fuscum to stabilise the application of these names. Six new species are described from the Indo-Pacific: R. acoronatum sp. nov., R. calyaceum sp. nov., R. oblongum sp. nov., R. inaequale sp. nov., R. karibu sp. nov. and R. sperkolae sp. nov. Species delimitation is supported by discrete combinations of morphological characters – particularly those of the tentacle and polyp sclerites – as well as multi-locus DNA barcoding and phylogenomic analyses of conserved elements (UCE and exon loci). Our findings highlight the diagnostic value of tentacle sclerites and reveal extensive species-level diversity that was previously obscured by insufficient morphological examination. The revised genus currently comprises 10 valid species, many of which display restricted geographic distributions, reflecting patterns of regional endemism in Indo-Pacific octocoral assemblages. These results underscore the importance of integrative taxonomy in uncovering hidden biodiversity.

 

Keywords: biogeography, DNA barcoding, endemism, Indo-Pacific, integrative taxonomy, neotype, soft corals, species delimitation, taxonomic revision, ultraconserved elements

Underwater images of Rhytisma species in their respective type localities.
(a, b) Rhytisma fulvum yellow colour and grey colour morphs, polyps partly expanded
(c) R. inaequale sp. nov., with expanded polyps (d) R. calyaceum sp. nov., with partially expanded polyps
(e) R. sperkolae sp. nov., mat morphology reflects underlying irregular reef substrate, polyps partly expanded. (f) R. monticulum with expanded polyps.

Photographs by (a–c, e) Y. Benayahu, (d, f) M. Ekins.

Underwater images of Rhytisma species in their respective type localities.
(a, b) Rhytisma karibu sp. nov., growing on seagrass blades and reef substrate (c, d) R. acoronatum sp. nov. (e, f) R. oblongum sp. nov. 

Photographs by (a, b, e, f) Y. Benayahu, and (c, d) M. Ekins.

Rhytisma acoronatum sp. nov., R. calyaceum sp. nov., 

R. oblongum sp. nov., R. inaequale sp. nov., 

R. karibu sp. nov. and R. sperkolae sp. nov. 

Kaveh Samimi-Namin, Yehuda Benayahu, Kathleen M. Durkin, Merrick Ekins, Andrea M. Quattrini and Catherine S. McFadden. 2026. Phylogenomics-guided Revision of the Genus Rhytisma Alderslade, 2000 (Octocorallia: Malacalcyonacea: Lemnaliidae), with Descriptions of Six New Sspecies. Invertebr Syst. 40(3); IS25068. DOI: doi.org/10.1071/IS25068   [30 March 2026]

[Herpetology • 2021] Hemidactylus cinganji, H. faustus, ... • Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820 (Squamata, Gekkonidae), with Description of Four New Species

  Hemidactylus cinganji
H. carivoensis
H. faustus 
H. pfindaensis 

 Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Vaz Pinto, 2021 

       

 DOI: 10.3897/vz.71.e64781 

 facebook.com/JavierLobonRovira

 

Abstract

The taxonomy of Angolan Hemidactylus has recently been revised. However, the lack of fresh material for some groups and regions, has led to the misidentification of some taxa and an underestimation of actual diversity in others. To shed light on the evolutionary history and systematics of Angolan Hemidactylus, we generated a new phylogenetic hypothesis for the group, and updated the taxonomy following an integrative approach. This resulted in the description of four new species (Hemidactylus pfindaensis sp. nov., H. faustus sp. nov., H. carivoensis sp. nov. and H. cinganji sp. nov.), the reevaluation of two recently described species (H. vernayi and H. paivae) and the synonymization of a recently described species (H. hannahsabinnae). We estimate divergence times for these lineages, providing a preliminary interpretation of their speciation process. Moreover, we suggest and outline 13 Angolan Main Biogeographic Units (AMBUs) in the area, defining a new biogeographic context for future works on Angolan herpetofauna. We now recognize eleven Hemidactylus species in Angola, and we provide here a new morphological key for Angolan Hemidactylus to help with identifications and species assignments in this group.

Keywords: Angola, biogeography, Gekkota, Hemidactylus, systematics, taxonomy

Hemidactylus bayonii-group 

Hemidactylus vernayi Ceríaco, Agarwal, Marques and Bauer 2020

Hemidactylus pfindaensis sp. nov.

