[Herpetology • 2026] Amolops ammachi, A. bella & A. sendenyu • Systematics, Molecular Phylogeny, Diversification, and Biogeography of Cascade Frogs (Anura: Ranidae: Amolops) from India: An Integrative Assessment with Description of Three New Species

 

 the Amolops marmoratus group from India

A, B. Amolops afghanus. C. Amolops ammachi sp. nov.. D–F. A. assamensis. G–I. Amolops bella sp. nov.. J–M. A. gerbillus. N, O. A. indoburmanensis; 

A. Amolops indoburmanensis haplo 1. 

C. A. cf. indoburmanensis sp. 1.  E. A. jaunsari. 

F, A. nepalicus. H. Amolops sendenyu sp. nov. 

Biju, Ellepola, Sarmah, Upadhyaya, Bisht, Tajo, Hanken & Garg, 2026

DOI: doi.org/10.3099/MCZ88.1

Photos: S. D. Biju.

 Abstract 

The Cascade Frog genus Amolops, currently comprising 95 described species, is a taxonomically challenging group of Asian frogs. Although the genus has been studied extensively across most of its range, the Indian members remain poorly understood, with sporadic species descriptions and no comprehensive taxonomic revision to date. Additionally, the current taxonomy of Amolops in India remains in flux; the true species diversity is potentially underestimated due to insufficient sampling efforts, misidentifications arising from limited investigations, and lack of a comprehensive comparative framework. In this study, we addressed these gaps through extensive sampling of Amolops across their range in India and by using an integrative morphological and molecular approach to investigate existing diversity. By using a multilocus data set of mitochondrial (16S, CO1, ND2) and nuclear (RAG1, TYR) markers, we present the first comprehensive Amolops phylogeny, including all known Indian species, which enables us to confidently establish species boundaries and infer evolutionary relationships. By assigning available names to well-supported lineages and testing multiple species-delimitation hypotheses, we identified a high level of undescribed cryptic diversity in the genus. Integrative analyses confirm three distinct unnamed lineages, which are formally described here as Amolops ammachi sp. nov., Amolops bella sp. nov., and Amolops sendenyu sp. nov. In addition, we report five new country records for India: three genetically conspecific with Amolops afghanus, Amolops nepalicus, and Amolops wangyali, and two shallowly divergent populations of A. medogensis and A. bellulus. Conversely, five nominal species do not represent distinct taxa and are here treated as junior subjective synonyms: Amolops chanakya synonym of Amolops wangyali; Amolops mahabharatensis synonym of Amolops jaunsari; Amolops senchalensis synonym of Amolops formosus; Amolops shillong synonym of Amolops assamensis; and Amolops terraorchis synonym of Amolops gerbillus. Molecular species delimitation reveals six additional candidate species from India; these lineages are left unnamed due to shallow divergence from established species and warrant further taxonomic investigation. Our time-calibrated phylogenetic analyses suggest that Amolops diversification began around the Eocene-Oligocene boundary, with a mean age of 31.5 Mya (median age: 32.5 Mya; 95% highest posterior density: 28.12–36.67 Mya). The most recent common ancestor (MRCA) of extant Amolops was widely distributed across Southeast China, mainland Southeast Asia, and the Malayan Peninsula during the early Oligocene. Through subsequent range contraction, subdivision, and jump dispersal, they spread into adjacent regions with dramatic rapid diversification toward the late Miocene. Biogeographic analyses suggest multiple waves of east-to-west colonization and highlight the role of early Himalayan uplift in mediating a complex “into-and-out-of-Himalaya” dynamic in structuring the contemporary distribution and high cryptic diversity in the genus. Mitochondrial haplotypes also show strong geographic structuring, indicating historical allopatric divergence by secondary contact among regions. These findings additionally improve our broader understanding of diversification and distribution patterns in this unique group.

KEYWORDS: Amphibia, Anura, cryptic diversity, evolutionary history, Integrative taxonomy, Himalayas, South and Southeast Asia, species delimitation, monticola Group, viridimaculatus Group, marmoratus Group, tadpole morphology

 

Dorsolateral views of species within the Amolops marmoratus group from India, in life.
A, B. A. afghanus (SDBDU 2023.6459 and SDBDU 2021.322, respectively). C. Amolops ammachi sp. nov. (ZSI A 16551). D–F. A. assamensis (SDBDU 2022.1100, SDBDU 2018.3843, and SDBDU 2025.6858, respectively). G–I. Amolops bella sp. nov. (SDBDU 2011.6959, SDBDU 2011.6913, and SDBDU 2011.6956, respectively). J–M. A. gerbillus (SDBDU 2024.6597, SDBDU 2024.6726, SDBDU 2021.1225, and SDBDU 2024.6598, respectively). N, O. A. indoburmanensis (SDBDU 2018.3870 and SDBDU 2022.5572, respectively). Photos: S. D. Biju.

