[Botany • 2026] Primula himalayana (Primulaceae) • A New Species from eastern Himalaya, India

 

Primula himalayana B. Hajong, Harsh Singh & P. Bharali, 

in Hajong, Singh et Bharali, 2026. 

DOI: doi.org/10.1002/njb.05052

 

Abstract

Primula himalayana sp. nov., a new species of Primulaceae, is described and illustrated from Tawang, Lutrem, 4238 m a.s.l., Arunachal Pradesh, India. It grows under Juniperus and Berberis scrub forest in an alpine meadow. The new species belong in Primula section Cordifoliae and is resembling P. gambeliana, but differs from the latter by the lamina puberulent with a white protuberance on the tips of the teeth, petiole 2‒3 times longer than lamina and white puberulent, scape long with many flowers, flowers yellow with orange-yellow center and along corolla tube, corolla lobes elliptic, anther basifixed, and ovary cylindrical without any teeth.

Keywords: Arunachal Pradesh, new taxon, Primrose, Primula sect Coridfoliae, Tawang, taxonomy

Primula himalayana B. Hajong, Harsh Singh & P. Bharali sp. nov.
 (A) Habit, (B) leaf, (C) side view of an umbel, (D) face view of an umbel, (Ei) bract, (Eii) sepals, (F) side view of a dissected ‘pin'-eyed morphs flower, (G) side view of a dissected ‘thrum'-eyed morphs flower, (H) face view of a dissected ‘pin'-eyed morphs flower, (I) face view of a dissected ‘thrum'-eyed morphs flower, (J) white puberulent on the corolla margin, (K) stamen, (L) carpel (left ‘thrum'-eyed morphs, right ‘pin'-eyed morphs).

Primula himalayana B. Hajong, Harsh Singh & P. Bharali sp. nov.
 (A–B) Habit, (C) face view of an umbel, (D) leaf;
Primula gambeliana G.Watt (E) Habit, (F) face view of an umbel, (G) leaf.

Primula himalayana B. Hajong, Harsh Singh & P. Bharali sp. nov.  

Diagnosis: A new species resembling P. gambelina due the appearance of leaves and flowers, but differing in lamina puberulent with a white protuberance on the tip of each teeth (versus glabrous and without any protuberances), petiole 2‒3 times longer than lamina and white-puberulent (versus 3‒5 times longer than lamina, glabrous), scape 15‒35 cm long with 4‒17 flowers in the umbel (versus 6‒15 cm long with 2‒8 flowers), flowers yellow with an orange-yellow center and ca 1.5‒1.9 cm long corolla tube (versus bright reddish-pink or purple-pink to violet-purple with yellow center, ca 1.2 cm long), corolla lobes elliptic (versus obcordate or broadly obovate), anthers basifixed (versus dorsi-basifixed), ovary quadrate–globose without any teeth (versus globose–cylindric with 5 or more apical teeth).

Etymology: The species epithet ‘himalayana' is derived from the Great Himalaya Mountains – a biodiversity hotspot and widely considered as the geographical origin as well as the centre of diversity and distribution of Primula.

Bipankar Hajong, Harsh Singh and Pankaj Bharali. 2026. Primula himalayana sp. nov., A New Species from eastern Himalaya, India. Nordic Journal of Botany. DOI: doi.org/10.1002/njb.05052 [11 March 2026]

[Mammalogy • 2026] Afronycteris rautenbachi • A New Species of African Pipistrelle-like Bat (Chiroptera: Vespertilionidae: Afronycteris)

Afronycteris rautenbachi

Kearney, de Vries & Markotter, 2026 

DOI: doi.org/10.11646/zootaxa.5768.1.1

Abstract 

The taxonomy of the small, sub-Saharan, insectivorous bat, Afronycteris helios (Heller, 1912), has been unresolved for decades. The name A. cf. helios was introduced in the literature to recognise bats found in east and southern Africa that were like A. helios but had glands on the uropatagium. Cranio-dental morphology, bacular morphology, and molecular genetics (albeit the latter two being without representation of A. helios), provided evidence to formally describe “A. cf. helios”, which is currently known from Kenya, Mozambique, and South Africa. Bayesian analyses based on cytochrome oxidase b, cytochrome c oxidase subunit 1, and 12S rRNA confirm that it belongs to the genus Afronycteris with A. nanus (Peter, 1852) and A. helios. These analyses also revealed genetic, bacular, and cranio-dental morphological differences within A. nanus, which are described here. Pending a more thorough geographic analysis, including all existing synonyms, A. cf. nanus has been introduced to refer to bats that were smaller than A. nanus. Although co-occurring in north-eastern parts of South Africa, A. cf. nanus has a more westerly distribution extending to west Africa, relative to the more easterly distribution of A. nanus. Afronycteris sp. nov. showed at least a 6.9%, 3.4% and 2.9% nucleotide difference to its nearest relative based on cytochrome oxidase b, cytochrome c oxidase subunit 1, and 12S rRNA respectively. It is cranio-dentally smaller than A. helios, slightly smaller than A. nanus, and larger than A. cf. nanus, with a distinct baculum, and a unique pair of glands on the uropatagium.