Etymology: The name “pfindaensis” derives from the local word “pfinda” which in Kikongo – the main language used in Uíge Province and northwestern Angola – refers to a “gallery forest” or a “continuous block of thick forest”, the main habitat type associated with the species.

Hemidactylus longicephalus-group

Hemidactylus paivae Ceríaco, Agarwal, Marques and Bauer 2020

A – From top, dorsolateral view of Hemidactylus cinganji sp. nov. (FKH0435), H. benguellensis (FKH0413) and H. carivoensis sp. nov. (MNCN 50543);
B – records of H. benguellensis–group (purple color represents records of H. cinganji sp. nov.; blue H. benguellensis; dark green H. carivoensis sp. nov.; circles represent genetically assigned records, while triangles represent historical or non-genetically assigned records by Ceríaco et al. 2020a; color stars depict type or neotype localities; black star depicts original type locality of H. benguellensis assigned by Bocage (1893);
C – habitat of H. cinganji sp. nov. at Ebanga; D – habitat of H. carivoensis sp. nov between Dombe–Echimina. E – habitat of H. benguellensis at Tchivinguiro.

 Photos Javier Lobón–Rovira (A–D) and William R. Branch (E).

Hemidactylus benguellensis-group

Hemidactylus cinganji sp. nov.

Etymology: The name “cinganji” is a widespread traditional word used in Angolan local languages that represents an ancestral spiritual entity that reincarnates assuming different physical forms in different places and occasions. This name is suitable as the new species corresponds to a taxon that was first described under a different name, then became lost and now resurfaces after its original name had been hijacked by a surrogate sister-species. The species epithet is used as a neuter singular noun in opposition to the generic name.

Hemidactylus carivoensis sp. nov.

Etymology: The species epithet “carivoensis” refers to the Farm Carivo, an old estate situated along the banks of the mid-lower Coporolo River on the coastal plain of Benguela Province, and where most of the type series was collected. The species proved to be common in the area, and by recognizing the farm, we also acknowledge the ongoing support from the owners to researchers, similar to the Chapmans nearly a century ago.

Species not assignable to specific species groups

Hemidactylus faustus sp. nov.   

  

Etymology: The name “faustus” applies to a Latin word that designate ‘good luck’, evoking the serendipitous discovery of this species. The species epithet is used as a masculine adjective singular. The first specimen was found by Beatriz Vaz Pinto, daughter of PVP, under a small stone which was removed while preparing a campsite. This unexpected find led to further collecting of this new and previously unrecorded form, albeit from a locality that had been regularly surveyed since the mid-19th century.  

 

 Javier Lobón-Rovira, Werner Conradie, David Buckley Iglesias, Raffael Ernst, Luis Veríssimo, Ninda Baptista and Pedro Vaz Pinto. 2021. Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820, with Description of Four New Species. Vertebrate Zoology. 71: 465-501. DOI: 10.3897/vz.71.e64781

  facebook.com/JavierLobonRovira/posts/3969307526513233

[Entomology • 2026] Staphylus neideae, S. ricardoi, ... • Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera: Hesperiidae: Pyrginae), with Descriptions of Four New Species

[M–P] Staphylus (C.) nicoleae Lemes, sp. nov.; [Q, R] S. (C.) neivae sp. nov.; 
[S, T] S. (C.) ricardoi Lemes, sp. nov.; [U–X] S. (C.) neideae Lemes, sp. nov.: 

in Lemes, Siewert, Mielke, Casagrande et Warren, 2026. 

DOI: doi.org/10.3897/asp.84.e179328 

Abstract

The taxonomy of the subgenus Staphylus (Capilla) is reviewed, including redescriptions of known species, identification keys, and detailed distribution maps. The female genitalia of Staphylus (Capilla) azteca (Scudder, 1872), S. (C.) caribbea (Williams & Bell, 1940), S. (C.) corumba (Williams & Bell, 1940), S. (C.) eryx Evans, 1953, S. (C.) tucumanus (Plötz, 1884) and S. (C.) tyro (Mabille, 1878) are described and illustrated for the first time. Four new species are described: Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, S. (C.) ricardoi Lemes, sp. nov. from Peru, S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil. Pholisora imperspicua Hayward, 1940 is a syn. nov. of Staphylus (C.) lizeri lizeri (Hayward, 1938) and Hesperia melangon epicaste Mabille, 1903 is a syn. nov. of Staphylus (C.) melangon melangon (Mabille, 1883). Neotypes are designated for Staphylus epicaste Mabille, 1903, Nisoniades tucumanus Plötz, 1884 and Staphylus fascia Hayward, 1933. Lectotypes are designated for Pholisora azteca Scudder, 1872, Bolla machuca Schaus, 1913, Helias tyro Mabille, 1878, Staphylus anginus Schaus, 1902, Hesperia melangon 1883, Hesperia musculus Burmeister, 1875 and Helias aurocapilla Staudinger, 1876.

