Tuesday, November 19, 2019

[Herpetology • 2019] Leptobrachella namdongensis • A New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Thanh Hoa Province, Vietnam


Leptobrachella namdongensis 
Chung, Nguyen, Luu, Nguyen & Jiang, 2019

RAFFLES BULLETIN OF ZOOLOGY. 67

Abstract
 Based on the results of molecular and morphological systematics analyses, a new species, Leptobrachella namdongensis, is recognised from Nam Dong Conservation Area in Thanh Hoa Province, Vietnam. Genetic divergences in the16S rRNA gene between the new form and other species in the genus range from 4.7 to 21.0% and average at approximately 24%. The new species is most similar to L. petrops and L. puhoatensis, but can be distinguished from them and other congeners by a combination of the following morphological characters: size medium (30.9 mm in the adult male and 32.1–35.3 mm in adult females); head wider than long; tympanum distinct; throat immaculate white, chest and belly with dark specking on outer margins; toes with basal webbing; supratympanic fold edged by a distinct black line; inter-orbital region with distinct black blotches; iris bicoloured, upper half copper, fading to golden in its lower half; head without dorsolateral markings. 

Key words: Nam Dong Conservation Area, phylogenetics, Vietnam, taxonomy


Fig. 3. Dorsolateral view (A) and ventral view (B) of the holotype (VNUF A.2017.37, male) of Leptobrachella namdongensis, new species in life. 
Photos by Vinh Q. Luu.

Fig. 6. Paratypes of Leptobrachella namdongensis, new species in life.
 A, female VNUF A.2017.95 (SVL 32.1mm); B, female VNMN2019.04 (SLV 33.0mm); C, female IEBR 4512 (SVL 32.2mm).
 Photos by Vinh Q. Luu.

Leptobrachella namdongensis, new species

Diagnosis. Leptobrachella namdongensis, new species is distinguished from its congeners by a combination of the following morphological characters: size medium (30.9 mm in the adult male and 32.1–35.3 mm in adult females); head wider than long; tympanum distinct; throat immaculate white, chest and belly with dark specking on outer margins; toes with basal webbing and without lateral fringe; supratympanic fold edged by a distinct black line; inter-orbital region with distinct black blotches; iris bicoloured, upper half copper, fading to golden in its lower half; head without dorsolateral markings.

Etymology. Specific epithet is in reference to the type locality of the Nam Dong Conservation Area. We recommend “Nam Dong litter frog” as the common English name and “Cóc mày nam động” as the Vietnamese name.

Distribution and ecological notes. Leptobrachella namdongensis, new species is currently known from Nam Dong Conservation Area and Pu Hu Nature Reserve (NR), Thanh Hoa Province, Vietnam. All specimens of the new species were found on rocks in small streams or near cascade streams in evergreen forest at elevations between 500 and 740 m a.s.l. (Fig. 7). Leptobrachella namdongensis, new species occurs sympatrically with Leptobrachella ventripunctata.

Fig. 7. Habitat of Leptobrachella namdongensis, new species in Nam Dong Conservation Area, Thanh Hoa Province, Vietnam.



Hoang Van Chung, Nguyen Thien Tao, Luu Quang Vinh, Nguyen Quang Truong and Jiang Jianping. 2019. A New Species of Leptobrachella Smith 1925 (Anura: Megophryidae) from Thanh Hoa Province, Vietnam. RAFFLES BULLETIN OF ZOOLOGY67;  536–556.  

     


10 comments:

  1. Do you want to fix the login and sign up errors in Binance and enjoy the services hassle-free? If you are unaware about the process, all you can do is dial Binance helpline number and acquire quality solutions from the team of professionals who is always there to guide you. These common issues can be fixed the moment you connect with the professionals as they will not only deliver the easiest solutions but also guide the whole process that can be easily executed and can be remembered for the long run.
    https://www.cryptowalletsupport.com/binance-support-number/

    ReplyDelete
    Replies
    1. Natural herbs have cured so many illness that drugs and injection cant cure. I've seen the great importance of natural herbs and the wonderful work they have done in people's lives. i read people's testimonies online on how they were cured of herpes, HIV, diabetics etc by Dr Ojoka herbal medicine, so i decided to contact the doctor because i know nature has the power to heal anything. I was diagnosed with HIV for the past 7 years but Dr Ojoka cured me with his herbs and i referred my aunt and her husband to him immediately because they were both suffering from herpes but to God be the glory, they were cured too .I know is hard to believe but am a living testimony. There is no harm trying herbs. He is also a spell caster, he cast spell to restore broken marriages and he cast good luck spells to prosper and excel dr.ojokarootandherbal@gmail com life. Contact Email  or call Whats App him +2348144172934



































































































      Natural herbs have cured so many illness that drugs and injection cant cure. I've seen the great importance of natural herbs and the wonderful work they have done in people's lives. i read people's testimonies online on how they were cured of herpes, HIV, diabetics etc by Dr Ojoka herbal medicine, so i decided to contact the doctor because i know nature has the power to heal anything. I was diagnosed with HIV for the past 7 years but Dr Ojoka cured me with his herbs and i referred my aunt and her husband to him immediately because they were both suffering from herpes but to God be the glory, they were cured too .I know is hard to believe but am a living testimony. There is no harm trying herbs. He is also a spell caster, he cast spell to restore broken marriages and he cast good luck spells to prosper and excel dr.ojokarootandherbal@gmail com life. Contact Email  or call Whats App him +2348144172934