Mammalia, molecular genetics, taxonomy, morphology, Afronycteris nanus, Afronycteris helios, Kruger National Park

Lateral head and shoulder view of Afronycteris rautenbachi in the hand, of the holotype TM 48535 (left), and TM 48537 (right), both from South Africa, Limpopo province, Kruger National Park, Makuleka Contract Park.

Images of ears (above) and tragi (below) of  Afronycteris rautenbachi, TM 48537 (left) and A. cf. nanus TM 48572 (right), both from South Africa, Limpopo province, Kruger National Park, Makuleka Contract Park.

Dorsal ventral and lateral views of the cranium and a lateral view of the mandible of the holotype, TM 48535, of Afronycteris rautenbachi from South Africa, Limpopo province, Kruger National Park, Makuleka Contract Park. All images were scaled to the same size. Scale bar = 3 mm.

Afronycteris rautenbachi sp. nov.

Diagnosis. Afronycteris rautenbachi is a small Vespertilionidae and among the smallest of the Vespertilionini, being most similar in size and appearance to A. nanus, A. cf. nanus and A. helios. Afronycteris rautenbachi has a pair of glands on the uropatagium on either side of the tail, near the body (Figure 7).

Etymology. This species is named in honour of Igantius (“Naas”) L. Rautenbach (1942–2024) who was head of the mammal section at Ditsong National Museum of Natural History (formerly Transvaal Museum) from 1968 to 1991, and then the director of the museum from 1991 to 1999. He conducted extensive field research on southern African mammals, including the northern part of the Kruger National Park, which considerably increased the size of the museum collection. Over the years he managed various research collaborations that started with field collection and resulted in numerous publications. The proposed English common name is Kruger tail-gland bat.

Teresa KEARNEY, Marinda DE VRIES and Wanda MARKOTTER. 2026. Description of A New Species of African Pipistrelle-like Bat (Chiroptera: Vespertilionidae: Afronycteris). Zootaxa. 5768(1); 1-28. DOI: doi.org/10.11646/zootaxa.5768.1.1 [2026-03-09]

[Botany • 2026] Sadiria kachinensis (Primulaceae: Myrsinoideae) • A New Species from Myanmar

Sadiria kachinensis Utteridge & Nob.Tanaka,

in Utteridge, Armstrong, Yoneda et Tanaka. 2026.  

DOI: doi.org/10.11646/phytotaxa.744.3.1

 

Abstract

Sadiria kachinensis from Kachin State, Myanmar is described and illustrated as a species new to science. It is morphologically similar to S. boweri and S. solanifolia but differs in several characters especially the smaller leaves on the vegetative shoots and the flowering branches, larger inflorescences and petals, and smaller sepals. In addition, this new species is compared to S. eugeniifolia var. burmanica, the only other member of Sadiria in Myanmar.

 Burma, Ericales, Indo-China, South-East Asia, taxonomy, Eudicots

Sadiria kachinensis Utteridge & Nob.Tanaka.
 A habit; B detail of leaf margin; C inflorescences; D flower; E single calyx lobe; F detail of overlapping corolla lobes; G flower opened to show stamens opposite the corolla lobes; H ovary and style (flower with corolla removed); I stamen abaxial view; J stamen adaxial view; K ovary; L fruit with persistent style.

Drawn from Armstrong et al. 4307 (TNS). Scale bars: A = 3 cm, C, D, G, H, L = 5 mm, E & F = 1 mm, I, J & K = 3 mm. Illustration by Kaoru Yoneda.

Sadiria kachinensis Utteridge & Nob.Tanaka.
A. habit; B. plagiotropic (flowering) branch with inflorescences and flowers pre-anthesis; C. plagiotropic branch with fruits; D. inflorescence/flower detail, note exserted styles in the flowers on the right hand side; E. fruit detail, note persistent style (dry and black).

 Photographed by Kate Armstrong.

Sadiria kachinensis Utteridge & Nob.Tanaka, sp. nov. 

