[Ichthyology • 2018] Gymnogeophagus peliochelynion • A New Mouth Brooder Species of Gymnogeophagus with Hypertrophied Lips (Cichliformes: Cichlidae) from A Tributary of the río Uruguay

Gymnogeophagus peliochelynion

Turcati, Serra-Alanis & Malabarba, 2018

DOI: 10.1590/1982-0224-20180118  

ABSTRACT

A new mouth breeder species of Gymnogeophagus is described from a tributary of the río Uruguay. It is distinguished from most species of the genus by the presence of hypertrophied lips, and from G. labiatus and G. pseudolabiatus by the color pattern. The presence of successive allopatric species of the Gymnogeophagus gymnogenys clade inhabiting the tributaries of the río Uruguay is discussed.

Keywords: Distribution; Endemism; Gymnogeophagus gymnogenys clade; New species; Río Uruguay

Fig. 2 Head of Gymnogeophagus peliochelynion (first column), G. pseudolabiatus (second column) and G. mekinos (third column) showing the entirely black hump in males in G. peliochelynion (vs. yellow with black margin), and upper lip not folded dorsally over anterior margin of snout (vs. upper lip folded dorsally in G. pseudolabiatus and undeveloped in G. mekinos).
G. peliochelynion from top to bottom, paratype, ZVC-P 13210, paratype, 76.3 mm SL; ZVC-P 7016, 89.9 mm SL; ZVC-P 13057, 90.2 mm SL.
 G. pseudolabiatus from top to bottom, paratype, UFRGS 7754, 102.0 mm SL; MHNM 4010, 88.8 mm SL; MHNM 4010, 95.3 mm SL.
G. mekinos from top to bottom, MHNM 3511, 105.1 mm SL; MHNM 3511, 97.2 mm SL; MHNM 4009, 121.3 mm SL. 

Fig. 4 Gymnogeophagus peliochelynion: top, holotype, male, ZVC-P 12493, 101.9 SL; bottom, paratype, female, UFRGS 8076, 77.2 SL. Both from arroyo de las Tunas on road 31, tributary of río Arapey Grande, Salto, Uruguay. Photographs taken just after collection and fixation in formalin.

Gymnogeophagus peliochelynion, new species

Diagnosis. The new species can be distinguished from the species of the Gymnogeophagus rhabdotus group and from G. balzanii by the shape of the caudal peduncle longer than deep (vs. deeper than long). It is distinguished from all congeners, except G. labiatus and G. pseudolabiatus, by the possession of thick lips. It differs from G. labiatus and G. lacustris by the lack of an oblique bar from the eye to the dorsal-fin origin (vs. oblique bar present), and by the color pattern of the caudal, dorsal and anal fins with dots (vs. caudal fin and posterior portion of anal fin with longitudinal hyaline stripes). It differs from G. pseudolabiatus and G. mekinos by the hump entirely black in males (Fig. 2; vs. yellow with black margin), and upper lip not folded dorsally over anterior margin of snout (vs. upper lip folded dorsally, usually with a well-developed medial lobe dorsally projected in G. pseudolabiatus).

....

Ecological notes. The new species was collected in rivers with clear water, usually with rocky or muddy bottom and little vegetation.

Etymology. The name peliochelynion is from the Greek pelios, meaning black and blue, and chelyne, meaning lip, in reference to the color of the lips of the new species. A name in apposition.

Fig. 5 Gymnogeophagus peliochelynion: above, paratype, male, ZVC-P 13210, paratype, 76.3 mm SL, río Arapey, Colonia Lavalleja, Paso Elías, Salto, Uruguay; below, female, ZVC-P 13057, 65.3 mm SL, Arroyo Sopas, Paso del Cementerio, Salto, Uruguay. Photographs of live specimens.

Andréia Turcati, Wilson Sebastián Serra-Alanis and Luiz R. Malabarba. 2018. A New Mouth Brooder Species of Gymnogeophagus with Hypertrophied Lips (Cichliformes: Cichlidae). Neotrop. ichthyol. 16(4). DOI: 10.1590/1982-0224-20180118 

 twitter.com/MJBernt/status/1072211391392727041

RESUMEN: Una nueva especie incubadora bucal de Gymnogeophagus es descripta de un tributario del Río Uruguay. Se distingue de la mayoría de las especies del género por la presencia de labios hipertrofiados, y de G. labiatus y G. pseudolabiatus por su patrón de coloración. Se discute la presencia de sucesivas especies alopátricas del clado Gymnogeophagus gymnogenys habitando los tributarios del Río Uruguay.

