Sunday, April 26, 2015

[Herpetology • 2015] Gegeneophis tejaswini • A New Species of Indian Caecilian Highlights Challenges for Species Delimitation within Gegeneophis Peters, 1879 (Gymnophiona: Indotyphlidae)

Gegeneophis tejaswini 
Kotharambath, Wilkinson, Oommen & Gower, 2015


A new species of indotyphlid caecilian amphibian, Gegeneophis tejaswini sp. nov., is described based on eight specimens from lowlands of the most northerly district of the state of Kerala in the southern part of the Western Ghats region, India. This species is distinguished from all other Gegeneophis in annulation characters and genetics (> 6% different from most similar nominal species for 883 base pairs of mitochondrial 12S and 16S rRNA gene sequence data). The high degree of morphological similarity of G. krishni, G. mhadeiensis and the new species underlines that, for some Gegeneophis, larger samples and/or new characters will be needed to further advance the taxonomy of this genus.

Keywords: caecilians, herpetology, India, taxonomy, Western Ghats

Kotharambath, Ramachandran, Mark Wilkinson, V. Oommen & David J. Gower. 2015. A New Species of Indian Caecilian Highlights Challenges for Species Delimitation within Gegeneophis Peters, 1879 (Amphibia: Gymnophiona: Indotyphlidae).
Zootaxa. 3948(1): 60–70. DOI: 10.11646/zootaxa.3948.1.4

[Herpetology • 2015] A Phylogeny of the Only Ground-dwelling Radiation of Cyrtodactylus (Squamata, Gekkonidae): Diversification of Geckoella across peninsular India and Sri Lanka

Fig. 1. Maximum likelihood phylogeny (ND2, RAG1, PDC) of Geckoella, with representative photographs of Geckoella species and a map of peninsular India and Sri Lanka showing sample localities (referenced in Table 1). Outgroups not shown (see Fig. S1 for complete tree); node support indicated by circles, solid fill = high support (ML bootstrap P75, Bayesian PP P 0.95), light fill = support only from Bayesian analyses (Bayesian PP P 0.95, ML bootstrap <75). Sample localities (number) follow the taxa labels.
Photographs of each species complex are approximately scaled by their maximum snout-vent length (Smith, 1935; Bauer and Giri, 2004; Somaweera and Somaweera, 2009). Colored branches and locality labels on the map indicate dry zone and wet zone clade membership. | DOI: 10.1016/j.ympev.2014.09.016

The subgenus Geckoella, the only ground-dwelling radiation within Cyrtodactylus, closely overlaps in distribution with brookii group Hemidactylus in peninsular India and Sri Lanka. Both groups have Oligocene origins, the latter with over thrice as many described species. The striking difference in species richness led us to believe that Geckoella diversity is underestimated, and we sampled for Geckoella across peninsular India. A multi-locus phylogeny reveals Geckoella diversity is hugely underestimated, with at least seven undescribed species, doubling previously known richness. Strikingly, the new species correspond to cryptic lineages within described Indian species (complexes); a number of these endemic lineages from the hills of peninsular India outside the Western Ghats, highlighting the undocumented diversity of the Indian dry zone. The Geckoella phylogeny demonstrates deep splits between the Indian species and Sri Lankan G. triedrus, and between Indian dry and wet zone clades, dating back to the late Oligocene. Geckoella and brookii group Hemidactylus show contrasting diversification patterns. Geckoella shows signals of niche conservatism and appears to have retained its ancestral forest habitat. The late Miocene burst in speciation in Geckoella may be linked to the expansion of rain forests during the mid-Miocene climatic optimum and subsequent fragmentation with increasing late Miocene aridification.