 Recognised in the genus Sadiria by the leaves (ortho- and plagiotropic) drying greenish and the shorter plagiotropic reproductive branches, and unique amongst those species in the combination of the following characters: 3–4 leaves in pseudowhorls along the erect (orthotropic) stems, these orthotropic leaves, elliptic and 9.5–13 × 3.5–4.5 cm; the lateral plagiotropic (reproductive) flower-bearing branches (4–)6.5–11 cm long, with 3(–5) leaves along the length of the branch; orthotropic and reproductive shoots leaves drying green; pendulous inflorescences sessile or with a very short rachis to 2 mm long; the corolla fused for approximately three-quarters of its length; stamens with anthers 2 mm long; and the style conspicuously exserted at anthesis and extending ca. 3 mm beyond the apex of the corolla lobes.

Timothy UTTERIDGE, Kate E. ARMSTRONG, Kaoru YONEDA and Nobuyuki TANAKA. 2026. Sadiria kachinensis (Myrsinoideae, Primulaceae), A New Species from Myanmar.  Phytotaxa. 744(3); 205-211. DOI: doi.org/10.11646/phytotaxa.744.3.1 [2026-03-10]

[Botany • 2026] Nymphanthus vietnamensis (Phyllanthaceae) • A New Species from Central Vietnam

 

Nymphanthus vietnamensis T.A.Le, Tak.Yamam. & Tagane, 

in Vu, T. A. Le, Nong, Dinh, V. H. Le, Yamamoto et Tagane, 2026. 

DOI: doi.org/10.11646/phytotaxa.742.3.6 

 facebook.com/LeTuanAnhBotany

 

Abstract

A new species of Nymphanthus, N. vietnamensis T.A.Le, Tak.Yamam. & Tagane is described from Quang Tri Province and Thua Thien Hue Province (currently Hue City), central Vietnam. It is similar to N. namkadingensis but distinguished by its branchlet with 14–30(–40) leaves (vs. 50–60 in N. namkadingensis), leaf blades with 3–6 pairs of secondary veins (vs. 8–12 pairs), pistillate pedicel of 8–9 mm long (vs. 10–16 mm long), disc of pistillate flowers with 4–6 free rectangular segments (vs. connate and annular), and subglobose capsules (vs. ellipsoid). A detailed description, photographs, and information on the distribution and ecology, vernacular name, phenology, and preliminary conservation status are provided for the species.

Flora of Vietnam, Dakrong, Phong Dien, Phyllantheae, taxonomy, Eudicots

Nymphanthus vietnamensis T.A.Le, Tak.Yamam. & Tagane sp. nov. 

TIEN CHINH VU, TUAN ANH LE, VAN DUY NONG, DIEN DINH, VAN HUONG LE, TAKENORI YAMAMOTO, SHUICHIRO TAGANE. 2026. Nymphanthus vietnamensis, A New Species of Phyllanthaceae from Central Vietnam.  Phytotaxa. 742(3); 237-242. DOI: doi.org/10.11646/phytotaxa.742.3.6 [2026-02-24]

  facebook.com/LeTuanAnhBotany/posts/3501787813307958

[Invertebrate • 2025] Quapawjapyx osage & Holjapyx nimiipuu • New Cave japygids (Diplura: Japygidae) from North America

Quapawjapyx osage Sendra gen. et sp. nov.

Holjapyx nimiipuu Sendra sp. nov.,

in Sendra, Jiménez-Valverde, Selfa et Cupello, 2026. 

DOI: doi.org/10.5852/ejt.2026.1043.3217 

Abstract

We studied a significant collection of Japygidae (Diplura) deposited in the Texas A&M University Insect Collection, sampled from several caves in North America, mostly in the early 21st century by a group of American speleologists. Among this biological material, a new genus and species, Quapawjapyx osage Sendra gen. et sp. nov., is described from four caves in Arkansas, and a new species, Holjapyx nimiipuu Sendra sp. nov., from five caves in Idaho; both are named in honor of native American tribes. Quapawjapyx osage can be distinguished from Indjapyx, Paurojapyx, and Parindjapyx – which share symmetrical cerci – by the combination of the shape and disposition of the glandular and sensory setae on the first urosternite, together with differences in cercal ornamentation. Holjapyx nimiipuu is characterized by the presence of two conspicuously large predental denticles on the right cercus. Both new taxa of cavernicolous japygids show slight cave-adaptation features, such as large body size, slight elongated appendages, and a small increase in the placoid sensilla of last antennomere. These two new taxa double the number of known cave-adapted japygids in North America, a seemingly low figure that is nonetheless comparable to other karst regions worldwide.

