[PaleoEntomology • 2026] Cretolala kachinensis (Hemiptera: Fulgoromorpha: Delphacoidea) • the First lalacid from mid-Cretaceous Kachin Amber: Morphological and Taphonomic Implications

Cretolala kachinensis 

Tang, ŠMÍDOVÁ, Ding, BODERAU, Fabrikant, Mähler, Deng, Nyunt, Fu, Jouault, Tomaschek, Szwedo & Jiang, 2026

 

 DOI: doi.org/10.11646/palaeoentomology.9.2.5

Abstract 

We report Cretolala kachinensis gen. et sp. nov., the first formally described representative of the family Lalacidae from mid-Cretaceous (~99 Ma) Kachin amber. The morphology and taphonomy of the specimen were studied using optical microscopy, micro-computed tomography (micro-CT), confocal laser scanning microscopy (CLSM), and scanning electron microscopy coupled with energy-dispersive X-ray spectroscopy (SEM‒EDS). Cretolala kachinensis gen. et sp. nov. exhibits a unique combination of forewing characters, including a strengthened, broad, and wrinkled costal margin; ScP+R fork proximal to both the Pcu+A1 fusion and the CuA fork; MP with five terminals; and CuA1 and CuA2 each with two terminals. Crossvein patterns differ between the left and right forewings, indicating that crossvein expression alone is not diagnostic. The forewing bears distinct maculation, with a face-like pattern dorsally and an extensive lateral pattern that may indicate aposematism, mimicry, or plant-like camouflage in an arboreal herbivore. Taphonomic analyses based on optical micrographs and material-dependent X-ray attenuation in micro-CT show that minerals are distributed throughout the fossil, coating anatomical surfaces and infilling pre-existing cavities and voids. These phases help stabilize morphology and influence the micro-CT reconstruction of soft-bodied structures. SEM–EDS analyses indicate that the body-coating and void-filling minerals are dominated by quartz and pyrite, with feldspars and aluminosilicate mixtures also present, locally enriched in K, Na, Mg, or Ca. This assemblage and its textures point to a polyphase paragenetic sequence, with early detrital infill and later authigenic mineralization. Our results suggest that minerals and some chemicals derived from the host sediments can be transported into and, in some cases, precipitated within amber inclusions, infilling voids and providing structural support that influences fossil preservation. Collectively, these phases occlude porosity, stabilize morphology, and enhance X-ray contrast, although they locally obscured primary tissue boundaries. Our results highlight multistage diagenesis in Kachin amber and underscore variability among inclusions in both mineral assemblages and the timing of mineralization/diagenetic events.

Hemiptera, Fulgoromorpha, Delphacoidea, Mesozoic, Burmese amber, Planthopper, taxonomy, taphonomy, mineral infilling, wing maculation

Cretolala kachinensis gen. et sp. nov., holotype (HMJ–2016–019).
A–D, Reflected-light micrographs. A, Right lateral view. B, Left lateral view. C, Dorsal view. D, Ventral view. E, F, Micro-computed tomography (micro-CT) reconstructions. E, Dorsal view. F, Ventral view. Scale bars = 1 mm.

Systematic palaeontology

Order Hemiptera Linnaeus, 1758

Suborder Fulgoromorpha Evans, 1946

Superfamily Delphacoidea Leach, 1815

Family Lalacidae Hamilton, 1990

Cretolala Jiang, Boderau, Jouault & Szwedo gen. nov.

  

Etymology. The generic name is formed from the combination of “Cretaceous” and generic name “Lalax”; the latter is derived from the Greek λάλαξ (“croaker”, “caller”), in reference to a lalacid planthopper from the Cretaceous. Gender is feminine.

Diagnosis (provisional, pending confirmation from additional specimens and in combination with specieslevel diagnostic characters). Tegmen with narrow base and distinctly widened apical portion; costal margin strengthened, broad and wrinkled. Pc+CP shifted away from anterior margin near tegmen base, parallel to CA, forming a narrow, sclerotized costal area. ScP+R forks basad both CuA fork and Pcu+A1 fusion. RA with two terminals; RP with three terminals; MP with five terminals; CuA1 and CuA2 with two terminals each. 

Cretolala kachinensis Jiang, Boderau, Jouault & Szwedo sp. nov.  

Material. Holotype: HMJ–2016–019, adult female in Kachin (Burmese) amber from northern Myanmar. Repository: Myanmar Gems Museum, Nay Pyi Taw, Myanmar. Condition: the specimen is incomplete, with the head and the dorsal part of the thorax missing. 

Etymology. The specific epithet is derived from “Kachin”, the name of the state in northern Myanmar where the amber originates, and is Latinized with the adjectival suffix -ensis, meaning from “Kachin”. 