Keywords: Mid-Miocene climatic optimum; Aridification; Cryptic species; Historical biogeography

• Species diversity vastly underestimated – 7 potentially new Geckoella from hills of peninsular India.
• Geckoella and brookii group Hemidactylus show contrasting historical diversification in peninsular India and Sri Lanka.
• Signals of niche conservatism in Geckoella, retention of terrestrial habit and forest habitat.
• Late Miocene diversification may be linked to mid-Miocene forest expansion and late Miocene aridification.

Ishan Agarwal and K. Praveen Karanth. 2015. A Phylogeny of the Only Ground-dwelling Radiation of Cyrtodactylus (Squamata, Gekkonidae): Diversification of Geckoella across peninsular India and Sri Lanka. Molecular Phylogenetics and Evolution. 82(A); 193–199. DOI: 10.1016/j.ympev.2014.09.016

Friday, April 24, 2015

[Paleontology • 2015] Gorgetosuchus pekinensis • A New Aetosaur (Archosauria, Suchia) from the Upper Triassic Pekin Formation, Deep River Basin, North Carolina, U.S.A., and its Implications for early Aetosaur Evolution

Aetosaurs are an extinct clade of quadrupedal, heavily armored archosaurs that had a worldwide distribution during the Late Triassic. Aetosaur fossils from the Upper Triassic Pekin Formation in the Deep River Basin of North Carolina (U.S.A.) consist primarily of isolated osteoderms and, rarely, more associated material. Here we describe a new genus and species, Gorgetosuchus pekinensis, based on an associated incomplete anterior carapace, consisting of a total of 19 nearly complete paramedian and lateral osteoderms from the first 10 rows of armor as well as some associated fragments. An important feature of Gorgetosuchus is that an articulated fifth row of cervical osteoderms almost encloses the neck, with prominent spines on both the dorsal and lateral osteoderms. This is a novel configuration among aetosaurs. Otherwise, NCSM 21723 preserves a mosaic of character states found in Longosuchus, Lucasuchus, or both taxa while simultaneously preserving several more plesiomorphic character states, such as cervical osteoderms that are wider than long. Our reevaluation of other Pekin Formation specimens that various authors have assigned to Desmatosuchus, Longosuchus, or Lucasuchus confirms that some possess characteristics of Lucasuchus, whereas others are not generically determinate. Incorporating Gorgetosuchus into existing phylogenies of aetosaurs results in a reshuffling of basal aetosaur relationships, but a variety of analyses consistently place Gorgetosuchus as a basal desmatosuchine. Using current taxonomic practices, there are at least three aetosaur genera in the Pekin Formation: Lucasuchus, Coahomasuchus, and Gorgetosuchus

Andrew B. Heckert, Vincent P. Schneider, Nicholas C Fraser and Richard A. Webb. 2015. A New Aetosaur (Archosauria, Suchia) from the Upper Triassic Pekin Formation, Deep River Basin, North Carolina, U.S.A., and its Implications for early Aetosaur Evolution. Journal of Vertebrate Paleontology. 35(1). DOI: 10.1080/02724634.2014.881831

[Paleontology • 2015] Wangosaurus brevirostris • A New Pistosauroid (Reptilia, Sauropterygia) from the late Ladinian Xingyi Marine Reptile Level, southwestern China

Wangosaurus brevirostris
Ma, Jiang, Rieppel, Motani & Tintori, 2015


Sauropterygia is a diverse group of Mesozoic marine reptiles that includes the Plesiosauria, one of the most successful clades from the Jurassic to the Cretaceous. The body plan of the Plesiosauria differs from that of basal sauropterygians by adaptations for life in the open ocean, as expressed by the propulsive, paddle-like limbs (Storrs, 1993; Sato et al., 2010; Benson et al., 2012). Triassic pistosauroids are believed to be important in the search for the origin of Plesiosauria (Rieppel et al., 2002; Sato et al., 2010). Until recently, the sister-group relationship between them was mainly based on cranial information, because of the generally poor preservation of Triassic pistosauroid specimens. Eight genera have been reported so far (Meyer, 1839; Yang, 1959, 1965; Storrs, 1991; Rieppel, 1997, 1998, 1999; Sander et al., 1997; Cheng et al., 2006; Dalla Vecchia, 2006), but some of these are known only from relatively complete cranial material (Augustasaurus Sander et al., 1997; Corosaurus Case, 1936; Cymatosaurus Fritsch, 1894; Pistosaurus Meyer, 1839; Yunguisaurus Cheng et al., 2006), whereas Yunguisaurus remains to date the only taxon represented by a complete skeleton.

