[Botany • 2026] Ophiorrhiza xinchengensis (Rubiaceae) • A New Species from Guangxi, China

Ophiorrhiza xinchengensis Y.Nong & G.Y.Wei, 

in Wei, J.-Y. Li, Chen, Y. Nong, Hu, Y.-J. Li, Ji, Xu et Y.-K. Nong. 2026. 

忻城螺序草  |  DOI: doi.org/10.3897/phytokeys.278.194094

Abstract

Ophiorrhiza xinchengensis (Rubiaceae), a new species from the middle part of Guangxi, China, is described and illustrated on the basis of morphological evidence. This new species resembles Ophiorrhiza longanensis in its densely pubescent stems, linear stipules, terminal or upper axillary inflorescences, but it can be distinguished by its elliptic or oblanceolate leaf blade, base attenuate (vs. oval, ovate to broadly ovate, base cuneate), its 2–3 mm long stipules (vs. 3.8–6.5 mm long), its 3–5 mm long bracteoles (vs. 6–10 mm long) its 10–12 mm long corolla tubes (vs. 2–3 mm long) and its filaments adnate at the middle of corolla tube, styles 12–15 mm long (vs. adnate to the base of corolla tube and styles ca. 3 mm long). The new species is currently known from a single population with approximately 60–100 mature individuals. Following IUCN Criteria, it is provisionally assessed as Critically Endangered. Photographs, an illustration and a distribution map are also provided.

Key words: Karst, new species, Ophiorrhiza, Spiradiclis, taxonomy

Ophiorrhiza xinchengensis Y.Nong & G.Y.Wei.
A. Habitat; B, C. Flowering plant; D. Stem, showing the linear stipules; E. Inflorescences; F. Fruiting plant; G. Dissected brevistylous flower; H. Dissected longistylous flower; I. Hypanthium, calyx, style and stigma; J. Ovary and calyx; K. Stigma; L. Bracts; M, N. Leaf (abaxially and adaxially); O. Capsule; P. Seeds.

Ophiorrhiza xinchengensis Y.Nong & G.Y.Wei, sp. nov.

 

Chinese name. “xīn chéng luó xù cǎo” (忻城螺序草).

Diagnosis. Ophiorrhiza xinchengensis is resembles Ophiorrhiza longanensis (R.J.Wang) Razafim. & Rydin in its densely pubescent stems, linear stipules, terminal or upper axillary inflorescences, but it can be distinguished by its elliptic or oblanceolate leaf blade, base attenuate (vs. oval, ovate to broadly ovate, base cuneate), its 2–3 mm long stipules (vs. 3.8–6.5 mm long), its 3–5 mm long bracteoles (vs. 6–10 mm long), its 10–12 mm long corolla tubes (vs. 2–3 mm long) and its filaments adnate at the middle of corolla tube, styles 12–15 mm long (vs. adnate to the base of corolla tube, styles ca. 3 mm long).

Etymology. The specific epithet “xinchengensis” refers to the type location of the new species.

 Gui-Yuan Wei, Jin-Yue Li, Rong Chen, You Nong, Qi-Min Hu, Ying-Jing Li, Xing-Yun Ji, Chuan-Gui Xu, Yun-Kai Nong. 2026. Ophiorrhiza xinchengensis (Rubiaceae), A New Species from Guangxi, China. PhytoKeys. 278: 1-14.  DOI: doi.org/10.3897/phytokeys.278.194094 [24-07-2026]

[Botany • 2026] Ophiorrhiza saxatilis (Rubiaceae) • A New Species from Guizhou, China

Ophiorrhiza saxatilis M.T.An & G.L.Li,

in Li, An, Wu, Yu, Qiu, Liu et Tan, 2026. 

 岩生蛇根草  ||  DOI: doi.org/10.3897/phytokeys.278.197480

Abstract

Ophiorrhiza saxatilis M.T.An & G.L.Li, sp. nov., a new species of the genus Ophiorrhiza (Rubiaceae) discovered in the karst regions of Guizhou, China, and provides its taxonomic basis from a morphological perspective. The new species is morphologically similar to Ophiorrhiza lignosa, but differs from the latter by having an inflorescence in that its young branches are uniformly pubescent (vs. bifariously pubescent in O. lignosa); the abaxial leaf surface is pubescent (vs. puberulent only along the midrib and secondary veins in O. lignosa); the bracteoles are 5–6 mm long (vs. 1.5–3 mm long in O. lignosa); the petioles are 2–5.8 mm long (vs. 6–20 mm long in O. lignosa); the stamens are inserted at the base of the corolla tube (vs. inserted near the middle of the corolla tube in O. lignosa); the flowers are distylous (vs. monomorphic with pin flowers in O. lignosa), and the styles of long-styled flowers are 6–8 mm long (vs. 10–11 mm long in O. lignosa). These stable and significant morphological differences, combined with a comprehensive and detailed morphological description of the new species and a comparison of diagnostic characters with similar species, firmly support the recognition of Ophiorrhiza saxatilis as a distinct new species.

Key words: Karst, Morphology, New Taxon, Ophiorrhiza, Rubiaceae

Ophiorrhiza saxatilis.
A, B. Habitat; C. Plant; D. Flowering branch; E. Dissected flowers: long-styled flower (above) and short-styled flower (below); F. Style of long-styled flower; G. Style of short-styled flower; H. Longitudinally dissected short-styled flower; I. Longitudinally dissected long-styled flower; J. Fruit; K. Infructescence; L. Adaxial leaf surface; M. Abaxial leaf surface; N. Branchlets.