Keywords: Japygoidea, cave-adapted fauna, Nearctic region, taxonomy

Class Diplura Börner, 1904

Superfamily Japygoidea Ewing, 1942

Family Japygidae Haliday, 1864

Quapawjapyx osage Sendra gen. et sp. nov.
A. Paratype, ♂ (TAMU-ENTO X1831026). B, D–E. Paratype, ♂ (TAMU-ENTO X1832015). C. Holotype, ♀ (TAMU-ENTO X1831147).
A. Habitus. B. Last antennomere; placoid sensilla remarked by spotted line. C. Lacinia with interior laminae and mandible. D. Dorsal portion on third antennomere with pores. E. Metathoracic claws.

Genus Quapawjapyx Sendra gen. nov.

Etymology: The generic name is in honor of the Quapaw Nation of Native Americans, who thrived along the lowlands of the Arkansas river five hundred years ago. Today, they live alongside much later European settlersfrom other nations.

Quapawjapyx osage Sendra gen. et sp. nov.

Etymology: The specific epithet is in honor of the Osage Nation of Native Americans, who now reside in Osage County, Oklahoma. They arrived and thrived a few hundred years ago in the northern mountains of Arkansas.

Holjapyx nimiipuu Sendra sp. nov., holotype,  ♀ (TAMU-ENTO X1831140).
A. Habitus. B. Last antennomere. C. Distal portion of urite X including cerci. D. Left cercus. E. Right cercus; placoidsensilla remarked by dash line.

 Holjapyx nimiipuu Sendra sp. nov.

Etymology: ‘Nimiipuu’ (‘we, the people’) is the name by which the Nez Perce Native American tribe refers to themselves. They have thrived in  the Pacific Northwest of North-America for  more than ten   thousand years and today live primarily on their tribal reservation in Idaho. We dedicate this new species to the Nimiipuu people and to their enduring struggle for survival.

Alberto Sendra, Alberto Jiménez-Valverde, Jesús Selfa and Mario Cupello. 2026. New Cave japygids (Diplura: Japygidae) from North America. European Journal of Taxonomy. 1043(1); 166–198. DOI: doi.org/10.5852/ejt.2026.1043.3217 

 

[Herpetology • 2026] Takydromus ultapaniensis • A New Species of Takydromus (Squamata: Lacertidae) from northeast India

 

Takydromus ultapaniensis

Chettri, Mothey, Ghatani, Deepak & Rai, 2026

DOI: doi.org/10.11646/zootaxa.5763.2.2 

 Researchgate.net/publication/401476446

Abstract

The diversity of lizards in the northeast Indian biodiversity hotspot is underestimated, as evidenced by many new species recorded in recent years. Here, we describe a new species of Takydromus from Ultapani, Kokrajhar District, Assam, India. This newly identified species is distinguished from its relatives in its distinctive morphological features and significant genetic variation in the mitochondrial 12S rRNA and COI genes. We have also added the missing molecular data for T. sikkimensis and T. khasiensis. In the light of these new data, we discuss the phylogenetic relationships of Takydromus species from the Indian subcontinent with their congeners.

Reptilia, Asian grass lizard, genetic divergence, phylogeny, range expansion, Takydromus, Ultapani

Takydromus ultapaniensis sp. nov. 

BASUNDHARA CHETTRI, RABINA MOTHEY, SUDEEP GHATANI, V. DEEPAK and ANANTA RAI. 2026. A New Species of Takydromus (Squamata: Lacertidae) from northeast India.  Zootaxa. 5763(2); 205-224. DOI: doi.org/10.11646/zootaxa.5763.2.2 [2026-03-03] 

Researchgate.net/publication/401476446_A_new_species_of_Takydromus_from_northeast_India

[Invertebrate • 2026] Dalhousiella yabukii & Leocratides watanabeae • Single Origin and Convergent Host Use of hexactinellid Sponge Symbiosis in Hesionidae (Annelida: Polychaeta) with Descriptions of Two New Deep-sea Species