Diagnosis. As for the genus, due to monotypy. Additional features include forewing coloration and venation details: costal margin with a continuous dark band along costal area; membrane bearing nearly continuous aligned patches and short transverse bands; a discrete insular macula on ScP+RA; a sub-elliptical isolated macula near distal Pcu; distad of forks of RP and MP, pigment is vein-following, tracking adjacent longitudinal and crossveins; postnodal cell centers unpigmented, ...

 Life reconstruction of Cretolala kachinensis gen. et sp. nov.

 (illustrated by Lucia Šmídová).

Ya-Ni TANG, Lucia ŠMÍDOVÁ, Ying-Ying DING, Mathieu BODERAU, Dolev FABRIKANT, Bastian MÄHLER, Jun-Chen DENG, Thet Tin NYUNT, Yan-Zh FU, Corentin JOUAULT, Frank TOMASCHEK, Jacek SZWEDO, Hui JIANG. 2026. Cretolala kachinensis gen. et sp. nov. (Hemiptera: Fulgoromorpha: Delphacoidea), the First lalacid from mid-Cretaceous Kachin Amber: Morphological and Taphonomic implications. Palaeoentomology. 9(2); 125-138. DOI: doi.org/10.11646/palaeoentomology.9.2.5 [2026-04-29]

[Arachnida • 2025] Pinelema aurata, P. cavernalis, P. onganh, ... • Hidden in the Dark: Five New Species of Pinelema (Araneae: Telemidae) from Caves in northern Vietnam with Remarks on the Genus Diversity and Conservation

Pinelema spp. in life. 
[A-C] Pinelema onganh, new species;
[D-F] P. cavernalis, new species;
[G-H] P. aurata, new species; [I] P. huifengi, new species 

Ballarin, Nguyen & Eguchi, 2025

RAFFLES BULLETIN OF ZOOLOGY. 73: 

 Abstract 

 Five new cave-dwelling species of the genus Pinelema Wang & Li, 2012 from northern Vietnam are described based on both sexes: Pinelema onganh, new species; P. cavernalis, new species; P. huifengi, new species; and P. nagaoi, new species, from Cao Bang Province, and P. aurata, new species, from Dien Bien Province. Three of these species exhibit marked morphological adaptations to subterranean life, including the absence of eyes, depigmentation, and leg elongation. For each species, detailed images of their diagnostic characters are provided. When possible, barcode sequences of the new species are obtained and compared with published Pinelema sequences in a pairwise analysis to assess the interspecific genetic variation of the genus. The significance of Pinelema species diversity in the Southeast Asian karst regions is also briefly discussed, along with conservation challenges linked to their strict habitat requirements.

 Key words. COI barcode, conservation, diversity, Southeast Asia, subterranean environment

Pinelema spp. in life. 
A, P. onganh, new species, male holotype; B, ditto, adult female; C, ditto, juvenile;
 D, P. cavernalis, new species, male holotype; E, ditto, adult female; F, ditto, adult female with egg sac;
G, P. aurata, new species, adult female; H, ditto, adult female with egg sac; I, P. huifengi, new species, adult female.

Francesco Ballarin, Anh D. Nguyen and Katsuyuki Eguchi. 2025. Hidden in the Dark: Five New Species of Pinelema (Araneae: Telemidae) from Caves in northern Vietnam with remarks on the Genus diversity and conservation. RAFFLES BULLETIN OF ZOOLOGY. 73: 548–567. [17 October 2025]

[Entomology • 2026] Rivetina archibaldi (Mantodea: Rivetinidae) • A New praying mantis from the Zagros Oak Forests of Iran

 Rivetina archibaldi  Kolnegari, 

in Kolnegari, Handal et Khormizi, 2026   

  DOI: doi.org/10.3897/jor.35.174839

Abstract

Iran, the world’s 18th-largest country, harbors a wide range of habitats and a rich diversity of Mantodea. The genus Rivetina Berland & Chopard, 1922 is represented in Iran by several morphologically similar ground-dwelling species. Here, we describe Rivetina archibaldi sp. nov. from the Zagros oak forests of Lorestan Province, western Iran. The new species is diagnosed based on external morphology and male genital structures and is compared with congeners occurring in Iran and Iraq. A dichotomous key to the Iranian species of Rivetina is also provided.

Keywords: Ground-dwelling mantis, Lorestan Province, Middle East, Persian Plateau

Head and pronotum of Rivetina archibaldi sp. nov. male (A, C) and female (B, D).

Live habitus of  Rivetina archibaldi sp. nov. (A, C) male and (B, D) female.

Family Rivetinidae Ehrmann & Roy, 2002

Subfamily Rivetininae Ehrmann & Roy, 2002

Tribe Rivetinini Ehrmann & Roy, 2002

Genus Rivetina Berland & Chopard, 1922

Rivetina archibaldi Kolnegari, sp. nov.