Ophiorrhiza saxatilis M.T.An & G.L.Li, sp. nov.

Diagnosis. Ophiorrhiza saxatilis is morphologically similar to O. lignosa Merr., but can be distinguished by several key characteristics (Table 2): uniformly pubescent young branches (vs. bifariously pubescent); white corolla lobes with ridges (vs. purple, without ridges); stamens inserted at the base of the corolla tube (vs. inserted near the middle of the corolla tube); flowers distylous (vs. monomorphic, with pin flowers); long-styled flowers 6–8 mm long (vs. 10–11 mm long); purple midrib on the abaxial leaf surface (vs. green); abaxial leaf surface pubescent (vs. puberulent only on the midrib and secondary veins); bracteoles 5–6 mm long (vs. 1.5–3 mm long); and petioles 2–5.8 mm long (vs. 6–20 mm long).

Etymology. The specific epithet is derived from the habitat of the new species, which occurs in a typical karst limestone environment. It mainly grows in harsh habitats such as rock crevices and stony slopes.

Local name. Simplified Chinese: 岩生蛇根草;
Chinese Pinyin: yán shēng shé gēn cǎo.

 Guang-Lin Li, Ming-Tai An, Xu Wu, Jiang-Hong Yu, Jian Qiu, Shao-Fei Liu and Cheng-jiang Tan. 2026. Ophiorrhiza saxatilis (Rubiaceae), A New Species from Guizhou, China. PhytoKeys. 278: 27-37.  DOI: doi.org/10.3897/phytokeys.278.197480 [24-07-2026]

[Botany • 2026] Petrocosmea anoleuca (Gesneriaceae) • A New Species from limestone cliffs in Yunnan, China

 

Petrocosmea anoleuca   M.Q.Han & C.Liu, 

in Liu, Cai, Zhong, Zhao, Wang et Han, 2026.

沧源石蝴蝶  ||  DOI: doi.org/10.1007/s12225-026-10406-z

 

Summary

Petrocosmea anoleuca M.Q.Han & C.Liu (Gesneriaceae), a new species from Yunnan, China, is described and colour images and a provisional IUCN conservation assessment are provided. The new species is similar to P. cavaleriei H.Lév., but differs in having subpeltate leaf blades, a white corolla tube base lacking spots, a darker purple throat, purple abaxial lip lobes and nearly straight, sparsely puberulent filaments.

Key Words: Flora of Yunnan, Karst habitats, taxonomy

Petrocosmea anoleuca, images and SEM images.
A – B habit, in natural habit; C – D flower, front view; E flower, side view; F flower bud; G corolla, longitudinally cut; H corolla, transversely cut; J peltate leaf blade, abaxial surface; K – L SEM; K rhizomatous stem; L ovary; M ovary, transversely cut; N pollen.

photos: a – b Cheng Liu and c – k Mengqi Han.

Petrocosmea anoleuca M.Q.Han & C.Liu sp. nov. 

Recognition. Similar to Petrocosmea cavaleriei, but differing in having subpeltate leaf blades (vs petiole attached at the base in P. cavaleriei), the base of the corolla tube white and lacking spots (vs with three antepetalous spots above the base), the throat inside abaxially darker purple (vs white), the abaxial lip lobes purple (vs pale purple, with two yellow spots at the sinuses between the lateral and median lobes) and filaments almost straight and very sparsely puberulent and with scattered capitate-glandular hairs (vs geniculate near the middle, forming an angle of about 160° and densely covered with capitate-glandular hairs).

Etymology. The term anoleuca derives from Greek roots, ano- (ἄνω, "above") and -leuca (λευκός, "white"), literally meaning "white above". The epithet of the new species refers to the white, upper section of the corolla when in bloom.

vernacular name. Chinese name: cāng yuán shí hú dié (沧源石蝴蝶).

Cheng Liu, Jie Cai, Liang-Liang Zhong, Jin-Chao Zhao, Qiang Wang and Meng-Qi Han. 2026. Petrocosmea anoleuca (Gesneriaceae), A New Species from limestone cliffs in Yunnan, China.  Kew Bulletin. DOI: doi.org/10.1007/s12225-026-10406-z  [21 July 2026]

[Herpetology • 2026] Gekko qiannanensis • A New Karst-adapted Gecko (Squamata: Gekkonidae) from southern Guizhou, China

Gekko (Japonigekko) qiannanensis Huang, Ma, Jiang & Wu,   

in Huang, Ma, Xiao, Zhang, Pu, Ran, Zhu, Jiang et Wu, 2026. 

黔南壁虎  ||  DOI: doi.org/10.3897/zse.102.189932

Abstract

Karst landscapes, which are characterized by their complex terrain, foster exceptionally high species diversity. During recent field surveys in Qiannan Buyi and Miao Autonomous Prefecture, Guizhou Province, China, a new karst-adapted gecko species was discovered. Using an integrative approach combining molecular and morphological analyses, it is described here as Gekko qiannanensis sp. nov. Phylogenetically, it clusters within a well-supported clade of the subgenus Japonigekko that includes the karst-adapted G. fengshanensis, G. liboensis, G. kwangsiensis, G. paucituberculatus, and the dry-hot valley species G. tesselatus. Morphologically, it can be distinguished from all known species in Japonigekko by a combination of the following traits: moderate body size, with snout–vent length 65.4–82.9 mm in adult males and 77.7–91.7 mm in adult females; two enlarged postmentals; nares in contact with the rostral, internasal absent; subdigital lamellae: fourth fingers 14–16, first toes 9–11, and fourth toes 14–17; moderately developed toe webbing, originating proximal to subdigital lamellae; interorbitals 28–38; midbody scale rows 134–155; ventral scale rows 42–49; precloacal pores 10–11 in males; a single postcloacal tubercle on each side; dorsum earth gray, with six dirty-white bands between the nape and sacrum. Notably, the new species can be distinguished from its karst-adapted congeners by its moderately developed toe webbing. The discovery of G. qiannanensis brings the number of recognized species in Japonigekko to 42 globally, of which 28 are now recorded from China. This finding highlights the role of karst habitat fragmentation in driving allopatric speciation and underscores the rich but still incompletely documented herpetofaunal diversity of South China’s karst regions.