Amolops ammachi sp. nov.

Etymology. The species is named in honor of the late Ms. Padmini Varkey, a renowned social worker and philanthropist from Kerala, India. She is fondly referred to as “Ammachi,” meaning “mother” in Malayalam, the language of Kerala. Her unconditional love and readiness to help those in need were widely admired. The species epithet ammachi is a tribute to her life and spirit; it is treated as an invariable noun in apposition to the generic name.

Suggested Common Name. Ammachi's Cascade Frog.

Amolops bella sp. nov.

Etymology. The specific epithet bella is from the Latin word bellus, meaning beautiful, and referring to the beauty of the frog. The species epithet is considered an invariable noun in apposition.

Suggested Common Name. Beautiful Cascade Frog.

Dorsolateral views of species within the Amolops marmoratus group from India, in life.
 A. A. indoburmanensis haplo 1 (SDBDU 2024.6670). B. A. indoburmanensis haplo 2, (SDBDU 2021.4629). C. A. cf. indoburmanensis sp. 1 (SDBDU 2009.140). D, E. A. jaunsari (SDBDU 2012.6991 and SDBDU 2012.153, respectively). F, G. A. nepalicus (SDBDU 2026.6929 and SDBDU 2026.6928, respectively). H. Amolops sendenyu sp. nov. (SDBDU 2024.5438). I–K. A. siju (SDBDU 2023.457, SDBDU 2022.1468, and SDBDU 2025.6832, respectively). L. A. yarlungzangbo (SDBDU 2022.6141). Photos: S. D. Biju.

Amolops sendenyu sp. nov.

Etymology. The species is named after Sendenyu Village, Nagaland State, where the type series was collected. The species epithet is considered an invariable noun in apposition.

Suggested Common Name. Sendenyu Cascade Frog.

S. D. Biju, Gajaba Ellepola, A. N. Dikshit Akalabya Sarmah, Radhakrishna Upadhyaya, Karan Bisht, Tage Tajo, James Hanken and Sonali Garg. 2026. Systematics, Molecular Phylogeny, Diversification, and Biogeography of Cascade Frogs (Ranidae: Amolops) from India: An Integrative Assessment with Description of Three New Species. Bulletin of the Museum of Comparative Zoology. 164(1); 1-144. DOI: doi.org/10.3099/MCZ88.1 (11 August 2026)

 

[PaleoIchthyology • 2026] Parastylephorus anatoides • A New fossil from the Eocene of Iran provides new insights regarding the evolution of the morphologically bizarre tube-eye fish

 

Parastylephorus anatoides 

Ridolfi, Schrøder, Bahrami, Yazdi, Roa-Varón, Marramà & Carnevale, 2026

 

DOI: doi.org/10.1002/spp2.70114

Artwork by Lucrezia Bonardo

Abstract

Stylephoriformes are a morphologically bizarre lineage of deep-sea fishes, currently represented by a single extant species, Stylephorus chordatus, the so-called tube-eye fish. The phylogenetic relationships of the Stylephoriformes were extensively debated until recent genomic studies proposed a sister-group relationship with the Gadiformes. The tube-eye fish exhibits a suite of highly derived features, including extreme cranial kinesis, a pipette-like oral apparatus specialized for suction feeding, and an elongate, naked ribbon-like body lacking scales. The evolutionary origin of these unique traits is largely unknown, in part due to the complete absence of a fossil record. Here, we describe the first-known fossil of the order Stylephoriformes, from Bartonian (c. 40 Ma) sediments of the Pabdeh Formation, Iran. On the basis of several unique morphological features (edentulous premaxillae and dentary, moderately enlarged pterygoid elements, reduced opercle, elongate ribbon-like body comprising more than 63 vertebrae, eight pectoral-fin rays, absence of basipterygia, and the presence of a complete series of well-developed lateral-line scales) the specimen is assigned to a new genus and species: Parastylephorus anatoides. The fossil exhibits a body physiognomy, oral apparatus and cranial architecture remarkably similar to those of the extant tube-eye, with the most derived features already present and closely comparable in morphology. This suggests that the new fossil represents a good example of long-term morphological stasis among marine teleosts, documenting the persistence of a highly specialized body plan for more than 40 myr.