Palabras Clave: Distribución; Endemismo; Clado Gymnogeophagus gymnogenys; Especie nueva; Río Uruguay

[Ichthyology • 2018] Austrolebias queguay • A New Species of Annual Killifish (Cyprinodontiformes, Rivulidae) Endemic to the lower Uruguay River Basin

Austrolebias queguay

Serra & Loureiro, 2018  

  DOI: 10.3897/zse.94.29115  

Abstract
In this article we describe a new species of the annual fish genus Austrolebias from the lower Uruguay river basin. The fusion of the urogenital papilla to the first anal fin ray in males and the pigmentation pattern, indicates a close relationship with the clade formed by A. bellottii, A. melanoorus, and A. univentripinnis. The new species can be differentiated from those by the following combination of characters: presence of well-defined light bands contrasting with the sides of the body, the distal portion of the anal fin dark gray, pelvic fins dark bluish green and bases united at about 50–80% on their medial margins, pectoral fins with iridescent blue sub-marginal band, and general coloration of body bluish green. The new species can only be found in wetlands of the Queguay river, an area included in the Uruguayan protected areas system and represents so far the only annual fish species endemic to the lower Uruguay river basin.

Key Words: Austrolebias bellottii species group, Systematics, La Plata basin

Austrolebias queguay sp. n. 
Figure 2. A. paratype male (ZVCP 11620); B. non type male, not preserved (right side, photo flipped)

Figure 2. A. Austrolebias queguay sp. n. paratype male (ZVCP 11620); B. A. queguay non type male, not preserved (right side, photo flipped); C. i non preserved male; D. A. univentripinnis male (UFRGS 18064, right side photo flipped); E. A. melanoorus topotype male (ZVCP13651); F. Detail of pectoral and pelvic fins of A. melanoorus; G. Detail of pectoral and pelvic fins of  A. queguay.

Figure 3. A. Austrolebias queguay sp. n. female: paratype ZVCP 11620; B. A. bellottii female (ZVCP 11560); C. A. univentripinnis female (UFRGS 18066); D. A. melanoorus topotype female (ZVCP 13651).

Austrolebias queguay sp. n.

Austrolebias sp. in Loureiro et al. (2018)

Diagnosis: The new species differs from all the other species of the genus except Austrolebias bellottii, A. univentripinnis and A. melanoorus, by the presence of the urogenital papilla attached to the anal fin in males (vs. free from the anal fin). It differs from A. bellottii and A. univentripinnis by the presence of well-defined light blue bands contrasting with the sides of the body in adult males (vs. vertical rows of light blue dots) (Fig. 2); from A. melanoorus, by the presence of dark gray coloration of the distal portion of the anal fin in males (vs. distal portion of anal-fin black), pelvic-fins dark bluish green (observed in ventral view) and bases united at about 50–80% on their medial margins (vs. dark gray and united about 50% or less), pectoral-fins with iridescent blue sub-marginal band (vs. sub-marginal band absent), and general coloration of the body bluish green (vs. grayish sky blue).

 Figure 6. Geographic distribution of Austrolebias queguay sp. n. (orange dots, Uruguay river basin), A. bellottii (yellow dots, Paraná and Uruguay river basins), A. melanoorus (red dots, Negro and Yaguarón river basins), and A. univentripinnis (green dots, Yaguaron river basin). Orange star indicates type locality.

Figure 7. Type locality of Austrolebias queguay sp. n., wetlands of middle Queguay river basin.

Etymology: The specific name, queguay, is in reference to Queguay river basin, the type locality of the new species, treated as a noun in apposition to the generic name.

Distribution: Austrolebias queguay is endemic to the wetlands of middle Queguay river basin (30 meters above sea level), Paysandú Department, Uruguay, which flows to the lower Uruguay river (Fig. 6).

Ecology: As many species of the family Rivulidae, A. queguay presents an annual life cycle which includes drought resistant eggs and diapausing embryos. All species of Austrolebias are obligate annuals (Berois et al. 2016). In the Pampa biome there is not a defined dry season, so dried environments can be found between mid spring to early fall (depending on the year), when evaporation is higher than precipitations (Williams 2006; García et al. 2017). Austrolebias species can be found in small grassland ponds and seasonal floodplain wetlands; however the new species has been found so far only in the latter environments (Fig. 7).