Key Words: herpetofauna, integrative taxonomy, new species, phylogeny, toe webbing

Holotype (JSU-HJ2025024, adult male) of Gekko qiannanensis sp. nov. after euthanasia.
 A. Dorsal view of body. B. Ventral view of body. C. Dorsal view of head. D. Ventral view of head. E. Right lateral view of the head. F. Ventral view of right hand. G. Ventral view of left hand. H. Ventral view of right foot. I. Ventral view of left foot. J. Ventral view of precloacal region. K. Right lateral view of basal tail.

Photos by Jie Huang and Yang Xiao. Scale bars: 10 mm.

Egg hatching of Gekko qiannanensis sp. nov.  
A. Discovery of gecko eggs at the corner of the wall, with 14 eggs already hatched and two remaining unhatched. B. Two hatching gecko eggs with attached wall plaster. C. One gecko egg in the process of hatching, side view. D. One gecko egg in the process of hatching, top view. E. The gecko juvenile 13 days after hatching.

Photos by Jie Huang. Scale bar: 10 mm.

Gekko qiannanensis Huang, Ma, Jiang & Wu, sp. nov.

 

Diagnosis. Gekko qiannanensis sp. nov. is assigned to the subgenus Japonigekko according to phylogenetic analysis. It can be distinguished from other congeners in Japonigekko by the following characters: moderate body size: SVL 65.4–82.9 mm in adult males, 77.7–91.7 mm in adult females; two enlarged postmentals; nares in contact with the rostral, internasal absent; subdigital lamellae: fourth fingers 14–16, first toes 9–11, fourth toes 14–17; moderately developed toe webbing, originating proximal to subdigital lamellae; interorbitals 28–38; Midbody scale rows 134–155; ventral scale rows 42–49; precloacal pores 10–11 in males; a single postcloacal tubercle on each side; dorsum earth gray, with six dirty-white bands between nape and sacrum.

Etymology and suggested Chinese and English name. The specific epithet, qiannanensis, is derived from the Chinese pinyin of its type locality in Qiannan Buyi and Miao Autonomous Prefecture. The Chinese name for the new species is suggested as “黔南壁虎” (pinyin: qián nán bì hǔ), and the English is suggested as “Qiannan Gecko.”

 Jie Huang, Shun Ma, Yang Xiao, Fangpeng Zhang, Zhongzhu Pu, Jingcheng Ran, Jingyi Zhu, Wansheng Jiang and Jianpu Wu. 2026. Gekko (Japonigekko) qiannanensis, A New Karst-adapted Gecko (Squamata, Gekkonidae) from southern Guizhou, China. Zoosystematics and Evolution. 102(4): 973-987. DOI: doi.org/10.3897/zse.102.189932 [20-07-2026]

[Mollusca • 2026] Cyllena hermes • From the Dark to the Light: A New Genus and Species of stygobiont hydrobiid (Caenogastropoda: Truncatelloidea: Hydrobiidae) from southern Greece

Cyllena hermes  Radea & Protopapas,

in Radea, Protopapas, Parmakelis et Koskeridou, 2026. 

DOI: doi.org/10.3897/subtbiol.57.189090

Abstract

A new stygobiont hydrobiid genus and species is described from the Stymphalia karst basin (Peloponnese, Greece). Cyllena hermes gen. et sp. nov. is distinguished from other valvatiform or valvatiform–planorboid stygobiont hydrobiids by a unique combination of shell and anatomical characters, supporting the establishment of a new monotypic genus. It is currently known from a single karstic spring connected to the subterranean hydrological network of the Stymphalia polje, indicating a highly localized distribution. Following a precautionary approach, and in accordance with IUCN criteria, the species is here considered to qualify as Vulnerable (VU D2). The discovery of the new genus contributes to the documentation of subterranean aquatic gastropod diversity in Greece and emphasizes the need for further targeted exploration of karst ecosystems in the region.

Keywords: Endemism, Hydrobiidae, IUCN, Peloponnese, karst aquifer

Cyllena hermes gen. et sp. nov.
Shell views of collected specimens. A, G, J, M, P dorsal views B, H, K, N, Q ventral views C–E, I, L, O, R lateral views F operculum A–E holotype.

 Scale bar: 1 mm.

Cyllena hermes gen. et sp. nov.
Views from a single immature specimen. A dorsal view B ventral view C, D lateral view E–G scanning electron micrographs of shell and protoconch respectively.

Scale bars: 1 mm (A–E); 100 μm (F); 50 μm (G).

Systematic description

Class Gastropoda Cuvier, 1795

Subclass Caenogastropoda Cox in Moore, 1960

Order Littorinimorpha Golikov & Starobogatov, 1975

Superfamily Truncatelloidea J.E. Gray, 1840

Family Hydrobiidae W. Stimpson, 1865

Cyllena Radea & Protopapas, 2026, gen. nov. 