Keywords: Parastylephorus anatoides, Bartonian, Iran, Pabdeh Formation, Paracanthopterygii

Parastylephorus anatoides gen. et sp. nov., holotype, IUGM 100897, from the Eocene of Babaheydar (Iran). Scale bar represents 10 mm.

SYSTEMATIC PALAEONTOLOGY

Order STYLEPHORIFORMES sensu Miya et al. 2007

Family STYLEPHORIDAE Swainson 1839

Genus Parastylephorus nov.

 

Derivation of name: The genus name is derived from the Greek prefix para- (παρά), meaning ‘near’, combined with Stylephorus, the name of the only currently recognized stylephoriform genus. The name reflects the shared morphological characteristics with Stylephorus.

 

Remarks: As discussed above, the Stylephoriformes are a lineage currently represented by a single extant species, Stylephorus chordatus, which possesses a number of highly derived anatomical features, including a strongly modified buccal apparatus, extremely reduced first two vertebral centra, and a continuous dorsal fin along the entire dorsal margin of the body. The body plan of Parastylephorus gen. nov. closely resembles that of Stylephorus, although it has a set of distinctive traits. These include the maxilla with a well-developed posterior process, the expanded metapterygoid and ectopterygoid, and the reduced size of the opercular bones. In addition, Parastylephorus gen. nov. possesses body squamation, consisting of thin and delicate lateral-line scales, which are completely absent in Stylephorus.

Parastylephorus anatoides sp. nov.

 

Derivation of name: The name is derived from the Latin word Anas, meaning ‘duck’, and the Greek suffix -oides (-οειδής) meaning ‘resembling’ or ‘having the form of’.

Holotype: IUGM 100897 a partially preserved specimen on a single plate 

 

Artistic reconstruction of Parastylephorus anatoides gen. et sp. nov. 

Note that the tubular eyes are based on the condition characteristic of the extant Stylephorus chordatus. 

Artwork by Lucrezia Bonardo.

Lorenzo Ridolfi, Ane Elise Schrøder, Ali Bahrami, Mehdi Yazdi, Adela Roa-Varón, Giuseppe Marramà and Giorgio Carnevale. 2026. A New fossil from the Eocene of Iran provides new insights regarding the evolution of the morphologically bizarre tube-eye fish. Papers in Palaeontology. DOI: doi.org/10.1002/spp2.70114 [11 August 2026]

[Herpetology • 2026] Lielaphis slashi • A New Species of Papuan Groundsnake (Squamata: Colubridae: Lielaphis) from New Guinea

 Lielaphis slashi  

Huang, Austin, Bernstein, Iova, Roberts, Frederick & Ruane, 2026

DOI: doi.org/10.11646/zootaxa.5866.2.6 

 

Abstract

The recently resurrected genus Lielaphis (Squamata: Colubridae) is comprised of 19 medium-sized, terrestrial snakes collectively regarded as groundsnakes. Although found primarily in New Guinea, a biologically megadiverse region with high conservation priorities, Lielaphis remain poorly studied due to their largely homogenous appearance, elusive nocturnal nature, limited records, and complex taxonomic history, leaving high potential for undescribed species still existing within this group. Using high-throughput sequence data of 12 previously described species of Lielaphis and representatives of its sister taxon, Stegonotus, we apply an integrative approach combining genetic analyses with morphological examinations using vouchered specimens to explore the cryptic diversity of Lielaphis. We place particular emphasis on populations occurring on and adjacent to mainland New Guinea. We identify both molecular and morphological evidence for a new species, Lielaphis slashi sp. nov. from the northeastern lowlands of mainland New Guinea, bringing the number of Lielaphis species to 20, along with a potential range extension of L. aplini. Despite the recent resurgence of interest in New Guinea’s snake diversity, our findings highlight persistent knowledge gaps, emphasizing the need for more comprehensive taxonomic studies of the speciose New Guinea herpetofauna.

Reptilia, Morphology, phylogenetics, Serpentes, systematics, taxonomy

Photographs of holotype Lielaphis slashi sp. nov. LSUMZ92332
showing dorsum (A), venter (B), ventral and dorsal sides of head (C), and right and left lateral sides of head (D).