 

Diagnosis.— The metazone minimum width is distinctly smaller in Rivetina archibaldi sp. nov. (mean = 1.78 mm) than in all examined congeners. The new species is morphologically similar to R. dolichoptera but differs in the shape of the pronotum, the shorter supra-anal plate, and distinct features of the male genitalia.

Etymology.— The specific epithet archibaldi is dedicated to Dr. George Archibald, a renowned conservationist who has devoted more than half a century of his life to the protection of wildlife, especially cranes (Aves, Gruidae), around the world. His lifelong efforts have played a central role in saving several crane species from extinction and in promoting the conservation of these legendary and symbolic birds across continents, including in the Middle East and Iran. The name honors his remarkable contribution to avian conservation and his inspiring legacy in global ornithology.

 Mahmood Kolnegari, Elias Handal and Mahdi Zare Khormizi. 2026. Rivetina archibaldi sp. nov. (Mantodea, Rivetinidae), A New praying mantis from the Zagros Oak Forests of Iran. Journal of Orthoptera Research. 35(2): 283-292. DOI: doi.org/10.3897/jor.35.174839 [12 Jun 2026]

[Botany • 2026] Primula chongqingensis (Primulaceae) • A New Species from Chongqing, China

Primula chongqingensis K.Liang & S.R.Yi, 

in Liang, Zhang et Yi, 2026.

DOI: doi.org/10.11646/phytotaxa.763.2.6 

 x.com/Jungle_KL

Abstract

Primula chongqingensis K.Liang & S.R.Yi, sp. nov., a new species of Primula sect. Ranunculoides from Chongqing, China, is described. The new species is currently known only from Qianjiang District, where it grows on wet moss-covered rock faces and dripping cliffs beside streams and below waterfalls in shaded valley woodland. It is most similar to P. ranunculoides in overall morphology, but differs by its smaller leaves with markedly fewer lateral segments, stable long-homostylous flowers, pedicels and calyces covered with short glandular hairs, and a relatively short calyx less than half as long as the corolla tube, usually reaching ca. one third of it. Filiform propagative shoots with apical bulblets are only occasionally produced in P. chongqingensis, whereas similar structures are consistently and abundantly produced in P. ranunculoides. These combined vegetative and reproductive differences support its recognition as a distinct species rather than an infraspecific taxon of P. ranunculoides. A morphological description, distribution information, provisional conservation assessment, a diagnostic comparison with P. ranunculoides, and a key to the species of Primula sect. Ranunculoides are provided.

Eudicots, narrow endemic, homostyly, morphological comparison, Qianjiang District, sect. Ranunculoides, taxonomy, Wuling Mountains

Primula chongqingensis K.Liang & S.R.Yi, sp. nov.

KE (Jungle) LIANG, HONG-JING ZHANG and SI-RONG YI. 2026. Primula chongqingensis (Primulaceae), A New Species from Chongqing, China.  Phytotaxa. 763(2);  193-200. DOI: doi.org/10.11646/phytotaxa.763.2.6 [2026-06-19]

 x.com/Jungle_KL/status/2067732578585460962

[Herpetology • 2026] Amolops minhlei • A New Species of Amolops (Anura: Ranidae) from Vu Quang National Park—An ASEAN Heritage Park of Vietnam

  

Amolops minhlei

A.V. Pham, C. V. Hoang, T. Q. Nguyen, V. T. H. Nguyen, T. T. T. Nguyen, K. D. Nguyen, Thai, N. V. Hoang & C. T. Pham, 2026 

 

 DOI: doi.org/10.11646/zootaxa.5760.2.4  

 facebook.com/PhamAnhdhsphn 

Abstract 

A new species of Amolops is described from Vu Quang National Park, Ha Tinh Province, North-central Vietnam based on morphological and molecular differences. Morphologically, Amolops minhlei sp. nov. is distinguishable from its congeners by a combination of the following morphological characteristics: SVL 31.8–35.8 mm in males, 36.7–45.8 mm in females; head longer than wide; vomerine teeth absent; snout long (SE/SVL 0.16–0.17 in males, 0.16 in females); tympanum distinct, round (TD/ED 0.41–0.42 in males, 0.40–0.43 in females); skin smooth; supratympanic fold indistinct; dorsolateral fold present; webbing formula I0–0II0–1/2III0–1/2IV1/2–0V; dorsum olive green or reddish brown with a few dark spots; a brown dorsolateral stripe extending from behind eye to above vent; dorsal surface of fore- and hind-limbs light grey with brown crossbars, throat, chest and belly white; males with external vocal sac and nuptial pad on finger I; and flanks grey brown with small white glandular spot. In phylogenetic analyses, the new species is strongly supported as a member of the A. monticola group. However, the new species differs from other members of the A. monticola group by 10.00–19.66% pairwise genetic divergence in the mitochondrial ND2 gene fragment and 1.56–6.67% in the mitochondrial 16S gene fragment.