Type species. Cyllena hermes sp. nov. by original designation.

Diagnosis. Shell minute, valvatiform to planorboid; umbilicus wide, with earlier whorls visible; soft body unpigmented; eyes absent; bursa copulatrix pyriform; a single seminal receptacle [proximal, rs2 (Radoman 1973)] large and irregular in shape; penis large, dorso-ventrally flattened, distally blunt with papilla and proximally folded, bearing a single, elongate, blunt, medial lobe.

Etymology. The generic name derives from the Greek mythology: Cyllena (Κυλλήνη in Greek) was a nymph of Mt Kyllini, Korinthia, Peloponnese, and the nurturer of the god Hermes. Gender feminine.

Cyllena hermes Radea & Protopapas, 2026, sp. nov.

Etymology. The specific name, used in apposition, is derived from Greek mythology. Hermes (Ἑρμῆς in Greek) was one of the twelve Olympian gods, the messenger of the other Olympian deities, and was born in a cave on Mt Kyllini.

Live specimens of Cyllena hermes gen. et sp. nov.
A–F live mature specimen, shown from the moment of emergence from the shell to active locomotion while foraging 
G–I another mature specimen; White arrows indicate the direction of movement J immature specimen floating at the water surface while feeding
K caddisfly case (Insecta, Trichoptera) constructed from gastropod shells, including shells of Cyllena hermes gen. et sp. nov. Scale bars: 1 mm.

Canella Radea, Dimitrios Protopapas, Aristeidis Parmakelis and Efterpi Koskeridou. 2026. From the Dark to the Light: A New Genus and Species of stygobiont hydrobiid (Caenogastropoda, Truncatelloidea) from southern Greece. Subterranean Biology. 57: 1-21. DOI: doi.org/10.3897/subtbiol.57.189090  [18 Jun 2026]

 

 

[Mollusca • 2022] Burmochlamys fasciola, B. albida, B. versicolor, ... • Uncovering local Endemism from southeastern Myanmar: Description of the New Karst-associated Terrestrial Snail Genus Burmochlamys (Eupulmonata: Helicarionidae)

 

Burmochlamys Pholyotha & Panha,
A Burmochlamys fasciola sp. nov. paratype showing left and right lateral views;  
D B. albida sp. nov. paratype; E B. moulmeinica sp. nov. paratype;
F B. versicolor sp. nov. paratype; G B. whitteni sp. nov. paratype 

in Pholyotha, Sutcharit, Lin et Panha, 2022.   

 DOI: doi.org/10.3897/zookeys.1110.82461 

Abstract

Salween River basin’s karst ecosystems in southeastern Myanmar remain largely unexplored and are likely to harbour a high terrestrial snail diversity that are often associated with high levels of snail endemism. Here, an outstanding group of new karst-associated terrestrial snails, Burmochlamys gen. nov., are discovered. A study of the comparative morphological and anatomical data reveals that the reproductive tract and radula of this new genus are closely related to the helicarionid genus Sophina Benson, 1859 but shell morphology (shape, size, and sculpture) and mantle extensions are distinct from the latter genus. Burmochlamys gen. nov. now consists of four known nominal species, B. cassidula comb. nov., B. cauisa comb. nov., B. perpaula comb. nov., and B. poongee comb. nov., and five new species; B. albida sp. nov., B. fasciola sp. nov., B. moulmeinica sp. nov., B. versicolor sp. nov., and B. whitteni sp. nov. The highlight is that the members of the new genus show site-specific endemism, being restricted to karstic habitat islands of the Salween River basin. In addition, the discovery supports that the unique and complex structure of Salween River basin’s karst ecosystems are habitats in which the terrestrial malacofauna have speciated and become endemic.

Keywords: Diversity, endemic, Indochina, land snail, limestone, Salween River basin, taxonomy

Family Helicarionidae Bourguignat, 1877

Subfamily Durgellinae Godwin-Austen, 1888

Living snails of Burmochlamys species
A Burmochlamys fasciola sp. nov. paratype CUMZ 14214 showing left and right lateral views B B. cassidula specimen CUMZ 14209
C B. poongee specimen CUMZ 14210 D B. albida sp. nov. paratype CUMZ 14212
E B. moulmeinica sp. nov. paratype CUMZ 14217 F B. versicolor sp. nov. paratype CUMZ 14219 G B. whitteni sp. nov. paratype CUMZ 14222.

Genus Burmochlamys Pholyotha & Panha, gen. nov.

 

Type species: Burmochlamys fasciola sp. nov., by original designation.

Etymology: The name combines Burmo in reference to Burma, the historical name of Myanmar, and the Greek word chlamys meaning mantle or cloak in reference to land snail with well-developed mantle extensions. Therefore, the generic name means the Burmese land snail with the well-developed mantle extensions. The gender of the new generic name is feminine.

Diagnosis: Shell subglobose to globose, small size, little high spire, and sculptured with spiral furrows and undulating radial lines. Snail with five well-developed mantle extensions; caudal horn raised. Genitalia with penial retractor muscle attached at tip of epiphallic caecum; gametolytic organ with rather short to moderate cylindrical duct and bulbous sac; well-developed dart apparatus; flagellum absent. Radula with large monocuspid central tooth and attached by two smaller teeth; laterals and marginals undifferentiated, large monocuspid, and at base of each tooth on outer side attached by a smaller tooth.