Black bars equal to 100 mm in (A) and (B), and equal to 10 mm in (C) and (D).

Photographs from T. Huang.

 Life photo of a juvenile  Lielaphis slashi sp. nov. (LSUMZ 97666, paratype).

Photograph from C. C. Austin.

Lielaphis slashi sp. nov. 

Slash’s groundsnake

Diagnosis. A new species of Lielaphis that can be diagnosed from its congeners by the combination of the following characteristics: a reddish-brown unpatterned dorsum with an unmarked white or cream venter that may be barely darker towards the subcaudal scales, 17 scale rows at midbody, absence of apical pits, 165−181 ventral scales and 83−107 divided subcaudal scales, two preocular scales, two postocular scales, eight to nine (primarily eight) supralabial scales, eight to ten (primarily ten) infralabial scales, supralabial and infralabial scales cream to light tan, supralabials darker, infralabials may be slightly mottled anteriorly. 

Etymology. The species epithet is a patronym to honor Mr. Saul Hudson, known popularly and professionally as the musician “Slash,” in recognition for his life-long support of museums and zoos, and especially for his love of snakes. Slash has frequently promoted and acted as an advocate for snakes across various media outlets over many years. Senior author SR also credits much of her scientific writing as having occurred while listening to Slash’s music, including that of Slash’s Snakepit, Guns N’ Roses, and Velvet Revolver

TIANQI HUANG, CHRISTOPHER C. AUSTIN, JUSTIN M. BERNSTEIN, BULISA IOVA, JACKSON R. ROBERTS, JEFFREY H. FREDERICK and SARA RUANE. 2026. A New Species of Papuan Groundsnake (Squamata: Colubridae: Lielaphis) from New Guinea.  Zootaxa. 5866(2); 305-319. DOI: doi.org/10.11646/zootaxa.5866.2.6 [2026-08-11]

[Botany • 2026] Lepidagathis argentea (Acanthaceae) • A New Species from Thailand, and Updated Synopsis Lepidagathis of Thailand

Lepidagathis argentea Kantachot & Chantar., 

 

in Kantachot et Chantaranothai, 2026.

สังกรณีเงิน  |  DOI: doi.org/10.20531/tfb.2026.54.2.07 

Abstract

In this synopsis of the genus Lepidagathis for the Flora of Thailand, nine species are recognized. This includes Lepidagathis argentea, a new species collected from limestone bedrock in Tak Province, and three species newly recorded for Thailand, L. dulcis, L. purpuricaulis and L. thorelii. Lepidagathis chiengmaiensis and L. thyrsiflora, are reduced to synonyms of L. incurva and L. fasciculata, respectively. The following names have been lectotypified: Lepidagathis chiengmaiensis, L. chlorostachya, L. dissimilis, L. dulcis, L. falcata and L. thorelii. A genus description with key to species, nomenclatural notes, distributional data and specimens examined are provided.

Keywords: Lamiales, lectotype, new record, new species, taxonomy

Drawing of Lepidagathis argentea Kantachot & Chantar.
 A. habit, B. inflorescence with flowers, C. external surface of calyx, D. bracteoles, E. sterile bracts, F. gynoecium, G. fertile bracts, H. flower, J. capsule and seeds, K. seed and embryo.

Drawn from Kladwong & Kantachot 2025-1 (KKU) by K. Tuanku.

Lepidagathis argentea Kantachot & Chantar.
A. habitat, B. habit, C.-D. inflorescences with flowers.

Photos by P. Kladwong, W. Kiewbang and S. Suddee.

Lepidagathis argentea Kantachot & Chantar., sp. nov.

Chortip Kantachot and Pranom Chantaranothai. 2026. Lepidagathis (Acanthaceae) of Thailand: New Species and Updated Synopsis. Thai Forest Bulletin (Botany), 54(2), 64–75. DOI: doi.org/10.20531/tfb.2026.54.2.07  [2026-07-02] 

[Botany • 2026] Hoya sinagtala (Apocynaceae: Asclepiadoideae) • A New, unusual Species from the Philippines

 

Hoya sinagtala Brillo, Cabactulan & Rodda, 

in Brillo, Cabactulan et Rodda. 2026. 