Three genera of pistosauroids have been reported from the Triassic marine reptile faunas of southwestern China. Chinchenia sungi Young, 1965, from the upper Middle Triassic of Qingzhen, Guizhou Province, and Kwangsisaurus orientalis Young, 1959, from the upper Lower or lower Middle Triassic of Wuming, Guangxi Province, are both extremely fragmentary and listed as Pistosauroidea incertae sedis (Yang, 1959, 1965; Rieppel, 1999, 2000). Yunguisaurus liae Cheng, Sato, Wu, and Li, 2006, from the late Middle Triassic of Chajiang, Guizhou Province, and Fuyuan, Yunnan Province, is the only one represented by a complete skeleton, which is suggestive of early stages of adaptation towards the plesiosaurian functional style (Cheng et al., 2006; Zhao et al., 2008; Sato et al., 2010, 2014). Thus, more complete material should be described to provide new information that would better resolve the origin of Plesiosauria.

Here, we report a new pistosauroid, Wangosaurus brevirostris, gen. et sp. nov., from the late Middle Triassic of Xingyi, Guizhou Province, which shows a combination of plesiomorphic and derived characters (Fig. 1). Due to the rarity of complete and articulated skeletons of pistosauroids, and the presence of enough unique characters gleaned from the dorsal view of this specimen to establish a new genus, we here provide a preliminary description and phylogenetic analysis.

Le-Tian Ma, Da-Yong Jiang, Olivier Rieppel, Ryosuke Motani and Andrea Tintori. 2015. A New Pistosauroid (Reptilia, Sauropterygia) from the late Ladinian Xingyi Marine Reptile Level, southwestern China. Journal of Vertebrate Paleontology. 35(1):1-6.
DOI: 10.1080/02724634.2014.881832

[Paleontology • 2015] Evidence for Sexual Dimorphism in the Plated Dinosaur Stegosaurus mjosi (Ornithischia, Stegosauria) from the Morrison Formation (Upper Jurassic) of Western USA

Fig 4. Hypothetical silhouettes of male and female Stegosaurus mjosi.
The wide morph exhibits more overlap between adjacent plates than does the tall morph, leading to a more continuous display surface. Sexual dimorphism in the size and shape and plates might have also occurred with other sexual differences such as sexual dichromatism.


Conclusive evidence for sexual dimorphism in non-avian dinosaurs has been elusive. Here it is shown that dimorphism in the shape of the dermal plates of Stegosaurus mjosi (Upper Jurassic, western USA) does not result from non-sex-related individual, interspecific, or ontogenetic variation and is most likely a sexually dimorphic feature. One morph possessed wide, oval plates 45% larger in surface area than the tall, narrow plates of the other morph. Intermediate morphologies are lacking as principal component analysis supports marked size- and shape-based dimorphism. In contrast, many non-sex-related individual variations are expected to show intermediate morphologies. Taphonomy of a new quarry in Montana (JRDI 5ES Quarry) shows that at least five individuals were buried in a single horizon and were not brought together by water or scavenger transportation. This new site demonstrates co-existence, and possibly suggests sociality, between two morphs that only show dimorphism in their plates. Without evidence for niche partitioning, it is unlikely that the two morphs represent different species. Histology of the new specimens in combination with studies on previous specimens indicates that both morphs occur in fully-grown individuals. Therefore, the dimorphism is not a result of ontogenetic change. Furthermore, the two morphs of plates do not simply come from different positions on the back of a single individual. Plates from all positions on the body can be classified as one of the two morphs, and previously discovered, isolated specimens possess only one morph of plates. Based on the seemingly display-oriented morphology of plates, female mate choice was likely the driving evolutionary mechanism rather than male-male competition. Dinosaur ornamentation possibly served similar functions to the ornamentation of modern species. Comparisons to ornamentation involved in sexual selection of extant species, such as the horns of bovids, may be appropriate in predicting the function of some dinosaur ornamentation.