B. cassidula comb. nov., B. cauisa comb. nov., 

B. perpaula comb. nov., and B. poongee comb. nov., 

and five new species; B. albida sp. nov., B. fasciola sp. nov., 

B. moulmeinica sp. nov., B. versicolor sp. nov., and B. whitteni sp. nov.

A adult of Burmochlamys moulmeinica sp. nov. while climbing slowly on the karst wall after raining at Lun Nya Pagoda B mating pairs of B. versicolor sp. nov. on the karst wall at Bayin Nyi Cave
C B. versicolor sp. nov. eaten by the carnivorous snail Carinartemis sp. on the karst wall at Bayin Nyi Cave D B. cassidula eaten by the carnivorous snail Haploptychius sp. on the karst wall at Kaw Gon Cave. 

 Arthit Pholyotha, Chirasak Sutcharit, Aung Lin and Somsak Panha. 2022. Uncovering local Endemism from southeastern Myanmar: Description of the New Karst-associated Terrestrial Snail Genus Burmochlamys (Eupulmonata, Helicarionidae).  ZooKeys. 1110: 1-37. DOI: doi.org/10.3897/zookeys.1110.82461 

[Botany • 2026] Ceropegia gengmaensis (Apocynaceae: Ceropegieae) • A New Species from Yunnan, China

 

Ceropegia gengmaensis  P.R.Luo, A.Liu & H.Sun,

in Luo, Liu, Tong, Zhang, Wang, Deng, Goyder et Sun, 2026. 

耿马吊灯花  ||  DOI: doi.org/10.3897/phytokeys.276.197333

Abstract

Ceropegia gengmaensis P.R.Luo, A.Liu & H.Sun, a new species from Gengma County, Yunnan, Southwest China, is described. Both morphological characteristics and chloroplast phylogenetic analysis strongly support its placement within C. sect. Chionopegia H.Huber. Molecular data further reveal that C. gengmaensis is closely related to C. salicifolia, C. mairei, and C. dolichophylla, yet it can be distinguished from them readily by its stem indumentum, leaf shape, and floral morphology, particularly the shape of the corolla tube, and features of the lobes. A detailed line drawing of this new species is also provided.

Key words: Ceropegieae, Chloroplast, Karst Landform, Morphology, Southwest China

Ceropegia gengmaensis. 
 A, B. Plant showing the rootstock; C, D. Upper and lower sides of leaf; E. Inflorescence with bases of adjoining leaves; F. Corolla tube dissected longitudinally; G. Side view of gynostegium (with sepals and pedicel shown); H. Top view of gynostegium with sepals shown; I. Ovary; J Pollinarium.

 Scale bars: 15 cm (A, B); 5 cm (C, D); 4 cm (E, F); 1 cm (G); 0.8 cm (H); 0.3 cm (I); 0.5 mm (J).

Morphological features of Ceropegia gengmaensis based on living plants collected from the type locality.
 A. Plant showing the rootstock; B, C. Leaf: B, abaxial surface and C, adaxial surface; D. Stem; E, F. Corolla: E, corolla from the outside with its base cut off and F, corolla with the tube dissected longitudinally; G. Small peduncle, part of the pedicel and petiole; H. White hairs on the inside of the base of the tube; I. Side view of gynostegium; J. Gynostegium from above; K. Pollinarium.

Scale bars: 15 cm (A); 5 cm (B, C); 4 cm (D, E, F); 1 cm (G, H, I); 0.8 cm (J); 0.5 mm (K).

Ceropegia gengmaensis P.R.Luo, A.Liu & H.Sun, sp. nov.

 

Diagnosis. Ceropegia gengmaensis shows affinities with C. salicifolia and C. dolichophylla, both of C. sect. Chionopegia . It differs from C. salicifolia by its erect to sprawling habit (vs. twining), elliptic leaves 2.0–5.5 cm (vs. lanceolate, 6–15 cm), densely pubescent stems bearing usually solitary flowers (vs. glabrous stems with sessile cymose many-flowered inflorescences), and narrower corolla lobes with only slight expansion above the throat (vs. lobes longer and wider, upper portion markedly expanded). From C. mairei it differs by the much less pronounced basal inflation of the tube (vs. a strongly inflated tube) and ... 

Etymology. Ceropegia gengmaensis is named after its type locality, Gengma County, Southwest Yunnan.

Vernacular name. the Chinese name is given as “耿马吊灯花” (gěng mă diào dēng huā) named after the type locality, Gengma County, Southwest Yunnan.

Peng-Rui Luo, Ang Liu, Rui Tong, Xin-Jian Zhang, Zi-Yi Wang, Tao Deng, David J. Goyder and Hang Sun. 2026. Ceropegia gengmaensis (Apocynaceae), A New Species from Yunnan, China. PhytoKeys. 276: 217-228. DOI: doi.org/10.3897/phytokeys.276.197333 [15 Jun 2026]

[Botany • 2026] Oreocharis fulvovillosa (Gesneriaceae) • A New Species from western Yunnan, China

Oreocharis fulvovillosa  M.Q.Han, H.Jiang & Y.Wang,

in Wang, Jiang, Xu et Han, 2026. 

德宏直瓣苣苔  ||  DOI: doi.org/10.3897/phytokeys.276.182136

Abstract

Oreocharis fulvovillosa, a new species of Gesneriaceae, is described and illustrated. This new species has only been discovered on the rock walls of an isolated karst hill at its type locality in Mang City, western Yunnan, China.