Amphibia, 16S rRNA, Amolops monticola group, Ha Tinh Province, molecular phylogeny, ND2, taxonomy

Amolops minhlei sp. nov.

Ếch bám đá minh

ANH VAN PHAM, CHUNG VAN HOANG, TRUONG QUANG NGUYEN, VAN THI HONG NGUYEN, THUY THU THI NGUYEN, KY DANH NGUYEN, TOAN CANH THAI, NGOC VAN HOANG and CUONG THE PHAM. 2026. A New Species of Amolops (Anura: Ranidae) from Vu Quang National Park—An ASEAN Heritage Park of Vietnam.  Zootaxa. 5760(2); 225-247. DOI: doi.org/10.11646/zootaxa.5760.2.4 [2026-02-23]

 facebook.com/PhamAnhdhsphn/posts/4486775618134704

  

[Fungi • 2026] Pleurocordyceps cornusynnemata • Taxonomy and Phylogeny of Pleurocordyceps (Hypocreales: Polycephalomycetaceae) associated with Ants and Cicadas from Malaysia, including A New Species and New Records

  

Pleurocordyceps cornusynnemata 

in Shahbaz, Mohadden, Rong, Wei et Seelan, 2026.

 DOI: doi.org/10.11646/phytotaxa.750.4.1

 

Abstract

Pleurocordyceps is a group of hyperparasites that infects invertebrates, plants, and other fungi. It belongs to the family Polycephalomycetaceae, a newly established family separated from Ophiocordycipitaceae, order Hypocreales which includes several genera such as Dingleyomyces, Paradingleyomyces, Perennicordyceps, Pleurocordyceps, and Polycephalomyces. Pleurocordyceps currently comprises 26 species reported from South American and Asian regions In Asia, Malaysia has diverse forest ecosystems and rich fungal diversity; however, no previous studies have been reported on the occurrence of Pleurocordyceps species. The aim of the study is to explore the Pleurocordyceps diversity tropical rainforests of Malaysia using morpho-molecular analyses. During a fungal survey in Malaysia, fungicolous fungus associated with ants and cicadas nymphs were collected and subjected to morphological and molecular assessments. Two genetic loci—the internal transcribed spacer (ITS) region and the translation elongation factor 1-alpha (TEF-1α) gene—were used to construct both individual and concatenated phylogenetic trees. The study describes a new species, Pleurocordyceps cornusynnemata sp. nov. (BORH/F03651) from an ant, and two new records of P. aurantiaca (BORH/F03651), and P. nipponica (UTHM-CEF003.1) from Diptera and cicada nymph, respectively. These findings expand the knowledge of the geographic distribution of Pleurocordyceps and provide the first baseline data for the genus in Malaysia, thereby improving understanding of the diversity and evolutionary relationships within Polycephalomycetaceae.

arthropod hosts, entomopathogenic fungi, Pleurocordyceps, synnema, three taxa, typification, Fungi

Pleurocordyceps cornusynnemata sp. nov. 

MUHAMMAD SHAHBAZ, FIRDZA ZULKARNAIN MOHADDEN, JEREMIAH SIA YIAO RONG, YAP JING WEI and JAYA SEELAN SATHIYA SEELAN. 2026. Taxonomy and Phylogeny of Pleurocordyceps (Polycephalomycetaceae, Hypocreales) associated with Ants and Cicadas from Malaysia, including A New Species and New Records.  Phytotaxa. 750(4); 219-237. DOI: doi.org/10.11646/phytotaxa.750.4.1 [2026-04-09]

[Fungi • 2026] Leptobacillium geminatum • A New Species of Leptobacillium and Additional Records of Simplicillium and Leptobacillium (Ascomycota: Cordycipitaceae) from Sabah, Malaysia

Leptobacillium geminatum

in Shahbaz, De Wint, Rong, Wei, Koid, Fayle, Tan, Abdullah, Palaniveloo et Seelan, 2026.