Dalhousiella yabukii  &  Leocratides watanabeae 

 Jimi, Downey, Mella & Hookabe, 2026

 DOI: doi.org/10.1093/zoolinnean/zlag028  

 x.com/nu_mbl

Abstract

Symbiotic associations between annelids and sponges are widespread and have evolved repeatedly across diverse families. However, their evolutionary history within Hesionidae remains poorly understood. Using the manned submersible Shinkai 6500, we conducted in situ observations and collected hesionid specimens from a hexactinellid sponge on the Getsuyo Seamount, north-west Pacific Ocean. An integrative taxonomic approach combining morphology and molecular phylogenetics revealed that they belong to two new species, Dalhousiella yabukii sp. nov. and Leocratides watanabeae sp. nov. Phylogenetic analyses suggest that sponge symbiosis in Hesionidae originated once, in the common ancestor of Dalhousiella and Leocratides. Nevertheless, the two new species are not each other’s closest relatives; instead, they form independent sister-clades, each including different congeners. Specimens of the two new species inhabited the same sponge host and probably shared similar ecological niches, suggesting convergent ecological specialization toward the limited resource of hexactinellid sponges in the deep sea. Such niche overlap and convergent evolution between closely related symbiotic species have not been previously reported in Hesionidae. Our findings shed new light on the origin and diversification of sponge symbiosis in Hesionidae and highlight the importance of integrative taxonomy and direct in situ observation for understanding deep-sea biodiversity.

Hesionini, taxonomy, symbiosis, Polychaeta, seamount

Dalhousiella yabukii sp. nov.  

Leocratides watanabeae sp. nov. 

Naoto Jimi, Rachel Downey, María Belén Arias Mella and Natsumi Hookabe. 2026. Single Origin and Convergent Host Use of hexactinellid Sponge Symbiosis in Hesionidae (Annelida: Polychaeta) with Descriptions of Two New Deep-sea Species. Zoological Journal of the Linnean Society. 206(3); zlag028. DOI: doi.org/10.1093/zoolinnean/zlag028 [09 March 2026] 

 x.com/nu_mbl/status/2030905993211810299

[Herpetology • 2021] Hemidactylus cinganji, H. faustus, ... • Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820 (Squamata, Gekkonidae), with Description of Four New Species

  Hemidactylus cinganji
H. carivoensis
H. faustus 
H. pfindaensis 

 Lobón-Rovira, Conradie, Iglesias, Ernst, Veríssimo, Baptista & Vaz Pinto, 2021 

       

 DOI: 10.3897/vz.71.e64781 

 facebook.com/JavierLobonRovira

 

Abstract

The taxonomy of Angolan Hemidactylus has recently been revised. However, the lack of fresh material for some groups and regions, has led to the misidentification of some taxa and an underestimation of actual diversity in others. To shed light on the evolutionary history and systematics of Angolan Hemidactylus, we generated a new phylogenetic hypothesis for the group, and updated the taxonomy following an integrative approach. This resulted in the description of four new species (Hemidactylus pfindaensis sp. nov., H. faustus sp. nov., H. carivoensis sp. nov. and H. cinganji sp. nov.), the reevaluation of two recently described species (H. vernayi and H. paivae) and the synonymization of a recently described species (H. hannahsabinnae). We estimate divergence times for these lineages, providing a preliminary interpretation of their speciation process. Moreover, we suggest and outline 13 Angolan Main Biogeographic Units (AMBUs) in the area, defining a new biogeographic context for future works on Angolan herpetofauna. We now recognize eleven Hemidactylus species in Angola, and we provide here a new morphological key for Angolan Hemidactylus to help with identifications and species assignments in this group.

Keywords: Angola, biogeography, Gekkota, Hemidactylus, systematics, taxonomy

Hemidactylus bayonii-group 

Hemidactylus vernayi Ceríaco, Agarwal, Marques and Bauer 2020

Hemidactylus pfindaensis sp. nov.

Etymology: The name “pfindaensis” derives from the local word “pfinda” which in Kikongo – the main language used in Uíge Province and northwestern Angola – refers to a “gallery forest” or a “continuous block of thick forest”, the main habitat type associated with the species.

Hemidactylus longicephalus-group

Hemidactylus paivae Ceríaco, Agarwal, Marques and Bauer 2020

A – From top, dorsolateral view of Hemidactylus cinganji sp. nov. (FKH0435), H. benguellensis (FKH0413) and H. carivoensis sp. nov. (MNCN 50543);
B – records of H. benguellensis–group (purple color represents records of H. cinganji sp. nov.; blue H. benguellensis; dark green H. carivoensis sp. nov.; circles represent genetically assigned records, while triangles represent historical or non-genetically assigned records by Ceríaco et al. 2020a; color stars depict type or neotype localities; black star depicts original type locality of H. benguellensis assigned by Bocage (1893);
C – habitat of H. cinganji sp. nov. at Ebanga; D – habitat of H. carivoensis sp. nov between Dombe–Echimina. E – habitat of H. benguellensis at Tchivinguiro.

 Photos Javier Lobón–Rovira (A–D) and William R. Branch (E).

Hemidactylus benguellensis-group

Hemidactylus cinganji sp. nov.