Key words: Flora of Yunnan, karst, yellow flowers

 

Oreocharis fulvovillosa sp. nov.
A. Habitat; B. Plant habit in natural habitat; C. Adaxial (left) and abaxial (right) leaf surfaces; D. Petiole with long fulvous indumentum; E. Rhizomatous stem; F. Flower in adaxial view; G. Flower in abaxial view; H. Flower in side view; I. Flower longitudinally cut, male stage with immature pistil; J. Flower transversely cut, male stage with immature pistil; K. Immature pistil and stamens with dehisced anthers cohering in pairs; L. Flower in front view; M. Capsules at different developmental stages: two immature capsules (left) and two dehisced capsules from the previous year (right); N. Pistils at different developmental stages: left, at flower opening with immature stigma; middle and right, at stigma maturity; O. Transverse sections of ovaries, all arranged with the dorsal calyx lobe positioned upwards: upper image, ovary at stigma maturity; lower images, ovary with immature stigma, showing transverse sections of the upper part of the ovary (left) and the lower part of the ovary (right).

Oreocharis fulvovillosa M.Q.Han, H.Jiang & Y.Wang, sp. nov.

 

Diagnosis. The new species is most similar to Oreocharis bullata, sharing yellow corollas and bullate leaf blades, but differs in having crenulate leaf margins (vs. double dentate), a slender corolla tube ca. 1 cm long (vs. tubular, ca. 2.7 cm long), and shorter capsules 1–1.9 cm long (vs. 3.0–4.2 cm long). It is also similar to Oreocharis fulva in corolla shape and colour but differs in having bullate leaf blades (vs. not bullate) and crenulate margins (vs. serrate to crenate).

Etymology. The epithet refers to the fulvous (yellowish) colour of the long, soft hairs on the petioles.

Vernacular name. Chinese Mandarin: dé hóng zhí bàn jù tái, (德宏直瓣苣苔).

Yang Wang, Hong Jiang, Bo Xu and Meng-Qi Han. 2026. Oreocharis fulvovillosa (Gesneriaceae), A New Species from western Yunnan, China. PhytoKeys. 276: 17-23.  DOI: doi.org/10.3897/phytokeys.276.182136 [08-06-2026]

 

[Botany • 2026] Syzygium khammouanense (Myrtaceae) • A New Species from central Laos

 

Syzygium khammouanense V.S.Dang, Tagane & Soulad.,

in Dang, Souladeth, Pham, Kongxaisavath, Phengmala, Sengthong, Souvannakhoummane, Vongthavone, Yamamoto, Tanaka, Takahashi et Tagane, 2026.

ຫວ້າຄຳມ່ວນ  ||  DOI: doi.org/10.1007/s12225-025-10351-3  

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Summary

A new species, Syzygium khammouanense V.S.Dang, Tagane & Soulad. (Myrtaceae), discovered from Khounkham District, Khammouane Province, central Laos, is described and illustrated. The new species is distinct from other Syzygium species by having 26 – 30 pairs of secondary veins and a strongly quadrangular hypanthium, 1 cm long. A description, provisional conservation assessment, photographs and vernacular name of the new species are provided.

Key Words: Indochina, Khounkham, limestone flora, plant diversity, taxonomy

Syzygium khammouanense V.S.Dang, Tagane & Soulad.
A flowering branch; B leaf, abaxial surface; C flower bud; D inflorescence; E flower, cutaway, showing stamens and style; F young fruits; G hypanthium, with style (left), and petals, adaxial surface (centre), abaxial surface (right). photos by: Shuichiro Tagane.

Syzygium khammouanense V.S.Dang, Tagane & Soulad. sp. nov. 

Etymology. The specific epithet “khammouanense” refers to the name of the province containing the type locality, in Laos.

Vernacular name. ຫວ້າຄຳມ່ວນ (Wa Khammouane, suggested here). “Wa” is the common Lao name for the genus Syzygium, and “khammouanense” represents the province’s name where the type material was collected.

 

Van-Son Dang, Phetlasy Souladeth, Quoc-Trong Pham, Deuanta Kongxaisavath, Kajonesuk Phengmala, Anousone Sengthong, Keooudone Souvannakhoummane, Thyraphon Vongthavone, Takenori Yamamoto, Nobuyuki Tanaka, Kotaro T. Takahashi and Shuichiro Tagane. 2026. A New Species of Syzygium P.Browne ex Gaertn. (Myrtaceae), S. khammouanense, from central Laos.  Kew Bulletin. DOI: doi.org/10.1007/s12225-025-10351-3 [21 May 2026] 

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[Ichthyology • 2026] Oreonectes weii • A New endemic Species (Cypriniformes: Nemacheilidae) within the Yangtze River Basin and its Monsoon-driven Evolutionary Diversification

Oreonectes weii

Luo, Ling, Cao, Zhou & Huang, 2026

 