DOI: doi.org/10.1002/nzb2.70107 

Abstract

Simplicillium and Leptobacillium are entomopathogenic fungal genera frequently associated with arthropods in tropical ecosystems and considered promising candidates for biological pest control. However, the diversity of these genera in tropical regions, particularly in Malaysia, has not been systematically investigated in the context of their taxonomy and phylogenetic relationships. This study aimed to investigate the taxonomic diversity and phylogenetic relationship of Simplicillium and Leptobacillium using integrated morphological and molecular analyses. During a survey of entomopathogenic fungi conducted in Malaysia from 2022–2023, parasitized insects were collected, including spiders and wasps infected by entomopathogenic fungi. Phylogenetic analyses were performed using concatenated datasets of the internal transcribed spacer (ITS) and translation elongation factor 1-alpha (TEF1-α) gene to construct phylogenetic trees. The study identified one novel species, Leptobacillium geminatum sp. nov., isolated from a spider. In addition, five species are reported here as new records for Malaysia: S. formicae, L. leptobactrum. symbioticum, and L. coffeanum isolated from spiders and S. puwenense, isolated from a wasp. Morphological characteristics together with molecular phylogenetic evidence supported the taxonomic placement of these taxa. These findings represent the first comprehensive documentation of Simplicillium and Leptobacillium species from Malaysia, significantly expanding the known diversity and geographical distribution of these entomopathogenic fungi in Southeast Asia.

Keywords: concatenation, fungal diversity, morphology, Northern Borneo, taxonomy

 Leptobacillium geminatum sp. nov.

Muhammad Shahbaz, Frederik C. De Wint, Jeremiah Sia Yiao Rong, Yap Jing Wei, Qian-Qun Koid, Tom Fayle, Yee Shin Tan, Sumaiyah Abdullah, Kishneth Palaniveloo, Jaya Seelan Sathiya Seelan. 2026. A New Species of Leptobacillium and Additional Records of Simplicillium and Leptobacillium from Sabah, Malaysia. New Zealand Journal of Botany. 64(2); DOI: doi.org/10.1002/nzb2.70107 [02 June 2026] 

[Botany • 2026] Microlicia almedae (Melastomataceae) • A New Species from the Chapada Diamantina, Bahia, Brazil

Microlicia almedae  R.B.Pacifico & Kriebel,

in Pacifico et Kriebel, 2026. 

DOI: doi.org/10.3897/phytokeys.276.193021

Abstract

A new species, Microlicia almedae, is described and illustrated from the species-rich campo rupestre vegetation of Chapada Diamantina, Bahia, Brazil. The taxonomic novelty is morphologically related to M. giuliettiana and M. pulchra, which are also restricted to Chapada Diamantina. Microlicia almedae differs from the latter two species in leaf shape and size, hypanthium indumentum, calyx lobe shape and size, and petal size. In addition, M. almedae differs from M. giuliettiana by its uniformly magenta petals and from M. pulchra by its capsules with deciduous columellae. Variation in leaf shape among M. almedae, M. pulchra, and M. giuliettiana was quantified using elliptic Fourier analysis, which revealed near-complete separation among the three species.

Key words: Campo rupestre, Lavoisiereae, Pico da Lapa Grande, Serra do Barbado, taxonomy

Microlicia almedae.
 A. Habit; B. Flower; C. Landscape with campo rupestre vegetation at Serra do Barbado, the type locality of M. almedae.

Voucher: L. Daneu et al. 796 (JABU). 

Photos: by L. Daneu (A, B); by E.A. Ramos (C).

Microlicia almedae.
A. Habit; B. Close-up of a branch; C. Leaf adaxial surface; D. Leaf abaxial surface; E. Detail of glandular-punctate adaxial surface; F. Detail of glandular-punctate indumentum on abaxial surface; G. Flower in lateral view; H. Flowering hypanthium and calyx lobes; I. Detail of the indumentum on the hypanthium; J. Petal in adaxial view; K. Antepetalous (left) and antesepalous (right) stamens; L. Gynoecium; M. Apex of style and stigma; N. Ovary in cross-section; O. Capsule enveloped by the hypanthium and calyx lobes; P. Seed in lateral view.

Illustration by Klei Sousa based on L. Daneu et al. 796 (JABU).

  

Microlicia almedae R.B.Pacifico & Kriebel, sp. nov.

Ricardo Pacifico and Ricardo Kriebel. 2026. Microlicia almedae (Melastomataceae), A New Species from the Chapada Diamantina, Bahia, Brazil. PhytoKeys. 276: 183-195. DOI: doi.org/10.3897/phytokeys.276.193021 [12 Jun 2026]

 

[Botany • 2026] Ceropegia gengmaensis (Apocynaceae: Ceropegieae) • A New Species from Yunnan, China

 

Ceropegia gengmaensis  P.R.Luo, A.Liu & H.Sun,

in Luo, Liu, Tong, Zhang, Wang, Deng, Goyder et Sun, 2026. 

耿马吊灯花  ||  DOI: doi.org/10.3897/phytokeys.276.197333

Abstract

Ceropegia gengmaensis P.R.Luo, A.Liu & H.Sun, a new species from Gengma County, Yunnan, Southwest China, is described. Both morphological characteristics and chloroplast phylogenetic analysis strongly support its placement within C. sect. Chionopegia H.Huber. Molecular data further reveal that C. gengmaensis is closely related to C. salicifolia, C. mairei, and C. dolichophylla, yet it can be distinguished from them readily by its stem indumentum, leaf shape, and floral morphology, particularly the shape of the corolla tube, and features of the lobes. A detailed line drawing of this new species is also provided.