DOI: doi.org/10.11646/phytotaxa.769.3.6  

  facebook.com/groups/philippineplants

 

Abstract

Hoya sinagtala Brillo, Cabactulan & Rodda sp. nov. (Apocynaceae, Asclepiadoideae) from the Philippines is described. When sterile, it resembles several species of Hoya section Acanthostemma, but its flowers make it readily distinguishable by its rotate corolla with a narrowly campanulate tube and free-spreading lobes and triangular-ovate corona lobes. No other species in the genus exhibits this unique combination of characters. The species was originally collected from the wild in Samar Island and subsequently widely distributed in cultivation under the informal name Hoya sp. Samar GN 0003 and Hoya sp. “Yellow Sapphire”; it is herein formally published after nearly seven years of horticultural circulation.

Eudicots, endemic, new species, Samar, taxonomy

Hoya sinagtala Brillo, Cabactulan & Rodda sp. nov.

SAMUEL C. BRILLO, DEREK D. CABACTULAN and MICHELE RODDA. 2026. Hoya sinagtala (Apocynaceae, Asclepiadoideae): A New, Unusual Species from the Philippines.  Phytotaxa. 769(3); 287-293. DOI: doi.org/10.11646/phytotaxa.769.3.6 [2026-08-13] 

 facebook.com/groups/philippineplants/permalink/28049135471405087

[Herpetology • 2026] Boulenophrys calciphila • A New Species of Boulenophrys Fei, Ye & Jiang, 2016 (Anura: Megophryidae) from Karst Landscapes in northern Guangdong, China

 

Boulenophrys calciphila Zeng, Wang, Qi, Song, Lyu & Wang,

in Zeng, J. Wang, Qi, Song, Cai, Zhu, Lin, Li, Zhou, H.-C. Yang, Lyu, Y.-P. Yang et Y.-Y. Wang. 2026. 

DOI: doi.org/10.3897/herpetozoa.39.e202590

Abstract

Karst landscapes across southern China represent global hotspots for vertebrate diversity and endemism, yet amphibian diversity within such habitats remains insufficiently documented. Herein, we describe Boulenophrys calciphila sp. nov., a new species discovered from fragmented karst hills in northern Guangdong Province, China, extending the documented habitat spectrum of the genus in Guangdong to limestone karst terrain. Detailed morphological comparisons coupled with phylogenetic analyses validate its taxonomic distinctness from all known congeners. This new discovery aligns with ongoing records of range-restricted anuran species uncovered from southern Chinese karst formations, reinforcing karst ecosystems as important reservoirs of amphibian diversity. Our results demonstrate that amphibian diversity within poorly-explored karst regions of northern Guangdong is substantially underestimated and call for focused conservation actions to safeguard these microendemic taxa and their unique evolutionary lineages.

Key Words: Boulenophrys calciphila sp. nov., conservation priority, karst habitat, micro-endemism, taxonomy

Life aspect of Boulenophrys calciphila sp. nov.
 the male holotype (GEP a596): lateral view (A1), dorsal view (A2), ventral view (A3), ventral view of hand (A4), ventral view of foot (A5);
and the female paratype (GEP a594): lateral view (B1), dorsal view (B2), ventral view (B3), ventral view of hand (B4), ventral view of foot (B5).

Photo by Zhao-Chi Zeng.

Boulenophrys calciphila Zeng, Wang, Qi, Song, Lyu & Wang, sp. nov.

 

Common name. Limestone Horned Toad (in English) 

 huī yán jiǎo chán (灰岩角蟾 in Chinese).

Diagnosis. (1) Moderate size (SVL 49.7–52.7 mm in two adult males, 56.3–59.2 mm in five adult females); (2) snout rounded in dorsal view, canthus rostralis well developed, tongue not notched distally; (3) tympanum distinct; (4) vomerine ridges present, vomerine teeth absent; (5) dorsal skin smooth with sparse granules, discontinuous X-shaped ridge on centre of dorsum, discontinuous dorsolateral ridges present, sparse large tubercles on flanks, dorsal limbs with discontinuous transverse ridges and tubercles; (6) outer margin of upper eyelid with a prominent horn-like ...

Etymology. The specific epithet calciphila combines the Latin calx, genitive calcis (limestone) with the Greek φίλος (phílos) (= loving), in reference to the association of this species with limestone karst habitats.