      Delete
  2. Email address in gemini can solve your numerous issues, so, is your email address verified in gemini? Verifying email address and linking it with the gemini account can fix many issues easily. If you have forgotten your password and need to fix it, you can send the email to your email ID and your issue will be solved in short. The team of skilled professionals provides solutions that are easy to implement and can be memorized for the long run. Dial gemini customer care number and get your issues resolved in the minimal time with perfection.
    https://www.cryptophonesupport.com/exchange/gemini/

    ReplyDelete
  3. Are you fed up of fixing email activation issues in trezor account? Do you want to fix your email activation error in trezor? Is your email not active in trezor and you are unable to get the complete information about the activity going on in your account? Well, you need to be extra careful to deal with such issues as your one wrong step can create issues. Dial trezor helpdesk number which is always available and is the easiest source of forwarding the errors to the professional team immediately. You can always connect with the team anytime for availing quality solutions from the team in no time.
    https://www.cryptowalletsupport.com/trezor-support-number/

    ReplyDelete
  4. i got hpv from my boyfriend and i was trying all i could do to get rid of this virus over 4 months now i have taking a lot of drugs from different medical doctor which didn't work at all, last month i was doing some research on google on how to get cured totally from this virus i found someone testimony on how she got cured totally from hpv with the help of doc. onokun natural herbs and she drop doc onokun email address, So i decide to give him a try and i contact him for help he told how to purchase the natural herbs and i did that, Do you believe i only took this herbs for 2 weeks and i got cured permanently from the hpv virus without no side effect. Thank you so much doc onokun for the help i will make sure i tell the whole world about you. contact him if you need his help too via: Dronokunherbalcure@gmail.com

    ReplyDelete
  5. Many people are making profits weekly and many are withdrawing their profits daily, weekly and even monthly. What are you waiting for? Contact Mrs Mary today. Send a message via gmail,,, (maryshea03@gmail.com) WhatsApp +15623847738 .Remember this is absolutely free!!!

    ReplyDelete
  6. Contact Dr aluya  at http://Draluyaspelltempl.website2.me if you want to get your Husband, Wife, Boyfriend and Girlfriend back now and regain your happiness.

    My Dear friends online, My name is walden Ann

      And i live in USA, New York, I have to give this miraculous testimony, which is so unbelievable until now. I had a problem with my Ex husband 2 years ago, which lead to our break up. I was not myself again, i felt so empty inside me, my love and financial situation became worst, until a close friend of mine told me about a spell caster who helped her in the same problem too his name is Doctor Aluy I email the spell caster and i told him my problem and i did what he asked of me, to cut the long story short. Before i knew what was happen, less than two days my husband gave me a call and told me that he was coming back to me i was so happy to have him back to me. The most interesting part of the story is that am pregnant. Thanks to Doctor Aluya for saving my marriage and for also saving others own too. Continue your good work, If you are interested to contact the great spell caster email address: aluya.48hoursspelltemple@gmail.com Good Luck

    ReplyDelete
  7. herpes is a serious and recurring disease which can't be cured through drugs or injections by the American doctors but the best way to deal with herpes is by taking natural herbs medicine for it and is only few American doctors that know about this herbal medicine from Dr Ofure.. I have read about Dr Ofure the great herbalist doctor from African who can cure disease with his powerful herbal medicine. for the people suffering from the following diseases, Herpes, Cancer, Also,Herpatitis, Diabetes, Hps,Infections ETC should contact him for his herbal medicine because i am a living testimony and i was cured of herpes. Although, i sent him what he requested and he sent me his medicine which i took for 1 weeks and today when i went for test i was tested herpes negative. you can reach him through his email drofureherbalhome@gmail.com or whatsapp number:+2348162868851 he will help you out

    ReplyDelete
  8. Hello everyone , here’s your opportunity for you to achieve your dreams of being a multi million dollar rich through trading , I once loss all I got through trading but was fortunate to come across a woman with great virtue and selfless heart (Mary ) i was introduce to her masterclass strategy while searching online which has revived me of all my losses and made me gain more and more . With her unique strategy you are entitled to daily signals and instant withdraw ,be rest assured of getting a refund of all your loss investment with any platform that has denied you in one way or the other in getting your money . Mrs Mary masterclass strategy is simply the best for beginners and those that are finding it difficult to succeed through trading she’ll help you with just a simple step . Email her ( maryshea03 @ Gmail .com) WhatsApp +1 562 384 7738 . Remember this is absolutely free!!!

    ReplyDelete
  9. This is the website I read about Doctor Life and contacted him to help me get my ex girlfriend back to marry me during the month of May this year and I have come back here to post about him. Doctor Life will help you solve your problem no matter what you are going through. My name is sebi from USA. Posting about a spell caster is very strange to me because I never believed i will be able to say that I was helped by a spell caster in bringing my ex girlfriend back to me after 3 years of no contact. I read about Doctor Life from other websites and contacted him to help me and in less than 48 hours my ex girlfriend called me and I was happy that he wants to get back to me. We met on Friday and he proposed to me,… It was the most beautiful ring. Please everyone out here, Contact Doctor Life to solve your problem for you and make you happy with your relationship that is hurting you. Love is the best feeling ever experienced. Email him at drlifetemple@gmail.com or directly on whats App him +2347010538590 or him website https://lifetempleforhahhpiness.webs.com

    ReplyDelete