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[18, 19, 7] ผีเสื้อขมิ้นดิ้นเงิน Cimeliomorpha jarujini,
[21] C. perspinosa, [22-23] C. inflata
Pinkaew & Horak, 2019
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Abstract
The enarmoniine genus Cimeliomorpha Diakonoff, 1966 is reviewed. Seven species are treated, and a key to the species is provided. Three named species, C. cymbalora (Meyrick), C. novarana (Felder & Rogenhofer), and C. egregiana (Felder & Rogenhofer), are redescribed, with the identity of C. novarana finally settled. Three new species, Cimeliomorpha jarujini, sp. n.; C. inflata, sp. n.; and C. perspinosa, sp. n., are described. Cimeliomorpha nabokovi Kuznetsov is included based on the original description. Illustrations of adults and genitalia of all species are provided, and the original hand-coloured images of C. egregiana and C. novarana are reproduced from a copy of the relevant book with unusually well-preserved plates.
Keywords: Lepidoptera, new species, Enarmoniini, Olethreutinae, Thailand, New Guinea
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FIGURES 4−9. Labial palpi of Cimeliomorpha spp.
4. C. cymbalora, female (Thailand, KKIC, np2895). 5. C. novarana, female (Thailand, KKIC, np5789). 6. C. egregiana, male (Amboyna, Doherty, 1892 No.41343, NHMUK). 7. Cimeliomorpha jarujini, male (Thailand, holotype). 8. C. perspinosa, male (West Papua, holotype). 9. C. inflata, male (Woodlark Id, holotype). |
Cimeliomorpha Diakonoff, 1966
Cimeliomorpha Diakonoff, 1966: 50, fig. 1. Obraztsov, 1968: 185, figs 9‒13; Robinson et al., 1994: 104, plate 17 fig. 5;
Kuznetsov, 1997: 801, fig. 7; Brown et al., 2005: 177.
Type species: Copromorpha cymbalora Meyrick, 1907, by original designation.
Diagnosis. Cimeliomorpha has an unmistakable two-toned forewing pattern with a uniformly yellow or white basal
half and a distal half with complex mostly black pattern on red-brown ground with raised silvery lines and spots,
and a hindwing that is either white or orange at the base, with a narrow to wide blackish band around its margin.
Anthozela Meyrick and some Loboschiza species also have a conspicuously two-toned forewing, but in the former
the basal half is yellow and either speckled with black or suffused with orange, and in the latter the hindwing is not
white or yellowish in its basal half. The genitalia of both sexes of Cimeliomorpha also indicate a close relationship
with Loboschiza, but not with Anthozela. The wing venation with widely separated Rs
and M1
and parallel M2
and
M3
in the hindwing (Fig. 3) is most similar to that of Irianassa Meyrick, but this is due rather to symplesiomorphy
than to a close relationship as evidenced by the very different genitalia in both sexes.
The cymbalora-group. Forewing length 5.8−7.2 mm. Frons, vertex, labial palpus, pronotal collar, tegulae and
mesonotum white. Forewing with basal 2/5 white; hindwing with basal half white; male genitalia with membrane
next to juxta without setae, cucullus tip pointed and ending in short strong spine; female genitalia with one small
signum, a granulate sclerite.
The egregiana-group. Forewing length 7.2−9.9 mm. Frons, vertex, labial palpus, pronotal collar, tegulae and mesonotum yellow. Forewing with basal 2/5 yellow; hindwing with basal part orange or at least with large orange patch in
centre; male genitalia with membrane next to juxta with numerous short setae, cucullus distally rounded and not ending
in a single long spine (C. jarujini with one strong spine centrally); female genitalia with two horn-shaped signa.
Cimeliomorpha cymbalora (Meyrick, 1907)
Distribution. India, Thailand (new record). Specimens from Thailand were collected from evergreen forest.