 Zhao-Chi Zeng, Jian Wang, Shuo Qi, Han-Ming Song, Yao-Yu Cai, Yun-Sheng Zhu, Shi-Shi Lin, Yuan-Hang Li, Dan-Yang Zhou, Hao-Cong Yang, Zhi-Tong Lyu, Yi-Ping Yang and Ying-Yong Wang. 2026. A New Species of Boulenophrys Fei, Ye & Jiang, 2016 (Anura, Megophryidae) from Karst Landscapes in northern Guangdong, China. Herpetozoa. 39: 291-302. DOI: doi.org/10.3897/herpetozoa.39.e202590 [12-08-2026]

[Paleontology • 2019] Indosinosuchus potamosiamensis • A New freshwater teleosaurid (Thalattosuchia: Teleosauridae) from the Jurassic of northeastern Thailand

Indosinosuchus potamosiamensis

Martin, S. Suteethorn, Lauprasert, Tong, Buffetaut, Liard, Salaviale, Deesri, V. Suteethorn & Claude, 2019

อินโดไซโนซูคัส โปเตโมสยามเอนซิส  || DOI: 10.1080/02724634.2018.1549059 

 Artwork by Kmonvish Lawan

 

ABSTRACT

The core of the fossil record of Teleosauridae, a family of thalattosuchian crocodylomorphs, is well known from western Tethyan marine deposits of the Jurassic. Outside this province, their fossil record is patchy and in need of revision, with specimens from Russia, Madagascar, and Asia. Peipehsuchus teleorhinus is known from the Early or Middle Jurassic of China and teleosaurid specimens have been mentioned or preliminarily described from two Jurassic localities in Thailand, yet they were not assigned to a given taxon. Thanks to recent field work, at least 10 individuals represented by cranial material were excavated and prepared from a single Jurassic locality known as Phu Noi in the lower Phu Kradung Formation of northeastern Thailand. Here, we describe these specimens, together with disarticulated postcranial elements, and erect a new taxon, Indosinosuchus potamosiamensis, gen. et sp. nov. Phylogenetic analyses confirm the teleosaurid affinities of the new species, which does not form an exclusive clade with the Chinese teleosaurid Peipehsuchus teleorhinus. The presence of teleosaurids at Phu Noi and a preliminary account of its faunal content favor a Middle to Late Jurassic age for the fossil-bearing horizon. In contrast, Cretaceous deposits in Thailand are characterized by goniopholidids and pholidosaurids, indicating a faunal turnover in Southeast Asia across the Jurassic–Cretaceous. As previously shown by isotope data, the new teleosaurid species was a resident of the freshwater environment and co-occurs with remains of exclusively terrestrial taxa such as sauropod, ornithopod, and theropod dinosaurs and freshwater tetrapods such as turtles and temnospondyls.

Indosinosuchus potamosiamensis, gen. et sp. nov.

 Jeremy E. Martin, Suravech Suteethorn, Komsorn Lauprasert, Haiyan Tong, Eric Buffetaut, Romain Liard, Celine Salaviale, Uthumporn Deesri, Varavudh Suteethorn and Julien Claude. 2019. A New freshwater teleosaurid from the Jurassic of northeastern Thailand. Journal of Vertebrate Paleontology. 64 (2): 239–260. e1549059. DOI: doi.org/10.1080/02724634.2018.1549059  [27 Feb 2019]

 facebook.com/Dinosaursdiary/photos/1606369629465840 

 twitter.com/Synapsida/status/1101165207257337856

[Paleontology • 2021] Priscaleclercera christae • Taxonomic Revision of Fossil Psilodercidae and Ochyroceratidae Spiders (Araneae: Synspermiata), with A New Species of Priscaleclercera from mid-Cretaceous Burmese Amber, northern Myanmar

 

Priscaleclercera christae

Magalhaes, Porta, Wunderlich, Proud, Ramírez & Pérez-González, 2021

DOI: doi.org/10.1016/j.cretres.2020.104751  

  x.com/6eyes8legs 

 Researchgate.net/publication/348281048

Highlights

• We revise the types of most known psilodercid and ochyroceratid spiders in amber.

• We describe a new Priscaleclercera species from Late Cretaceous Burmese amber.

• Priscaleclercera is a crown-group psilodercid, close to Althepus and Leclercera.

• Late Cretaceous Burmese amber contains 11 species of psilodercids.

• Arachnolithulus, preserved in Dominican amber, is an Ochyroceratinae.