Key words: Ceropegieae, Chloroplast, Karst Landform, Morphology, Southwest China

Ceropegia gengmaensis. 
 A, B. Plant showing the rootstock; C, D. Upper and lower sides of leaf; E. Inflorescence with bases of adjoining leaves; F. Corolla tube dissected longitudinally; G. Side view of gynostegium (with sepals and pedicel shown); H. Top view of gynostegium with sepals shown; I. Ovary; J Pollinarium.

 Scale bars: 15 cm (A, B); 5 cm (C, D); 4 cm (E, F); 1 cm (G); 0.8 cm (H); 0.3 cm (I); 0.5 mm (J).

Morphological features of Ceropegia gengmaensis based on living plants collected from the type locality.
 A. Plant showing the rootstock; B, C. Leaf: B, abaxial surface and C, adaxial surface; D. Stem; E, F. Corolla: E, corolla from the outside with its base cut off and F, corolla with the tube dissected longitudinally; G. Small peduncle, part of the pedicel and petiole; H. White hairs on the inside of the base of the tube; I. Side view of gynostegium; J. Gynostegium from above; K. Pollinarium.

Scale bars: 15 cm (A); 5 cm (B, C); 4 cm (D, E, F); 1 cm (G, H, I); 0.8 cm (J); 0.5 mm (K).

Ceropegia gengmaensis P.R.Luo, A.Liu & H.Sun, sp. nov.

 

Diagnosis. Ceropegia gengmaensis shows affinities with C. salicifolia and C. dolichophylla, both of C. sect. Chionopegia . It differs from C. salicifolia by its erect to sprawling habit (vs. twining), elliptic leaves 2.0–5.5 cm (vs. lanceolate, 6–15 cm), densely pubescent stems bearing usually solitary flowers (vs. glabrous stems with sessile cymose many-flowered inflorescences), and narrower corolla lobes with only slight expansion above the throat (vs. lobes longer and wider, upper portion markedly expanded). From C. mairei it differs by the much less pronounced basal inflation of the tube (vs. a strongly inflated tube) and ... 

Etymology. Ceropegia gengmaensis is named after its type locality, Gengma County, Southwest Yunnan.

Vernacular name. the Chinese name is given as “耿马吊灯花” (gěng mă diào dēng huā) named after the type locality, Gengma County, Southwest Yunnan.

Peng-Rui Luo, Ang Liu, Rui Tong, Xin-Jian Zhang, Zi-Yi Wang, Tao Deng, David J. Goyder and Hang Sun. 2026. Ceropegia gengmaensis (Apocynaceae), A New Species from Yunnan, China. PhytoKeys. 276: 217-228. DOI: doi.org/10.3897/phytokeys.276.197333 [15 Jun 2026]

[Botany • 2026] Argostemma baotingense (Rubiaceae) • A New Species from Hainan Island, South China

 

Argostemma baotingense S.P.Dong & H.B.Yang, 

 in Dong, Yuan, Wang et Yang, 2026. 

DOI: doi.org/10.3897/phytokeys.276.195404 

Abstract

Argostemma baotingense, a new species of Rubiaceae from Hainan Island, southern China, is described. This new species resembles A. bachmaense but can be distinguished by its suberect or creeping stem, linear leaves with a greenish-white abaxial surface, white and glabrous inflorescence, and glabrous calyx. An identification key and photographs of the nine Argostemma species in China are provided.

Key words: Argostemma, new species, taxonomy

Argostemma baotingense.
A. Plant in natural habitat; B. Flowering plant; C. Flowers; D. Calyx; E. Leafy branch, adaxial view; F. Leafy branch, abaxial view; G. Leaves, adaxial view; H. Leaves, abaxial view; I, J. A node with opposite leaves and stipules; K. Flower, apical view; L. Flower, bottom view; M, N. Inflorescence; O. Androecium; P. Pistil.

Argostemma baotingense S.P.Dong & H.B.Yang, sp. nov.

Diagnosis. The new species is morphologically similar to A. bachmaense T.V.Do but differs by the suberect or creeping stem (vs. erect), the linear leaves (vs. oblanceolate to spatulate), glabrous inflorescence (vs. pubescent), glabrous calyx (vs. pubescent). A detailed morphological comparison between the two species is provided in Table 1. Photographs of seven Argostemma species in China have been provided (Fig. 2). A key to all the nine species of Argostemma in China is also provided at the end of the text.