魏氏岭鳅  ||  DOI: doi.org/10.3897/zse.102.186155 

Abstract

A new species Oreonectes weii sp. nov., is described that was collected from Shanggao County, Jiangxi Province, China, located at the upper reaches of the Yangtze River Delta. This naming honors the globally distinguished conservation biologist Fu-Wen Wei for his exceptional and pioneering contributions to biodiversity conservation and research. Morphologically, the new species can be distinguished from its congeners by a combination of meristic and morphological characters, including fin-ray counts, body coloration, eye normal, number of lateral-line pores, and gill-raker counts. Genetically, it forms a distinct lineage in the mitochondrial Cyt b-based phylogeny and exhibits a genetic distance of 6.3% from its sister species, O. polystigmus. We further estimated divergence time within the genus Oreonectes, which indicates an origin in the Late Oligocene (~27.06 Ma) and a most recent common ancestor at ~13.91 Ma. Our biogeographic analyses suggest that the Guijiang-Hejiang River Basin likely served as a source area for the genus’ dispersal into adjacent basins, and that the new species probably originated from a dispersal event of its ancestral population from the Pearl River Basin to the Yangtze River Basin during the Late Miocene (~6.78 Ma). Lineage-divergence dynamics indicate that cladogenesis began around 28 Ma, accelerated markedly at ~18 Ma, peaked at ~6 Ma, and subsequently showed a gradual decline. The current diversity pattern of Oreonectes may have been shaped primarily by dispersal mediated by enhanced precipitation under the East Asian monsoon climate, with subsequent erosion-induced geographical isolation likely promoting speciation and diversification within the genus.

Key Words: Biogeography, cryptic species, morphology, taxonomy, phylogeny, Yangtze River

Ecological photographs and habitats.
A, B. Oreonectes weii sp. nov.; C. O. guidongensis; D. O. polystigmus;
E. Outside the cave; F. Inside the cave.

Oreonectes weii sp. nov.

Diagnosis. Oreonectes weii sp. nov. can be distinguished from all other congeners by the following combination of characters: (1) coloration pattern present and irregular black spots scattered on the lateral body surface and caudal fin; (2) eyes normal; (3) well-developed posterior chamber of air-bladder; (4) cephalic lateral-line system with 4 + 10 infraorbital canal pores and six preoperculo-mandibular canal pores; (5) 9–10 branched pectoral-fin rays, length 16.0–24.3% SL; (6) tip of pelvic fin not reaching the anus; (7) caudal fin truncated, with 14 branched caudal-fin rays; (8) 10 inner gill rakers on the first gill arch; (9) tip of maxillary barbel not reaching the posterior margin of the operculum.

Etymology. The specific epithet “weii” is a genitive noun derived from the surname of Academician Fu-Wen Wei (魏辅文), an eminent conservation biologist. The name is given in honor of his outstanding contributions to the study and conservation of biodiversity in China. We suggest the English common name for this species is the “Wei’s Mountain Loach”, and its Chinese name is “Wèi Shì Lǐng Qiū” (魏氏岭鳅).

 Tao Luo, Wen-Qing Ling, Hai-Lin Cao, Jiang Zhou and Guang-Ping Huang. 2026. A New endemic Species Oreonectes weii sp. nov. (Cypriniformes, Nemacheilidae) within the Yangtze River Basin and its Monsoon-driven Evolutionary Diversification. Zoosystematics and Evolution. 102(3): 751-765. DOI: doi.org/10.3897/zse.102.186155 [14 May 2026]

[Diplopoda • 2026] Alafanahydesmus lavasoa • Description before Extinction: A New Genus and Ninth Species of indigenous Polydesmida (Dalodesmidae) from Madagascar

Alafanahydesmus lavasoa  

Golles, Yoo & Wesener, 2026

DOI: doi.org/10.11646/zootaxa.5807.1.4 

Abstract

Madagascar’s microendemism is not restricted to large rainforests, but also still exists in small, isolated fragments of natural vegetation. One such fragment is the Lavasoa-Ambatotsirongorongo protected area in southeastern Madagascar, where less than 92 ha of humid forest continue to face deforestation, but to which several species are endemic and critically endangered, including three species of millipedes. Here, we describe Alafanahydesmus lavasoa gen. nov., sp. nov. (Polydesmida: Dalodesmidae), a locally endemic Polydesmida only found in the Lavasoa-Ambatotsirongorongo area. With our here described addition, Polydesmida remain rare on Madagascar, with only four genera and nine known species, all belonging to the family Dalodesmidae. Alafanahydesmus lavasoa gen. nov., sp. nov. shares its small size (10–25 mm) and light/brown colouration with Dalodesmus Cook, 1896, Eutubercularium Brölemann, 1916, and Phymatodesmus de Saussure & Zehntner, 1897, but differs conspicuously in somatic (paraterga projecting weakly before tergite 6, elongated shape of metatergal tubercles, paraprocts (pa) with dorsal setose knobs) and sexual characters (stoutly fused femorites and a unique five-branched gonopod with a retrorse branch). Here, we describe Alafanahydesmus lavasoa gen. nov., sp. nov. integratively, providing the first molecular data for Malagasy Dalodesmidae—via partial cytochrome c oxidase I (COI) sequences (GenBank PX922879–PX922581) and pioneer the first microcomputed tomography (μCT) images for a member of the family, alongside with multilayer photographs and scanning electron microscopy images.

Myriapoda, biodiversity, cytochrome c oxidase I gene, Lavasoa, µCT, microendemism

Alafanahydesmus lavasoa gen. nov., sp. nov., Living specimens, at type locality Lavasoa. Photos taken by Moritz Einhaus in 2025.
A. Adult female walking, dorsolateral view. B. Anterior half of body and head, lateral view. C. Midbody segments and legs, close anterodorsal view. D. Rolled-up female showing egg-care behaviour, lateral view. Not to scale.

Alafanahydesmus lavasoa gen. nov., sp. nov., holotype male (ZFMK MYR14588), focus-stacked images.
A. Head and first segments, dorsal view. B. Telson, lateral view. C. Telson, ventral view. D. Mid-body, dorsal view. E. Posterior half of body, dorsal view. F. Gonopods, ventral view. G. Gonopods, ventrolateral view. H. Gonopods, lateral view. Not to scale.