Evan Thomas Saitt. 2015. Evidence for Sexual Dimorphism in the Plated Dinosaur Stegosaurus mjosi (Ornithischia, Stegosauria) from the Morrison Formation (Upper Jurassic) of Western USA.
PLoS ONE.  DOI: 10.1371/journal.pone.0123503

Thursday, April 23, 2015

[Herpetology • 2015] Rhacophorus indonesiensis | Katak Pohon Totol • A New Species of Tree Frog Genus Rhacophorus (Amphibia, Anura) from Sumatra, Indonesia

Rhacophorus indonesiensis Hamidy & Kurniati, 2015
Adult male paratype MZB Amph 21847 and a non-vouchered female from Birun, showing nocturnal coloration
photo: Mediyansyah || DOI: 10.11646/zootaxa.3947.1.3

A small-sized tree frog of the genus Rhacophorus is described on the basis of 18 specimens collected from three different localities on Sumatra Island, Indonesia. Rhacophorus indonesiensis sp. nov. is divergent from all other Rhacophorus species genetically and morphologically. The new species is distinguished from its congeners by a combination of: the presence of black spots on the ventral surfaces of the hand and foot webbing, an absence of vomerine teeth, a venter with a white kite-shaped marking, raised white spots on the dorsum or on the head, and a reddish brown dorsum with irregular dark brown blotches and distinct black dots. With the addition of this new species, fifteen species of Rhacophorus are now known from Sumatra, the highest number of species of this genus in the Sundaland region. However, with the increasing conversion of forest to oil palm cultivation or mining, the possibility of the extinction of newly described or as yet undiscovered species is of great concern.
Key words: Tree frog, Sundaland, pond-breeding, elephant wallow

 Rhacophorus indonesiensis | Katak Pohon Totol

Hamidy, Amir & Hellen Kurniati. 2015. A New Species of Tree Frog Genus Rhacophorus from Sumatra, Indonesia (Amphibia, Anura).
Zootaxa. 3947(1): 49-66. DOI: 10.11646/zootaxa.3947.1.3

[Herpetology • 2015] Limnonectes nguyenorum • More of the Same: A Diminutive New Species of the Limnonectes kuhlii complex (Anura: Dicroglossidae) from northern Vietnam

FIGURE 2. Two syntopic species of Limnonectes from northern Vietnam.
Holotype of Limnonectes nguyenorum (VMNH A.2015.1; adult male; SVL = 43.81 mm): (A) dorsal, (B) ventral, (C) plantar view of foot, (D) Palmar view of hand, (E) lateral view of head, (F) magnified view of tubercles on leg (reference in image A). Mature ova from female paratype (VMNH A.2015.3).
Specimen of L. bannaensis from type locality of L. nguyenorum (IEBR A. 2015.38; adult male; SVL = 64.50 mm): (H) lateral view of head, (I) magnified view of tubercles on leg (reference in image J), (J) dorsal, (K) ventral. All specimens imaged as they appear in alcohol.


A new species in the dicroglossid genus Limnonectes known only from Ha Giang province, Vietnam is described. Analysis of DNA sequence data from the mitochondrial 12S and 16S gene regions places the species within the Limnonectes kuhlii Complex and demonstrates it to be the sister taxon to an Indochinese clade containing L. isanensis, L. jarujini, L. megastomias, and L. taylori. The new species occurs in syntopy with L. bannaensis. Both molecular and morphological data support the recognition of this lineage as a new species. Notably, the relatively diminutive size of this species distinguishes Limnonectes nguyenorum sp. nov. from all other members of the L. kuhlii Complex.