Cimeliomorpha novarana (Felder & Rogenhofer, 1875)
Distribution. Nicobar Island, Thailand (new record), Malaysia, Indonesia. Specimens from Thailand were collected from evergreen and dry evergreen forest.
Cimeliomorpha nabokovi Kuznetsov, 1997
Distribution. Vietnam.
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FIGURES 16−23. Adults of egregiana group (scale bars = 2 mm).
16. C. egregiana, male (Amboyna, Doherty, 1892 No.41343, NHMUK). 17. C. egregiana, female (Amboyna, Doherty, 1892 No.41344, NHMUK). 18. Cimeliomorpha jarujini, male holotype (Thailand). 19. C. jarujini, female paratype (Thailand, KKIC, np1852). 20. C. perspinosa, male holotype (West Papua). 21. C. perspinosa, female paratype (Papua New Guinea, NHMUK). 22. C. inflata, male holotype (Woodlark Id). 23. C. inflata, female paratype (Woodlark Id, ANIC). |
Cimeliomorpha egregiana (Felder & Rogenhofer, 1875)
Distribution. Ambon Island (Maluku Islands), Indonesia.
Cimeliomorpha jarujini, sp. n.
Diagnosis. This species differs from members of the cymbalora group by the yellowish basal 2/5 of the forewing and a mostly orange hindwing. It is similar to all other species of the egregiana group in forewing pattern, but in the hindwing the short crescent-shaped blackish band is diagnostic: it reaches neither the base of the costa nor the anal angle, or at most only as some scattered scales, unlike all other species of the egregiana group. The valva of C. jarujini is most similar to that of C. perspinosa, but the cucullus of the former is less densely setose and has a central single large spine.
Etymology. This species is named after the late Jarujin Nabhitabhata, the first director of The Thailand Natural History Museum.
Distribution. Southern Thailand (Nakhon Si Thammarat). Specimens were collected from evergreen forest.
Cimeliomorpha perspinosa, sp. n.
Diagnosis. This species differs from members of the cymbalora group by the yellow basal 2/5 of the forewing and a mostly orange hindwing. It is similar to the other species of the egregiana group in forewing pattern. The hindwing above is similar to that of C. egregiana with the black band reaching to the anal angle but not beyond, while in C. jarujini it ends before the anal angle, and in C. perspinosa it extends also along the anal margin. A diagnostic difference on the ventral side of the hindwing separates C. perspinosa with the subelliptical patch on costa medially from C. egregiana with a longitudinal blackish patch on costa. The valva of C. perspinosa is most similar to that of C. jarujini, but the cucullus of C. perspinosa has a dense tuft of equally large spines, whereas C. jarujini has a single, much larger spine.
Etymology. The specific epithet refers to the cluster of bristles on the cucullus.
Distribution. New Guinea: West Papua (Indonesia) and New Britain (Papua New Guinea).
Cimeliomorpha inflata, sp. n.
Diagnosis. This species differs from members of the cymbalora group by the yellow basal 2/5 of the forewing and the partially orange hindwing. From other species in the egregiana group it differs by the very distinct but short, black, longitudinal fine striation in the ocelloid patch, and by the orange scales in the hindwing restricted to a bipartite median patch. The valva of C. inflata is unique, with a large and swollen cucullus, and the female genitalia are diagnostic with two elliptical patches of scale sockets on the membranous lamella postvaginalis and a very long colliculum.
Etymology. The specific epithet refers to the swollen appearance of the cucullus (inflata=swollen).
Distribution. Papua New Guinea (St. Matthias Island near New Britain, and Woodlark, Goodenough and Bougainville islands) and Solomon Islands (Guadalcanal and The Nggela Islands).
Nantasak Pinkaew and Marianne Horak. 2019. Revision of the Genus
Cimeliomorpha Diakonoff (Lepidoptera: Tortricidae).
Zootaxa. 4615(3); 457–480. DOI:
10.11646/zootaxa.4615.3.3
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