Abstract

Psilodercidae contains some 200 known extant species of small spiders that live in tropical rainforests and caves and is mainly restricted to the Oriental biogeographic realm. Interestingly, at least ten different fossil species have been described from inclusions in Late Cretaceous Burmese amber. This suggests the family has been diversifying in the region for ∼100 million years. In this paper, we revise the taxonomy of fossil species of this family and the closely related Ochyroceratidae, based on the re-examination of type specimens of described species. We find that ten Cretaceous species described in the psilodercid genera Priscaleclercera and Aculeatosoma are valid, and present new illustrations of their type material. The genus Propterpsiloderces apparently does not belong to Psilodercidae, but rather to a family representing a stem lineage within Scytodoidea. Ochyroceratidae is represented in the fossil record by a single ochyroceratine genus from Miocene Dominican amber, Arachnolithulus. The original combination of the extant species Leclercera spinata Deeleman-Reinhold, 1995 is reinstated by transferring it from Priscaleclercera, leaving this latter genus as a strictly fossil taxon. Finally, we present the description of a hitherto unknown species of Priscaleclercera from mid-Cretaceous Burmese amber, Priscaleclercera christae sp. nov., bringing the total number of congeners to ten. By studying its morphological features, we argue that Priscaleclercera is a crown-group Psilodercidae, closely related to the Althepus–Leclercera lineage. The high species diversity of Priscaleclercera indicates that Psilodercidae was already a diverse component of the Oriental tropical forests during the Cretaceous.

Keywords: Aculeatosoma, Arachnolithulus, Cretaceous, Myanmar, Synspermiata

Priscaleclercera christae sp. nov.

 

 Ivan L.F. Magalhaes, Andrés O. Porta, Jörg Wunderlich, Daniel N. Proud, Martín J. Ramírez and Abel Pérez-González. 2021. Taxonomic Revision of Fossil Psilodercidae and Ochyroceratidae Spiders (Araneae: Synspermiata), with A New Species of Priscaleclercera from mid-Cretaceous Burmese Amber, northern Myanmar. Cretaceous Research. In Press, 104751. DOI: doi.org/10.1016/j.cretres.2020.104751 [5 January 2021]

 twitter.com/6eyes8legs/status/1346826645638946825 

 Researchgate.net/publication/348281048_Taxonomic_revision_of_fossil_Psilodercidae_and_Ochyroceratidae_spiders_with_a_new_species_of_Priscaleclercera_from_mid-Cretaceous_Burmese_amber

 

[Botany • 2026] Pterolepis arenacea (Melastomataceae: Melastomateae) • A New Species from the Brazilian Cerrado

Pterolepis arenacea D.Nunes & P.J.F.Guim.,

 in Silva, Figueira, Schindler, Simon et Guimarães, 2026.

DOI: doi.org/10.24823/ejb.2026.2140

Abstract

Pterolepis arenacea is described and illustrated as a new species to the Estação Ecológica Serra Geral do Tocantins and its  surroundings, in the Brazilian states of Bahia and Tocantins. This species is restricted to microhabitats of sandstone outcrops surrounded by savanna vegetation. Pterolepis arenacea is the first known species of the genus exhibiting a dorsal appendage on the stamens. Morphologically, Pterolepis arenacea is most similar to species of the P. polygonoides complex, composed of P. polygonoides, P. saturejiformis and P. trimera. Furthermore, Pterolepis arenacea shares vegetative similarities with the  haplostemonous species P. haplostemona and P. xaxa. Detailed line drawings, field photographs, and scanning electron microscopy images are provided. The conservation status of Pterolepis arenacea is preliminarily assessed as Data Deficient (DD) under the IUCN criteria.

Keywords: Estação Ecológica Serra Geral do Tocantins, Jalapão, Matopiba, Pterolepis polygonoides, sandstone outcrops

 Field images of  Pterolepis arenacea D.Nunes & P.J.F.Guim., sp. nov.
A, Individuals growing in sandstone outcrop cavities; B, detail of a stem, showing the grey-green abaxial surface leaf blades; C, individuals with both flowers and fruits; D, flowering individual; E and F, details of the flowers and floral bud.

Photographs by M. Figueira (A, C and D; all from M. Figueira et al. 1773) and M. F. Simon (B, E and F; all from R.R. Souza et al. 269).

 Pterolepis arenacea D.Nunes & P.J.F.Guim., sp. nov.
 A, Flowering branch; B, a small branched specimen; C, branch node; D, abaxial surface of the leaf blade; D′, detail of the abaxial surface of the leaf blade base; E, adaxial surface of the leaf blade; E′, detail of the adaxial surface of the leaf blade base; F, front view of the flower, exposing the inner surfaces of the petals and the androecium; G, lateral view of the flower, showing the outer surfaces of the hypanthium and perianth; H, inner surface of the petal, highlighting the venation; I–I′′, antesepalous stamens (I, lateral view of the stamen; I′, front view of the stamen; I′′, detail of the dorsal and ventral appendages); J, gynoecium; K, capsule covered by the hypanthium and persistent sepals; L, exposed capsule; M, open capsule exposing persistent placenta.