Shu-Peng Dong, Lang-Xing Yuan, Qing-Long Wang and Hu-Biao Yang. 2026. Argostemma baotingense (Rubiaceae), A New Species from Hainan Island, South China. PhytoKeys. 276: 253-260. DOI: doi.org/10.3897/phytokeys.276.195404   [18 Jun 2026]

[Botany • 2026] Polystemma cualense (Apocynaceae: Asclepiadoideae) • A New Species from western Mexico

Polystemma cualense  C.D.Sánchez, S.Islas & A.Black, 

in Sánchez-Sánchez, Islas-Hernández et Black, 2026. 

DOI: doi.org/10.3897/phytokeys.276.176723 

Abstract

A new species of Polystemma is described from the Sierra El Cuale, Jalisco, Mexico. Polystemma cualense is distinguished from its congeners, particularly P. horconesense, by a gynostegial corona higher than the gynostegium, longer external corona lobes, and a black gynostegium. Morphological data were obtained by examining herbarium specimens, consulting digital images from iNaturalist, and reviewing specialized literature. A preliminary conservation assessment based on IUCN criteria indicates that it is a Critically Endangered (CR) species due to its restricted area of occupancy and threats from deforestation, agriculture, and livestock grazing. The discovery of P. cualense increases the known diversity of Polystemma in Jalisco and highlights the floristic importance of the Sierra El Cuale.

Key words: Asclepiadoideae, high diversity, Jalisco, Sierra El Cuale

Polystemma cualense.
A. Complete plant; B. Inflorescence; C. Detail of gynostegium; D. Pollinarium. E. Immature follicle with sepal remnants.

Polystemma cualense C.D.Sánchez, S.Islas & A.Black, sp. nov.

  

Diagnosis. Polystemma cualense resembles P. horconesense in having flowers similar in size and shape. Polystemma cualense differs in having a gynostegial corona higher than the gynostegium (vs. gynostegial corona the same height as the gynostegium), external gynostegial corona lobes 3–4 mm length (vs. 0.5–0.6 mm length) and gynostegium black (vs. gynostegium green-yellowish).

 Cristóbal Daniel Sánchez-Sánchez, Carla Sofía Islas-Hernández and Adam W. Black. 2026. Polystemma cualense (Apocynaceae), A New Species from western Mexico. PhytoKeys. 276: 245-251. DOI: doi.org/10.3897/phytokeys.276.176723   [18 Jun 2026]

 

[PaleoIchthyology • 2026] Asioaspis brachyotus • A New Genus of Polybranchiaspiformes (Galeaspida: stem-Gnathostomes) from the Lower Devonian of Yunnan, China

 

Asioaspis brachyotus

 R.-R. Zhang, N. Zhang, Li, Zhu & Gai, 2026

 

VERTEBRATA ALASIATICA. 64(2);   

 

A new genus and species, Asioaspis brachyotus gen. et sp. nov. of Polybranchiaspiformes, is described from the Lower Devonian Lochkovian Xishancun Formation near Miandian Reservoir, Qujing City, Yunnan Province, China. The new genus is characterized by an elongated ovoid headshield with serrated lateral margins, a small subcircular median dorsal opening, short and laterally projecting cornual processes, an exceptionally broad ventral rim, 12 pairs of branchial fossae, and coarse granular tubercles (~3/mm²). It displays a mosaic of characters of the most primitive polybranchiaspiform (Platylomaspis), derived Polybranchiaspiformes, and even the basal members of the galeaspid family Dayongaspidae. Phylogenetic analysis resolves A. brachyotus as an intermediate taxon between the most primitive Gumuaspidae and other derived Polybranchiaspiformes. Although A. brachyotus also shares the broad ham-brim-like ventral rim with the Gumuaspidae, its other characters are more like other derived Polybranchiaspiformes. In particular, its median dorsal opening and paired orbital openings are not highly close-set on the top of the headshield as in the Gumuaspidae, but rather significantly separated from each other and close to the margin of the ventral rim as in the Pentathyraspidae, Duyunolepididae, and Polybranchiaspidae. These proportions indicate that it was unlikely to have lived a semi-infaunal lifestyle and are more consistent with the epibenthic habit typical of most galeaspids, representing an evolutionary transition from the semi-infaunal benthic to epibenthic lifestyle within Polybranchiaspiformes.

Photograph of Asioaspis brachyotus gen. et sp. nov. from Lower Devonian Xishancun Formation, Qujing, Yunnan in ventral view
A. an incomplete headshield, paratype, IVPP V26676. 2; B. a nearly complete dorsal headshield, holotype, V26676. 1
A. an incomplete dorsal headshield, IVPP V26676.5; B. an incomplete dorsal headshield, V26676.4b

Subclass Galeaspida Tarlo, 1967 

Supraorder Polybranchiaspidida Janvier, 1996 

Order Polybranchiaspiformes Liu, 1965 

Family Asioaspidae fam. nov. 

Type genus Asioaspis gen. nov. 