 Wilson S. Serra and Marcelo Loureiro. 2018. Austrolebias queguay (Cyprinodontiformes, Rivulidae), A New Species of Annual Killifish Endemic to the lower Uruguay River Basin. Zoosystematics and Evolution. 94(2): 547-556.  DOI: 10.3897/zse.94.29115

[PaleoMammalogy • 2018] Neoglyptatelus uruguayensis • A New Species of Neoglyptatelus (Xenarthra, Cingulata) from the Late Miocene of Uruguay Provides New Insights on the Evolution of the Dorsal Armor in Cingulates

Neoglyptatelus uruguayensis 

Fernicola, Rinderknecht, Jones, Vizcaíno & Porpino, 2018

DOI: 10.5710/AMGH.02.12.2017.3150 
 twitter.com/Ameghiniana

Abstract

The genus Neoglyptatelus Carlini, Vizcaíno and Scillato-Yané has been considered a member of Glyptatelinae, a group encompassing the purportedly basal-most glyptodonts. It is up to now represented by two species from Colombia: Neoglyptatelus originalis Carlini, Vizcaíno and Scillato-Yané, from the middle Miocene (a carapace fragment, isolated osteoderms and postcranial bones), and Neoglyptatelus sincelejanus Villarroel and Clavijo, from the middle or late Miocene (a partial carapace and a caudal armor). More scarce material assigned to this genus was recovered from the late Miocene of Uruguay and Brazil. In this article, we describe a new species, Neoglyptatelus uruguayensis, from the late Miocene Camacho Formation, Uruguay, based on an almost complete carapace and several postcranial bones. We conducted a phylogenetic analysis based on 167 morphological characters (23 new ones and 144 from previous analysis) scored for 19 taxa, encompassing some of the best known glyptodontid genera, one pampathere and four armadillos (including the enigmatic genus Pachyarmatherium Downing and White). In the most parsimonious tree that was obtained, Neoglyptatelus forms a clade with Pachyarmatherium (Pachyarmatheriidae), which is the sister group of the glyptodonts + pampatheres clade; consequently, it is not a glyptodont, as previously believed. This result, together with the known stratigraphic and geographic distribution of Neoglyptatelus and Pachyarmatherium, suggests that this new cingulate clade originated in South America and that Pachyarmatherium reached North America during the Plio—Pleistocene. The carapace of Neoglyptatelus and Pachyarmatherium comprises pelvic and scapular shields overlapping each other without separate intervening transverse mobile bands, an arrangement that differentiates both genera from the remaining cingulates.

Keywords: Carapace, Movement, Pachyarmatheriidae, Glyptatelinae, Phylogeny, Caparazón, Movimiento, Pachyarmatheriidae, Glyptatelinae, Filogenia

Figure 2. Complete specimen of the holotype of Neoglyptatelus uruguayensis (MHNM 1642).
1, dorsal view; 2, lateral view; 3, life reconstruction.

Scale bar= 50 mm.

SYSTEMATIC PALEONTOLOGY 

Order CINGULATA Illiger, 1811

Family PACHYARMATHERIIDAE new rank 

Type genus. Pachyarmatherium Downing and White, 1995.

Genus NEOGLYPTATELUS Carlini, Vizcaíno, and Scillato-Yané, 1997

Type species. Neoglyptatelus originalis Carlini, Vizcaíno, and ScillatoYané, 1997

Neoglyptatelus uruguayensis sp. nov.

Derivation of name. uruguayensis; from Uruguay, to emphasize its geographic provenance.

 Juan C. Fernicola, Andrés Rinderknecht, Washington Jones, Sergio F. Vizcaíno and Kleberson Porpino. 2018. A New Species of Neoglyptatelus (Mammalia, Xenarthra, Cingulata) from the Late Miocene of Uruguay Provides New Insights on the Evolution of the Dorsal Armor in Cingulates [UNA NUEVA ESPECIE DE NEOGLYPTATELUS (MAMMALIA, XENARTHRA, CINGULATA) DEL MIOCENO TARDÍO DE URUGUAY PROPORCIONA NUEVAS PERSPECTIVAS SOBRE LA EVOLUCIÓN DEL CAPARAZÓN DORSAL EN LOS CINGULADOS]. Ameghiniana. 55(3):233-252. DOI: 10.5710/AMGH.02.12.2017.3150