Keywords: dicroglossid, fanged frog, Limnonectes nguyenorum sp. nov., mitochondrial DNA, morphology, species complex

Distribution and ecology. Based on the specimens examined, Limnonectes nguyenorum is only known from Vi Xuyen District in northwestern Ha Giang Province, northern Vietnam (Fig. 3). As with other members of the L. kuhii Complex, L. nguyenorum is found along streams in submontane evergreen forests (Fig. 4). The new species seems to occur at elevations approximately between 600 and 900 m, corresponding generally to the transition between lowland and montane forests. Based upon the presence of gravid females of both L. nguyenorum and L. bannaensis during collecting events, it is possible that these species breed at similar times and under similar

Etymology. The specific name, nguyenorum, is the plural possessive form of the family name Nguyen. This species is named in honor of two herpetologists who have contributed greatly to our understanding of Vietnamese herpetology and biodiversity, Truong Quang Nguyen and Tao Thien Nguyen. It is rare to find siblings working together in herpetological research, and more so to find brothers as productive as these. We commend them for their efforts and recognize that without them this and many other amphibians and reptiles in Vietnam would remain undescribed.

David S McLeod, Scuyler Kurlbaum and Ngoc Van Hoang. 2015. More of the Same: A Diminutive New Species of the Limnonectes kuhlii complex from northern Vietnam (Anura: Dicroglossidae). Zootaxa. 3947(2):201-214. DOI: 10.11646/zootaxa.3947.2.4

David S. McLeod Lab at James Madison University:

Wednesday, April 22, 2015

[Herpetology • 2015] Isanophis gen. nov. • On the Taxonomic Status of the Thai Endemic Freshwater Snake Parahelicops boonsongi, with the Erection of A New Genus (Squamata: Natricidae)

Isanophis boonsongi (Taylor & Elbel, 1958)
 Boomsong’s Stream Snake | 
Isan Keeled Stream Snake
 Isanophides / Isan Gebirgswassernattern


Parahelicops boonsongi Taylor & Elbel, 1958 is known from only three specimens from Thailand. It has been placed either in the genus Parahelicops Bourret, 1934, along with Parahelicops annamensis Bourret, 1934, or in the genus Opisthotropis Günther, 1872. We compared its morphological characters with those of P. annamensis and with three other relevant genera, Opisthotropis, Pararhabdophis Bourret, 1934, and Paratapinophis Angel, 1929. Parahelicops boonsongi is phenotypically distinct from Parahelicops annamensis, Opisthotropis, and all other natricine genera. We consequently erect a new genus, Isanophis gen. nov., to accommodate Parahelicops boonsongi.

Keywords: Serpentes, Natricidae, Parahelicops, Parahelicops annamensis, Isanophis gen. nov., Opisthotropis, Pararhabdophis, Paratapinophis

Etymology. The generic nomen Isanophis is derived from the word “Isan”, the Thai name of the north-eastern region of Thailand, and from the Greek word ophis, meaning “snake”. It describes the limited known range of this genus. This generic nomen is masculine in gender.

David, Patrick, Olivier S. G. Pauwels, Truong Q. Nguyen & Gernot Vogel. 2015. On the Taxonomic Status of the Thai Endemic Freshwater Snake Parahelicops boonsongi, with the Erection of A New Genus (Squamata: Natricidae).
Zootaxa. 3948(2): 203–217. DOI: 10.11646/zootaxa.3948.2.3
Taylor, Edward H. & Elbel, Robert E. 1958. Contribution to the Herpetology of Thailand. Univ. Kansas Sci. Bull. 38 (13): 1033-1189