Drawn by Carla de Lima from R.R. Souza et al. 269 (A and C–M) and M. Figueira et al. 1773 (B).

 Distribution and habitat of  Pterolepis arenacea D.Nunes & P.J.F.Guim., sp. nov. 
A, Distribution map for Pterolepis arenacea (yellow triangles) (Brazilian states: BA, Bahia; MA, Maranhão; TO, Tocantins). B, Aerial view of the vegetation in the Estação Ecológica Serra Geral do Tocantins (EESGT), showing a predominance of savannas and grasslands interspersed with rocky outcrops. C and D, Habitat where Pterolepis arenacea was found in the municipality of Formosa do Rio Preto, Bahia.

Photographs: Sérgio E. Noronha (B) and M. Figueira (C and D).

D. N. Silva, M. Figueira, B. Schindler, M. F. Simon and P. J. F. Guimarães. 2026. PTEROLEPIS ARENACEA (MELASTOMATACEAE: Melastomateae), A New Species from the Brazilian Cerrado.  Edinburgh Journal of Botany. (83); 1-14 pp. DOI: doi.org/10.24823/ejb.2026.2140 [2026-07-13]

 Researchgate.net/publication/409266877_PTEROLEPIS_ARENACEA_A_NEW_SPECIES_FROM_THE_BRAZILIAN_CERRADO

[Herpetology • 2026] Hebius yangi • A New Species of the Hebius atemporalis complex (Serpentes: Natricidae) from southeastern Yunnan, China

Hebius yangi

 Liu, J. Wang, Hou, Deng, Zhang, M. Wang & Rao, 2026

Yang’s Keelback | 杨氏腹链蛇  ||  DOI: doi.org/10.3897/zookeys.1289.189681

Abstract

A new species of the Hebius atemporalis complex is described from Wenshan Prefecture, Yunnan Province, China, based on morphological and molecular evidence. The new species can be distinguished from other species of the complex by the differences in the ratio of tail length to total length, the number of ventral and subcaudal scales, the number of maxillary teeth, and the coloration of the iris and anterior body. Phylogenetically, the new species is separated from other species of the complex by an uncorrected p-distance of 6.0–7.8% in the mitochondrial cytochrome b gene. This study further highlights the cryptic diversity in the H. atemporalis complex and the need to re-examine previous records of H. atemporalis and specimens previously considered H. atemporalis from central-southern China, as well as northeastern Vietnam.

Key words: Complex, cytb, keelback snake, morphology, taxonomy, Wenshan Prefecture

The holotype (KIZ2025058) of Hebius yangi sp. nov. in life.
A. Dorsal view; B. Ventral view.

Close-up views of the head of the holotype (KIZ2025058) of Hebius yangi sp. nov. in life.
A. Dorsal view; B. Ventral view; C. Left view; D. Right view.

Hebius yangi sp. nov.

Diagnosis. Body size relatively small, SVL 237–306 mm; tail moderately long, TaL/TL 0.32–0.33; dorsal scale rows 17-17-17, all keeled except the outermost rows; ventrals 130–131; subcaudals 80–85, paired; supralabials six, third and fourth entering orbit; infralabials seven; maxillary teeth 32–34, gradually enlarged posteriorly; dorsal surface grayish brown to blackish brown with a series of neatly arranged small orange spots on dorsolateral body on each side; ventral surface of head white, scattered with black streaks on scale sutures; iris brownish gray; lip region black and white alternately; neck collar present, discontinuous.

Etymology. The specific epithet is in commemoration of the late renowned Chinese herpetological taxonomist Prof. Dr Datong Yang. We suggest “Yang’s Keelback” as the common English name and “杨氏腹链蛇” as the common Chinese name.

 Shuo Liu, JiShan Wang, Mian Hou, Tao Deng, Chaowu Zhang, Mo Wang and Dingqi Rao. 2026. A New Species of the Hebius atemporalis complex (Squamata, Natricidae) from southeastern Yunnan, China. ZooKeys. 1289: 141-157. DOI: doi.org/10.3897/zookeys.1289.189681 [11-08-2026]