 twitter.com/Ameghiniana/status/1032306918990249984

facebook.com/Ameghiniana/photos/2120901014837479

Resumen: El género Neoglyptatelus Carlini, Vizcaíno and Scillato-Yané ha sido considerado un miembro de los Glyptatelinae, un grupo que supuestamente incluye a los gliptodontes basales. Actualmente, está representado por dos especies halladas en Colombia: Neoglyptatelus originalis Carlini, Vizcaíno and Scillato-Yané, del Mioceno medio (fragmento de caparazón, osteodermos aislados y elementos postcraneanos) y Neoglyptatelus sincelejanus Villarroel y Clavijo del Mioceno medio o tardío (parte de un caparazón y coraza caudal). Osteodermos aislados asignados a este género fueron recuperados en el Mioceno tardío de Uruguay y Brasil. En este trabajo, nosotros describimos una nueva especie, Neoglyptatelus uruguayensis, del Mioceno tardío de la Formación Camacho, Uruguay, a base de una coraza casi completa y distintos elementos postcraneanos. Nuestro estudio filogenético se sustenta en 167 caracteres óseos (23 nuevos y 144 utilizados previamente) relevados en 19 taxones, que incluyen los géneros de gliptodontes mejor conocidos, un pampaterio y cuatro armadillos, incluyendo Pachyarmatherium Downing y White. En el árbol más parsimonioso obtenido Neoglyptatelus forma con Pachyarmatherium un clado (Pachyarmatheriidae) hermano del grupo formado por pampaterios + gliptodontes, por lo cual no es un gliptodonte como se sostenía previamente. Este resultado, junto al conocimiento estratigráfico y geográfico de Neoglyptatelus y Pachyarmatherium, sugiere que los Pachyarmatheriidae se habrían originado en América del Sur y que durante el Plio—Pleistoceno Pachyarmatherium arribó a América del Norte. El caparazón de Neoglyptatelus y Pachyarmatherium contiene un escudo escapular y otro pélvico que solapan directamente entre sí, sin mediar bandas movibles transversas; este arreglo morfológico es único entre los cingulados.

[Herpetology • 2017] Liolaemus gardeli • A New Species of Lizard of the L. wiegmannii group (Iguania: Liolaemidae) from the Uruguayan Savanna

Liolaemus gardeli 

Verrastro, Maneyro, da Silva & Farias, 2017

 DOI:  10.11646/zootaxa.4294.4.4 ภาพ

Abstract

Lizards of the Liolaemus wiegmannii group comprise 11 species that are widely distributed east of the Andes, occurring in Argentina, Uruguay, and Brazil. Here we analyzed a population of the genus Liolaemus, found in the Pampa region of Uruguay, on isolated sand dunes along the Tacuarembó River. We conducted biometric, meristic, and genetic comparisons between this population and other populations of L. weigmannii from Argentina and Uruguay, and the other species of the L. weigmannii complex. Our morphological and genetic analyses showed that this population represents to a new species of the genus Liolaemus, belonging to the L. wiegmannii group. To date, its known distribution is extremely restricted.

Keywords: Reptilia, endemic species, phylogeny, sand habitat, Pampa biome

  

 Laura Verrastro, Raúl Maneyro, Caroline M. da Silva and Iraia Farias. 2017. A New Species of Lizard of the L. wiegmannii group (Iguania: Liolaemidae) from the Uruguayan Savanna. Zootaxa. 4294(4); 443–461. DOI:  10.11646/zootaxa.4294.4.4

[Ichthyology • 2016] Gymnogeophagus terrapurpura • A New Species of Gymnogeophagus Miranda Ribeiro (Teleostei: Cichliformes) from Uruguay

Gymnogeophagus terrapurpura 

Loureiro, Zarucki, Malabarba & González-Bergonzoni, 2016 

DOI: 10.1590/1982-0224-20150082

ABSTRACT

We describe a new species of a substrate-brooding Gymnogeophagus , based on coloration characters. The new species can be distinguished from the remaining substrate-brooding species in the genus by the unique pigmentation of the dorsal fin which consists of light blue, diagonal stripes over a red background in the spiny section and a combination of round, elliptic, and elongated bright blue spots over a red background in the soft section. It can be further distinguished from all other species of Gymnogeophagus by the following combination of characters: a discontinuous bright blue band above the upper lateral line in the humeral area, light blue roundish spots over a red to orange background on the anal fin, and conspicuous bright blue horizontal bands on body. The new species inhabits a wide range of freshwater habitats in the lower rio Uruguay basin, Rio de la Plata coastal drainages and Atlantic Ocean coastal drainages in Uruguay.