Etymology: The name “cinganji” is a widespread traditional word used in Angolan local languages that represents an ancestral spiritual entity that reincarnates assuming different physical forms in different places and occasions. This name is suitable as the new species corresponds to a taxon that was first described under a different name, then became lost and now resurfaces after its original name had been hijacked by a surrogate sister-species. The species epithet is used as a neuter singular noun in opposition to the generic name.

Hemidactylus carivoensis sp. nov.

Etymology: The species epithet “carivoensis” refers to the Farm Carivo, an old estate situated along the banks of the mid-lower Coporolo River on the coastal plain of Benguela Province, and where most of the type series was collected. The species proved to be common in the area, and by recognizing the farm, we also acknowledge the ongoing support from the owners to researchers, similar to the Chapmans nearly a century ago.

Species not assignable to specific species groups

Hemidactylus faustus sp. nov.   

  

Etymology: The name “faustus” applies to a Latin word that designate ‘good luck’, evoking the serendipitous discovery of this species. The species epithet is used as a masculine adjective singular. The first specimen was found by Beatriz Vaz Pinto, daughter of PVP, under a small stone which was removed while preparing a campsite. This unexpected find led to further collecting of this new and previously unrecorded form, albeit from a locality that had been regularly surveyed since the mid-19th century.  

 

 Javier Lobón-Rovira, Werner Conradie, David Buckley Iglesias, Raffael Ernst, Luis Veríssimo, Ninda Baptista and Pedro Vaz Pinto. 2021. Between Sand, Rocks and Branches: An Integrative Taxonomic Revision of Angolan Hemidactylus Goldfuss, 1820, with Description of Four New Species. Vertebrate Zoology. 71: 465-501. DOI: 10.3897/vz.71.e64781

  facebook.com/JavierLobonRovira/posts/3969307526513233

[Diplopoda • 2026] Riukiaria langyaensis • A New Species of the Genus Riukiaria (Polydesmida: Xystodesmidae) from eastern China, with the characterization of its complete mitochondrial genome

Riukiaria langyaensis  Han & Wang, 

in Han, Liu, Wang et Dong, 2026. 

DOI: doi.org/10.3897/zookeys.1272.182977

Abstract

This study describes a new millipede species of the genus Riukiaria from China: Riukiaria langyaensis sp. nov., and provides an identification key to all 19 Riukiaria species currently known from China. A systematic review was conducted on the 34 described Riukiaria species, and these species were classified into five categories based on the morphology of the male gonopod. The complete mitochondrial genome of this new species was sequenced and annotated for the first time, representing the first mitogenomic data reported for the genus Riukiaria. The mitogenome of R. langyaensis sp. nov. is 14,954 bp in length, containing 13 protein-coding genes, 22 tRNA genes, 2 rRNA genes, and a control region, with an overall A+T content of 65.8%. All 37 mitochondrial genes are encoded on the Heavy strand. The results help clarify the confused species relationships within Riukiaria and provide fundamental data for phylogenetic studies of Xystodesmidae.

Key words: Complete mitochondrial genome, identification key, millipede, mitogenome, Riukiaria, species description, taxonomy

Riukiaria langyaensis sp. nov. 
A. Live coloration; B. Live specimen under UV light.

Riukiaria langyaensis sp. nov. 
A. Posterior part of body, dorsal view; B. Midbody rings, dorsal view; C. Anterior part of body, dorsal view; D. Telson, ventral view; E. Both gonopods, ventral view; F. Rings 16–19 and telson, dorsal view. Scale bars: 1.0 mm.

Riukiaria langyaensis Han & Wang, sp. nov.

  

Diagnosis. This species closely resembles Riukiaria mundyi Korsós, Nakamura & Tanabe, 2011 in tergal features and body coloration. However, Riukiaria langyaensis sp. nov. differs by having a lighter body color than R. mundyi, with antennae and walking legs exhibiting a wheat-colored hue and the paratergal margins appearing translucent. In terms of gonopod structure, the prefemorite of Riukiaria langyaensis sp. nov. is similar to that of Riukiaria contigua (Wang, 1957), but its acropodite is more inflated and nearly spherical, with an indistinctly forked tip, and the prefemoral processes are completely missing.