Tuesday, April 21, 2015

[Ichthyology • 2015] Description of Danio absconditus and Redescription of D. feegradei (Teleostei: Cyprinidae), from the Rakhine Yoma Hotspot in south-western Myanmar

Danio absconditus Kullander & Britz, 2015
Danio feegradei Hora, 1937


Danio feegradei Hora is redescribed based on recently collected specimens from small coastal streams on the western slope of the Rakhine Yoma, ranging from the Thade River drainage southward to slightly north of Kyeintali. Danio absconditus, new species, is described from the Kyeintali Chaung and small coastal streams near Gwa, south of the range of D. feegradei. Both species are distinguished from other Danio by the presence of a dark, elongate or round spot at the base of the caudal fin and a cleithral marking composed of a small black spot margined by a much smaller orange spot. Danio feegradei is characterized by the colour pattern, with series of white spots along the otherwise dark side; D. absconditus by about 7-–11 dark vertical bars on the abdominal side. Within Danio, the presence of a complete lateral line, cleithral spot, and 14 circumpeduncular scales is shared with D. dangila and similar species, but these character states may be plesiomorphic as suggested by the shared presence of cleithral spot and complete lateral line in Devario and Betadevario. In other Danio the cleithral spot is absent, the lateral line is short or absent, and the circumpeduncular scale count is lower (10–12). Twenty teleost species are reported from streams on the western slope of the Rakhine Yoma, all probably endemic. The parapatric distribution of D. absconditus and D. feegradei is unique within the genus, and may be partly explained by changes in eustatic sea levels.

Keywords: colour pattern, freshwater fish, morphology, species discrimination, taxonomy

Kullander, Sven O. & Ralf Britz. 2015. Description of Danio absconditus, new species, and Redescription of Danio feegradei (Teleostei: Cyprinidae), from the Rakhine Yoma Hotspot in south-western Myanmar.
Zootaxa. 3948(2): 233–247. DOI: 10.11646/zootaxa.3948.2.5

Monday, April 20, 2015

[Herpetology • 2015] An Integrative Taxonomic Review of the Agamid Genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with Descriptions of New Montane and Insular Endemics; Bronchocela shenlong & B. rayaensis

Bronchocela shenlong
Grismer, Wood, Lee, Quah, Anuar, Ngadi & Sites, 2015
DOI: 10.11646/zootaxa.3948.1.1


An integrative taxonomic analysis is used to identify and describe two new species of the agamid genus Bronchocela (Kuhl) from Peninsular Malaysia: an upland species Bronchocela shenlong sp. nov. from Bukit Larut, Perak in the Bintang Mountain Range and Parit Falls, Cameron Highlands, Pahang in the Titiwangsa Mountain Range and an insular species, Bronchocela rayaensis sp. nov., from Pulau Langkawi, Kedah off the northwest coast on the border with Thailand. Both species are diagnosed from each other and all other species of Bronchocela on the basis of body shape, scale morphology, and color pattern. The analysis also demonstrates the remarkable genetic similarity of B. cristatella (Kuhl) throughout 1120 km of its range from northern Peninsular Malaysia to western Borneo despite its highly variable coloration and pattern. The two new species are appended to a rapidly growing list of newly described lizard species (60 to date) from Peninsular Malaysia tallied within the last decade.

Keywords: Peninsular Malaysia, Integrative taxonomy, Bronchocela, Langkawi Island

The Dragon of the Mountains. Introducing Bronchocela shenlong, a new species of green crested lizard from the highlands of Peninsular Malaysia that was named after one of the great dragons of Chinese mythology. It can be differentiated from its lowland congener B. cristatella by the broad white stripe on the jawline, green tympanum and red stripe along the lower edge of the tail and posterior margin of the thigh
photograph: Evan Quah
Grismer, L. L., JR. P. L. Wood, Cheol H. Lee, Evan S. H. Quah, Shahrul Anuar, Ehwan Ngadi & Jack W. Sites, Jr. 2015. An Integrative Taxonomic Review of the Agamid Genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with Descriptions of New Montane and Insular Endemics. Zootaxa. 3948(1): 1–23. DOI: 10.11646/zootaxa.3948.1.1