Keywords: Butterflies, Cytochrome oxidase subunit 1, Neotropical, Skippers

Species of Staphylus (Capilla) in dorsal and ventral views.  
A–D S. (C.) ascalon: A, B male (Brazil, São Paulo, Teodoro Sampaio, Morro do Diabo, OM 17.996); C, D female (Brazil, Minas Gerais, Passa Quatro, DZ 46.813).
E, F S. (C.) buena: male (Bolivia, Santa Cruz, Buenavista, holotype, CMNH 7188).
G–J S. (C.) caribbea: G, H male (Costa Rica, Limón Province, La Bomba, MGCL 1112082); I, J female (Costa Rica, Limón Province, Germania, MGCL 1112078).
K–N S. (C.) eryx: K, L male (Brazil, Pará, Santarém, DZ 46.566); M, N female (Brazil, Pará, Santarém, MGCL 1112109).
O–R S. (C.) corumba: O, P male (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.033); Q, R female (Brazil, Pará, Santo Antônio do Tauá, Reserva Sonho Azul, DZ 45.005).
S, T S. (C.) lizeri album: male (Brazil, Minas Gerais, Carmo do Rio Claro, OM 2.788). U–X S. (C.) lizeri lizeri: U, V male (Peru, Madre de Dios, Tambopata Reserve, DZ 46.176); W, X female (Colombia, Meta, Villavencio, Bosque Bavaria, DZ 45.526).

 Scale bar = 1 cm.

Species of Staphylus (Capilla) in dorsal and ventral views.
A–D S. (C.) tucumanus: A, B male (Argentina, Salta, Pichanal, DZ 45.125); C, D female (Paraguay, Presidente Hayes, Estancia Victoria, Rio Norte Lindo, OM 37.963).
E–H S. (C.) azteca: E, F male (El Salvador, La Libertad, Santa Tecla, MGCL 1112133); G, H female (Mexico, Chiapas, San Jerónimo, DZ 45.085).
I–L S. (C.) tyro: I, J male (Venezuela, Aragua, Ocumare de la Costa, DZ 22.893); K, L female (Venezuela, Aragua, Ocumare de la Costa, DZ 46.608).

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944).
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

M–P Staphylus (C.) nicoleae sp. nov.: M, N male (Colombia, Cundinamarca, El Boqueron, holotype, DZ 45.163); O, P female (Colombia, no specific locality, paratype, NHMUK015052554);
Q, R S. (C.) neivae sp. nov.: male (Brazil, Paraíba, São José da Mata, Campina Grande, holotype, DZ 44.944). 
S, T S. (C.) ricardoi sp. nov.: male (Peru, Tingo Maria, holotype, OM 41.643).
U–X S. (C.) neideae sp. nov.: U, V male (Brazil, Minas Gerais, Brumadinho, holotype, OM 55.562); W, X female (Brazil, Minas Gerais, Brumadinho, paratype, OM 55.476).

Scale bar = 1 cm.

Staphylus (Capilla) nicoleae Lemes sp. nov. from Colombia and Venezuela, 

S. (C.) ricardoi Lemes, sp. nov. from Peru, 

S. (C.) neideae Lemes sp. nov. and S. (C.) neivae sp. nov. from Brazil.  

 José Ricardo Assmann Lemes, Ricardo Russo Siewert, Olaf Hermann Hendrik Mielke, Mirna Martins Casagrande and Andrew David Warren. 2026. Integrative Taxonomic Revision of Capilla Grishin, 2023, Subgenus of Staphylus Godman & Salvin, 1896 (Lepidoptera, Hesperiidae, Pyrginae, Carcharodini), with Descriptions of Four New Species. Arthropod Systematics & Phylogeny. 84: 123-173. DOI: doi.org/10.3897/asp.84.e179328 [24 Feb 2026]

[Mammalogy • 2026] Found Alive After 6,000 years: Modern Records of An ‘Extinct’ Papuan Marsupial, Dactylonax kambuayai (Marsupialia: Petauridae), with A Revision of the Systematics and Zoogeography of the Genus Dactylonax

Dactylonax kambuayai (Aplin, 1999) 

Pygmy Long-fingered Possum

in Flannery, Aplin, Bocos, Koungoulos et Helgen, 2026. 