 Zewu Han, Xin Liu, Jiajia Wang and Yan Dong. 2026. Description of A New Species of the Genus Riukiaria (Diplopoda, Polydesmida, Xystodesmidae) from eastern China, with the characterization of its complete mitochondrial genome. ZooKeys. 1272: 315-335.  DOI: doi.org/10.3897/zookeys.1272.182977 [10 Mar 2026]

[Ichthyology • 2026] Oreichthys elianae & Glyptothorax sentimereni • New Species (Teleostei: Sisoridae, Cyprinidae) from Nagaland, Northeast India

 

Oreichthys elianae  &

Glyptothorax sentimereni

 Praveenraj & Vijayakrishnan, 2026

DOI: doi.org/10.11646/zootaxa.5763.2.5

Abstract

Two new species of fishes are described from the tributaries of the Brahmaputra River in Nagaland, Northeast India. Glyptothorax sentimereni, described from the Dikhu River, is distinguished from congeners in having the following combination of characters: ventral surfaces of pectoral spine and first pelvic-fin ray with plicae, serrated posterior edge of dorsal-fin spine, thoracic adhesive apparatus completely encinctured by striae, length of dorsal-fin spine 13.7–16.1% SL, body depth at anus 17.1–19.6% SL, length of dorsal-fin base 12.0–13.1% SL, post-adipose distance 20.8–22.2% SL, head depth 16.5–17.0% SL and dorsal-to-adipose distance 21.8–23.5% SL. Oreichthys elianae, described from the Tsurang River, is distinguished from its congeners by a combination of characters that include 19+1 scales (with 5 pored scales) in longitudinal series, ½3/1/2½ scales in transverse row, 9 predorsal scales, 14–15 pores on cheek, a black spot on the caudal-fin base, and reddish fins.

Pisces, Brahmaputra River, Cypriniformes, Diversity, endemic fish, Siluriformes

 

Glyptothorax sentimereni, a rheophilic torrent catfish from the Dikhu River, Nagaland

Oreichthys elianae, from the Tsurang River in Mokokchung, loses its coloration immediately under stress

Jayasimhan PRAVEENRAJ and Balaji VIJAYAKRISHNAN. 2026. New Species of the Genera Glyptothorax (Teleostei: Sisoridae) and Oreichthys (Teleostei: Cyprinidae) from Nagaland, Northeast India.  Zootaxa. 5763(2); 255-267. DOI: doi.org/10.11646/zootaxa.5763.2.5 [2026-03-03]

[Paleontology • 2026] Tanyka amnicola • An aberrant Stem Tetrapod from the early Permian of Brazil

 

Tanyka amnicola

 Pardo, Marsicano, Smith, Cisneros, Angielczyk, Fröbisch, Kammerer & Richter, 2026

 

DOI: doi.org/10.1098/rspb.2025.2106

Illustration by Vitor Silva

Abstract

Early evolutionary history of tetrapods is typically divided into two major phases: an initial diversification of archaic stem tetrapod groups, and a sudden replacement by temnospondyl amphibians and amniotes following a late Carboniferous dry interval termed the Carboniferous Rainforest Collapse (CRC). However, the degree to which this scenario applies to the early tetrapods of Gondwana is uncertain. Here, we report Tanyka amnicola, gen. et sp. nov., an archaic stem tetrapod from the early Permian of Brazil characterized by strong torsion of the mandibular ramus and a remarkable battery of enlarged denticles on a strongly arched coronoid. The new taxon is assigned to the tetrapod stem based on the presence of a denticulate adsymphyseal and elevated hook-like glenoid surface without a postglenoid area. Phylogenetic analysis shows affinities between this species and stem tetrapods more proximal to the tetrapod crown group, particularly Eucritta and the Laurussian baphetids. This is the second stem tetrapod group known to have survived until the end of the early Permian in Gondwana despite local extirpation in Laurussia, implying that current hypotheses of Carboniferous tetrapod turnover are oversimplified. The unique jaw morphology suggests adaptations to either specialized processing of small invertebrates or consumption of some plant material, demonstrating that stem tetrapods continued to explore new niche space into the Permian of Gondwana.

Keywords: Permian, Tetrapoda, palaeobiogeography

Holotype jaw of Tanyka amnicola, MAP-PV 662.
(a) MAP-PV 662 in dorsal view; (b) interpretive drawing of MAP-PV 662 in dorsal view;
(c) MAP-PV 662 in ventral view; (d) interpretive drawing of MAP-PV 662 in ventral view.

adsym, adsymphyseal; an, angular; ar, articular; c1, first coronoid; c2, second coronoid; c3, third coronoid; ct, foramen for chorda tympani; d, dentary; laf, lateral angular flange; Mf, Meckelian fenestra; par, prearticular; pMf, pre-Meckelian foramina; pspl, postsplenial, sa, surangular; sfp, symphyseal fang pair of dentary; spl, splenial.

Tetrapodamorpha Ahlberg, 1991

Tanyka amnicola gen. et sp. nov.

Holotype. MAP-PV 662, nearly complete left mandible. Accessioned at Museu de Arqueologia e Paleontologia (MAP) at Universidade Federal do Piauí, Teresina, Brazil.