Sunday, April 19, 2015

[Herpetology • 2015] Gekko kwangsiensis • A New Species of the Genus Gekko Laurenti (Squamata: Sauria: Gekkonidae) from Guangxi, China

Gekko kwangsiensis Yang, 2015


A new species of the genus Gekko is described on the basis of six specimens from Wuming county of Guangxi, southern China. Gekko kwangsiensis sp. nov. is distinguished from other congeners by a combination of the following characters: body relatively small (SVL 64.2–69.7 mm in adults), slender; nares in contact with rostral; internasal absent or single; postmentals two (rarely three), enlarged; interorbital scales between anterior corners of the eyes 29–31; dorsal tubercle rows 9–11; ventral scales between mental and cloacal slit 185–208; midbody scale rows 143–156; ventral scale rows 41–45; subdigital lamellae on first toe 11–13, on fourth toe 13–18; finger and toe webbing weakly developed; tubercles absent on upper surface of fore limbs and hind limbs; precloacal pores nine or ten in males, absent in females; postcloacal tubercle single; tubercles present on dorsal surface of tail base; subcaudals enlarged; dorsal surface of body with 9 or 10 thin light bands between nape and sacrum, and dorsal surface of tail with remarkable black and white bands. Data on the natural history of the new species are provided, and the number of species in the genus Gekko recorded from China is now 17.

Keywords: Gekko, taxonomy, Guangxi Zhuang Autonomous Region

 Jian-Huan Yang. 2015. A New Species of the Genus Gekko Laurenti (Squamata: Sauria: Gekkonidae) from Guangxi, China.
Zootaxa. 3936(2):287–295. DOI: 10.11646/zootaxa.3936.2.9

[Herpetology • 2015] Cnemaspis adii • A New Species of Cnemaspis (Sauria: Gekkonidae) from northern Karnataka, India

 Cnemaspis adii 
Srinivasulu, Kumar & Srinivasulu, 2015


A new species of rupicolous gecko of the genus Cnemaspis is described from Hampi, Karnataka, southern India. Cnemaspis adii sp. nov. is diagnosable from all the Indian congeners in possessing the following suite of characters: medium-sized Cnemaspis, SVL less than 35 mm (31.7–34.9). Dorsal scales on the trunk homogeneous, small, granular and feebly keeled. Spine-like tubercles absent on the flanks. Mental subtraingular, two pairs of postmentals, primary pair separated by a single chin shield. Ventral scales on the trunk smooth, imbricate; 22–26 scales across the belly. Supralabial I narrowly in contact with nasal. Dorsal aspect of forelimbs and hindlimbs are weakly unicarinate. Lamellae under the digit IV of pes 20–22. Males with two precloacal pores, two femoral pores on each side of the thigh. The existence of the species in a World Heritage Site with continuous anthropogenic interference ascertains the robustness of the species and need for additional herpetofaunal explorations to reveal the total diversity of species of the genus Cnemaspis in peninsular India.

Keywords: Cnemaspis, description, Hampi, Gekkonidae, Karnataka, India

Srinivasulu, Chelmala, Gandla C. Kumar & Bhargavi Srinivasulu. 2015. A New Species of Cnemaspis (Sauria: Gekkonidae) from Northern Karnataka, India.
Zootaxa. 3947(1): 85–98. DOI: 10.11646/zootaxa.3947.1.5

Hyderabad researcher discovers new species of gecko, Cnemaspis adii, in ruins of Hampi in Karnataka
A new species of the day gecko, a type of lizard usually found in warm climates, has been spotted at the ruins of the World Heritage Site of Hampi in Karnataka. The gecko has been named Cnemaspis adii after a young herpetology researcher from Hyderabad, Aditya Srinivasulu.