Differential diagnosis: The Asioaspidae is distinguished from other families of Polybranchiaspiformes by its short, ear-like, and laterally projecting cornual processes. It resembles the Gumuaspidae in having a broad ventral rim, but differs in lacking a rostral process. It differs from the Duyunolepididae, Pentathyraspidae, and Polybranchiaspidae by the presence of a broad ventral rim and further differs from the Duyunolepididae by bearing a pair of broad inner cornual processes. Therefore, a new family, the Asioaspidae fam. nov., is herein established.

Genus Asioaspis gen. nov.  

Type species Asioaspis brachyotus gen. et sp. nov. 

Etymology: Asioaspis is derived from Asio (referring to owls of the genus Asio characterized by prominent ear tufts) plus aspis (Greek shield). 

Diagnosis: Medium-sized polybranchiaspiform characterized by an ovoid headshield with serrated lateral margins; the rostral margin arcuate, but lacking a rostral process; the cornual processes short and projecting laterally; the inner cornual processes large and broad leaf-shaped; median dorsal opening small and subcircular in shape; orbital opening large, round, and dorsally positioned; the sensory canals are well developed, V-shaped posterior supraorbitals and four pairs of lateral transverse canals issuing from the lateral dorsal canal; ventral rim of headshield remarkably broad, 13.5–14.1 mm in width on each side (approximately one half of the width of the entire headshield); a large dome-shaped oralobranchial chamber (a composite chamber with both oral and branchial functions) with 12 pairs of branchial fossae; and ornamentation consists of coarse tubercles (~3/mm²)

 Asioaspis brachyotus gen. et sp. nov. 

Etymology: The brachyotus means short-eared, which is in reference to its short and laterally projecting cornual processes that resemble a pair of short ears on the headshield, as in Asio.

Type locality and horizon: Miandian Reservoir in the Triathlon Sports Park, Qilin District, Qujing City, Yunnan Province, China; Xishancun Formation, lower Lochkovian, Lower Devonian.

Morphological comparison of Polybranchiaspiformes, Dayongaspidida, and Huananaspidiformes

A. Asioaspis; B. Dayongaspis (Zhang et al., 2024); C. Foxaspis (Gai et al., 2023); D. Platylomaspis (redrawn after Gai et al., 2018); E. Rhegmaspis (Gai et al., 2015); F. Rhegmaspis (Gai et al., 2015); G. Nanningaspis (redrawn after Gai et al., 2018)

A–E, G. in dorsal views; F. in ventral view.  

Phylogenetic diagram of Dayongaspidida, Polybranchiaspiformes, and Huananaspidiformes
Dayongaspis (Zhang et al., 2024); Platylomaspis (redrawn after Gai et al., 2018); Nanningaspis (redrawn after Gai et al., 2018); Polybranchiaspis (Liu, 1965); Foxaspis (Gai et al., 2023); Rhegmaspis (Gai et al., 2015).

No scaled

 Life restoration of Asioaspis brachyotus gen. et sp. nov. 

ZHANG Rui-Rui, ZHANG Ning, LI Qiang, ZHU Min and GAI Zhi-Kun. 2026. Asioaspis, A New Genus of Polybranchiaspiformes (Galeaspida, stem-Gnathostomes) from the Lower Devonian of Yunnan, China.  VERTEBRATA PALASIATICA. 64(2); 137-150.  DOI: 10.19615/j.cnki.2096-9899.260113 [20 April 2026] 

https://www.vertpala.ac.cn/EN/10.19615/j.cnki.2096-9899.260113

[Entomology • 2026] Leptogenys distincta & L. gastrolucida • Two New Species of Ant Genus Leptogenys (Hymenoptera: Formicidae) from India, with an updated key to the known Oriental species

Leptogenys distincta  
 Leptogenys gastrolucida

Baidwan, Kapoor & Bharti, 2026

 

DOI: doi.org/10.1080/00222933.2026.2657496

 

ABSTRACT

Two new species of the genus Leptogenys are described based on the worker caste, namely Leptogenys distincta sp. n. and Leptogenys gastrolucida sp. n. from India. The male of L. gastrolucida sp. n. is also reported and described. An updated dichotomous key to the 60 known species of Leptogenys from the Oriental region is provided.

 

KEYWORDS: Taxonomy, Indo-Burma, biodiversity, description

 

 Leptogenys distincta sp. n.  

Leptogenys gastrolucida sp. n.  

Parvinder Singh Baidwan, Rakeshwar Kapoor and Himender Bharti. 2026. Two New Species of Ant Genus Leptogenys (Hymenoptera: Formicidae) from India, with an updated key to the known Oriental species. Journal of Natural History. 60(25-28); 1357-1397. DOI: doi.org/10.1080/00222933.2026.2657496 [08 Jun 2026]