Abbreviations: Co = collum; db = dichotomous branch; ep = epiproct; h = head; hy = hypoproct; o = ozopore; pa = paraproct; pb = paramedian branch; rb = retrorse branch; sl = subsecuriform solenomere; T = tergite.

 Genus Alafanahydesmus new genus 

Type species: Alafanahydesmus lavasoa sp. nov.

Etymology: The generic name is derived from the Malagasy words ala (forest) and fanahy (soul, spirit, inner essence), combined with the suffix -desmus, commonly used for Polydesmida. The name is treated as masculine.

Distribution: Currently only known from south-eastern Madagascar, the isolated rainforest of the Lavasoa-Ambatotsirongorongo area (Fig. 1).

Alafanahydesmus lavasoa gen. nov., sp. nov., Micro-CT scans, paratype male, (ZFMK MYR14544).
A. Lateral view. B. Anterior body half, showing detailed tergite pattern, dorsal view. C. Segments T9–T12 with stigma openings, ozopores, and  lateral  tergite  projections,  lateral  view. D.  Isolated  second  thoracic  segment  bearing  reduced  second  leg  pair. E.  Close-up  of  sternite  with  posterior  sternite  processes,  ventral  view. F. Telson  and  associated  structures,  ventral  view.  Not  to  scale.

Abbreviations: Co = collum; cx = coxa; ep = epiproct; fe = femur; h = head; hy = hypoproct; o = ozopore; pa = paraproct; pf = prefemur; po = postfemur; sti = stigma opening; s = setae; stp = sterite process; str = sternite; T = tergite; ta = tarsus; ti = tibia.

TIZIANO GOLLES, JEHOON YOO and THOMAS WESENER. 2026. Description before Extinction: A New Genus and Ninth Species of indigenous Polydesmida from Madagascar (Polydesmida: Dalodesmidae).  Zootaxa. 5807(1); 109-124. DOI: doi.org/10.11646/zootaxa.5807.1.4  [2026-05-08]

[Botany • 2026] Impatiens arakanensis & I. cirrosa (Balsaminaceae) • Two New Species from the Arakan Mountain Range of Myanmar [Contributions to the Flora of Myanmar X]

Impatiens cirrosa Nob.Tanaka & T.Sugaw.,

 

in Tanaka, Aung et Sugawara, 2026.  

 DOI: doi.org/10.51033/jjapbot.ID0361 

Photographed by T.Sugawara.

 

Abstract

Two new species, Impatiens arakanensis Nob.Tanaka & T.Sugaw. (Balsaminaceae) and I. cirrosa Nob.Tanaka & T.Sugaw., from the Arakan Mountain Range, Magway Region, southwestern Myanmar, are described here. Impatiens arakanensis is similar to I. decipiens Hook.f. in floral morphology and coloration, but differs in having axillary inflorescence (vs. terminal), smooth dorsal petals (vs. with horn-like appendage), and pubescent lower sepal (vs. glabrous). Impatiens cirrosa is similar to I. tripetala Roxb. in floral morphology, but differs in having purple lower sepals (vs. orange to pale orange) with filamentous cirrus at apex, connate lateral united petals (vs. free) and white ovary with red spots (vs. green). Both of the two new species likely belong to Impatiens subgenus Impatiens sect. Uniflorae.

Keywords: Arakan, balsam, Burma, new taxon, sect. Uniflorae, taxonomy

Impatiens arakanensis.
A. Habit. B. A diagonal front view of flower. C. A front view of flower. D. A side view of flower. E. Dissected flower, showing lower sepal, dorsal petal, and lateral united petals. Scale bars: 10 mm.

Photographed by T.Sugawara.

Impatiens arakanensis Nob.Tanaka & T.Sugaw., sp. nov.   

 Impatiens arakanensis is similar to I. decipiens in floral morphology and purplish coloration, but distinguished from it by having an axillary inflorescence (vs. terminal inflorescence), smooth dorsal petals (vs. with horn-like appendage), and pubescent lower sepal (vs. glabrous).

Etymology: The specific epithet “arakanensis” refers to the name of the Arakan Mountain Range in which its type locality is located. 

Impatiens cirrosa.
A. Habit. Arrows showing a filamentous cirrus. B, C. A diagonal front view of flower, showing a filamentous cirrus. D. A front view of flower. E. Dissected flower: a. Lower sepal. b. A young fruit with pedicel. c, d. Lateral united petals. e. Dorsal petal. Scale bars: 10 mm.

Photographed by T.Sugawara.

Impatiens cirrosa Nob.Tanaka & T.Sugaw., sp. nov.  

 Impatiens cirrosa is similar to I. tripetala in floral morphology, but distinguished from it by having purple lower sepals (vs. orange to pale orange) with filamentous cirrus at apex, connate lateral united petals (vs. free) and white ovary with red spots (vs. green).

Etymology: The specific epithet “cirrosa” refers to the most distinctive character of Impatiens cirrosa, which is a very distinct species with long filamentous cirrus at the apex of the lower sepal and lower margin of lamina (Figs. 3B, E, 5).

Nobuyuki Tanaka, Mu Mu Aung and Takashi Sugawara. 2026. Contributions to the Flora of Myanmar X: Two New Species of Impatiens (Balsaminaceae) from the Arakan Mountain Range of Myanmar. J. Jap. Bot. 101(2); 88-96. DOI: doi.org/10.51033/jjapbot.ID0361 [April 20, 2026]