Records of the Australian Museum. 78; 

DOI: doi.org/10.3853/j.2201-4349.78.2026.3003

 painting by Peter Schouten. 

Abstract

The Pygmy Long-fingered Possum, Dactylonax kambuayai, is the smallest of the striped possums (the petaurid subfamily Dactylopsilinae). It is a ‘Lazarus species’, found living when previously known only from fossils. Recently collected museum specimens, and observations and photographs of living individuals, correspond taxonomically to the only previously reported specimens of D. kambuayai, which are fossil remains from a nearby Holocene location, all occurring in low- to mid-elevation rainforests on the Vogelkop Peninsula of New Guinea. Together with the recent discovery of another Lazarus possum, ‘Petauroides’ ayamaruensis, they represent the only marsupials known as modern animals only from low elevations (below 1000 m) on the Vogelkop Peninsula. Both of these species are diminutive species with enigmatic natural histories. In reviewing the systematics of this species, we demonstrate the distinctness of Dactylonax as a genus-level taxon, and clarify species boundaries in this genus. Dactylonax palpator, the type species of Dactylonax, was previously considered to be a widespread montane taxon, but is recognized here as two separate species: D. palpator is restricted to the Arfak Mountains of the Vogelkop, while D. ernstmayri is widely distributed on the New Guinean Central Cordillera and the Huon Peninsula at elevations above 800 m. Dactylonax palpator replaces, or co-occurs with, D. kambuayai at elevations between around 900 m and 1,400 m on the Vogelkop. We hypothesize that colonization of montane habitats on the Vogelkop by a D. kambuayai-like ancestor gave rise to the larger bodied D. palpator, and dispersal of this lineage to montane habitats in the Central Cordillera gave rise to the highly specialized D. ernstmayri. The fact that the montane taxon of the Vogelkop Dactylonax species couplet was able to migrate eastwards, while the lowland taxon was not, suggests that an unusual elevational zoogeographic filter was in play. We observe that D. kambuayai is thus far recorded from sites where the other small petaurid of New Guinea, Petaurus papuanus, is not recorded or is uncommon.

Keywords: Marsupialia; Petauridae; Dactylonax; kambuayai; palpator; ernstmayri; Lazarus taxon; Vogelkop; New Guinea; systematic review

Photograph of a living female Dactylonax kambuayai, Klalik area, Vogelkop.

Photo: Carlos Bocos.

Dactylonax kambuayai (Aplin, 1999)

Hands of the four extant genera (and three subfamilies) of family Petauridae.
Palmar surfaces of the manus, showing key differences in phalangeal proportions, claws, palmar pads, furring, and patagial development in
(a) Dactylopsila, subfamily Dactylopsilinae (represented by Dactylopsila trivirgata); (b) Dactylonax, subfamily Dactylopsilinae (represented by Dactylonax palpator);
(c) Gymnobelideus, represented by G. leadbeateri (Gymnobelideus McCoy, 1867 is type genus of the new subfamily proposed here: Gymnobelideinae Helgen & Flannery, subfam. nov., Gymnobelideus leadbeateri McCoy, 1867 is type species of Gymnobelideus, the new subfamily and its type genus are both monotypic);
(d) Petaurus, subfamily Petaurinae (represented by P. norfolcensis). Modified from Flannery (1994:61–65). Drawings by Peter Schouten.

Dactylonax kambuayai (Aplin, 1999)—a painting by Peter Schouten.
 This artwork is based on the two modern specimens in the Australian Museum (M.56796 adult female and pouch young M.56797).

Tim F. Flannery; Kenneth P. Aplin; Carlos Bocos; Loukas G. Koungoulos; Kristofer M. Helgen. 2026. Found Alive After 6,000 years: Modern Records of An ‘Extinct’ Papuan Marsupial, Dactylonax kambuayai (Marsupialia: Petauridae), with A Revision of the Systematics and Zoogeography of the Genus Dactylonax. Records of the Australian Museum. 78; 17-34. DOI: doi.org/10.3853/j.2201-4349.78.2026.3003 [06 March 2026]