Type locality and horizon. Pedra de Fogo riverbank, MAP locality PB 156, south of Pastos Bons, Maranhão, lower Pedra de Fogo Formation, early Permian.

 

Etymology. Guarani: tañykã, meaning ‘jaw’ or ‘chin’, and Latin: amnicola, meaning ‘living in or next to the river’ the latter being a reference both to the river bed in Pastos Bons where the holotype was found and also to the presumed aquatic habits of the stem tetrapod.

Diagnosis. Stem tetrapod of moderate size. Adsymphyseal and all coronoids covered in a thickly ankylosed and arched denticle pad much wider than the tooth row. Occlusal surface of jaw, including coronoid denticle plate, faces more labially than dorsally when jaw is in neutral position. Ventral margin of jaw ramus wider than dentary-coronoid area. Angular with prominent sculptured flange. Prearticular braces articular medially with a large triangular process. Meckelian foramen small (less than one-third the depth of the prearticular).

Jaw rotation during mandibular adduction in Tanyka.
(a) Jaw in closed position, lateral view; (b) jaw in closed position, anterior view;
(c) jaw in open position, lateral view; (d) jaw in open position, anterior view.

General skull shape speculative, based on Baphetes

Illustration showing Tanyka amnicola in life, eating underwater plants.

 Artwork: Vitor Silva

 

Jason D. Pardo; Claudia A. Marsicano; Roger Smith; Juan Carlos Cisneros; Kenneth D. Angielczyk; Jörg Fröbisch; Christian F. Kammerer; Martha Richter. 2026. An aberrant Stem Tetrapod from the early Permian of Brazil. Proc Biol Sci. 293 (2066): 20252106. DOI: doi.org/10.1098/rspb.2025.2106 [04 Mar 2026]

  

[Herpetology • 2026] Sphenomorphus lingnanensis • A New skink Species of the Genus Sphenomorphus Fitzinger, 1843 (Squamata: Scincidae) from Central Guangdong, South China

Sphenomorphus lingnanensis

Guo, Zhong, Li, Xu, Wang, Li, Yu & Ji, 2026 

  

DOI: doi.org/10.3897/zse.102.183764

Abstract

We collected nine specimens of the genus Sphenomorphus Fitzinger, 1843 (Squamata, Scincidae) from Liuxihe National Forest Park in central Guangdong, South China, which were found not to correspond to any currently recognized species. Here, we describe this new species, Sphenomorphus lingnanensis sp. nov., integrating morphological and mitochondrial DNA (COI and 16S rRNA) evidence. Phylogenetic analyses reveal the new species to be a sister taxon to S. indicus with uncorrected p-distances of 0.132 in COI and 0.021 in 16S rRNA. The p-distance in COI between the new species and S. indicus exceeds that between S. scutatus and S. solomonis (0.057), and the p-distance in 16S rRNA exceeds that between S. cyanolaemus and S. sabanus (0.005). Morphologically, this new species is distinguished from five congeners (including S. indicus) with comparable data by: (1) possessing 56–58 ventral scale rows (versus 65–69 in S. indicus); and (2) occupying a distinct position in the morphospace defined by the first two principal components derived from seven meristic variables. The description of S. lingnanensis sp. nov. raises the number of recognized Sphenomorphus species in China to nine. Sphenomorphus lingnanensis sp. nov. is currently known only from two proximate localities (approximately 5 km apart) in central Guangdong, where it is sympatric with the oviparous congener S. incognitus.

Key Words: Guangdong, mitochondrial DNA, morphology, new species, Scincidae, South China, Sphenomorphus, taxonomy

Sphenomorphus lingnanensis sp. nov. from Central Guangdong, China.
The adult female holotype (A, WZULXHTXSP1), a juvenile paratype of (B, WZULXHTXSP2)

Sphenomorphus lingnanensis sp. nov.
Close-up views of the adult female holotype (WZULXHTXSP1; left, from A1–D1) and
a juvenile paratype (WZULXHTXSP3; right, from A2–D2) 

Sphenomorphus lingnanensis sp. nov.

 Kun Guo, Jun-Jie Zhong, Yi-Peng Li, Wang Xu, Wei-Min Wang, Yu-Dong Li, Xin-Fan Yu and Xiang Ji. 2026. A New skink Species of the Genus Sphenomorphus Fitzinger, 1843 (Squamata, Scincidae) from Central Guangdong, South China. Zoosystematics and Evolution. 102(2): 411-420. DOI: doi.org/10.3897/zse.102.183764 [05 Mar 2026]