Keywords: Cichlidae; Gymnogeophagus meridionalis; Gymnogeophagus rhabdotus; Substrate brooding; Systematics

Fig. 2: Non-type live specimens of (A) Gymnogeophagus terrapurpura from rio Santa Lucía basin, Canelones Department, Uruguay,
(B) G. rhabdotus and (C) G. meridionalis. (B) and (C) both from rio Negro basin, Tacuarembó Department, Uruguay. 

Sexual dimorphism. There is no evident sexual dimorphism except from slight differences in size. In the breeding season males are usually larger than females (10%-50% larger).

Geographic distribution. Gymnogeophagus terrapurpura is distributed from the rio Arapey (in the middle rio Uruguay basin) to all tributaries of the lower rio Uruguay basin and in the eastern coastal drainages of the Río de la Plata estuary and Atlantic Ocean in Uruguay (Fig. 6).

Ecological notes. Gymnogeophagus terrapurpura has been found in a large variety of habitats including shallow lakes, floodplain lakes, rivers, and streams, mainly associated with cobble, sandy or muddy substrates. Gymnogeophagus terrapurpura is an omnivore, feeding on zooplankton, macroinvertebrates, and periphyton (to a lesser extent), reducing greatly its feeding activity during autumn and winter when water temperature decreases, as described in a low-impacted lowland stream in Florida Department by González-Bergonzoni et al. (2016, named therein as "Gymnogeophagus sp.") and in a eutrophic shallow lake of Montevideo by Yafe et al. (2002, named therein as G. rhabdotus). Gymnogeophagus terrapurpura has been observed to brood on the substrate and exhibits substantial parental care, including aggressive territorial behavior in both males and females during the reproductive season, from November to January.

Etymology. The specific epithet refers to the G. E. Hudson novel "La Tierra Purpúrea" (as it is known in the Spanish translation), in which the main character makes a trip through the same region where the new species occurs. A noun in apposition.

Conservation status. Gymnogeophagus terrapurpura is relatively frequent and abundant in the rio Negro, the middle and lower rio Uruguay basin, and in the oriental coastal drainages of Río de la Plata estuary and Atlantic Ocean in Uruguay. No specific threats were detected, and the species can be categorized as Least Concern (LC) according to IUCN criteria (International Union for Conservation of Nature (IUCN), 2011).

Marcelo Loureiro, Matías Zarucki, Luiz R. Malabarba and Iván González-Bergonzoni. 2016.  A New Species of Gymnogeophagus Miranda Ribeiro from Uruguay (Teleostei: Cichliformes).
Neotropical Ichthyology. 14(1); e150082.  DOI: 10.1590/1982-0224-20150082

RESUMEN: Describimos una nueva especie de Gymnogeophagus con incubación de sustrato, en base a caracteres cromáticos. La nueva especie puede ser diferenciada de las restantes especies del género por la exclusiva pigmentación de la aleta dorsal, la que consiste de bandas oblicuas de color celeste sobre fondo rojo en la porción espinosa y una combinación de manchas redondas, elípticas, o alargadas azul brillantes sobre fondo rojo en la porción blanda. Además puede ser distinguida de las otras especies de Gymnogeophagus por la siguiente combinación de caracteres: banda horizontal azul iridiscente situada por encima de la línea lateral superior discontinua en la región humeral, manchas azules brillantes redondeadas sobre fondo rojo anaranjado en la aleta anal, bandas horizontales azul brillante en el cuerpo conspicuas. La nueva especie habita una amplia gama de hábitats de agua dulce en el sector inferior de la cuenca del río Uruguay, los drenajes costeros del Río de la Plata y drenajes costeros del Océano Atlántico en Uruguay.

[Arachnida • 2017] Allocosa marindia • On the Taxonomy of southern South American Species of the Wolf Spider Genus Allocosa (Araneae: Lycosidae: Allocosinae)

Allocosa marindia  Simó, Lise, Pompozzi & Laborda, 2017

Living specimen of Allocosa marindia sp. nov. from Playa Pascual, San José, Uruguay.

  DOI:  10.11646/zootaxa.4216.3.4 

Abstract

Three species of the genus Allocosa Banks, 1900 from southern South America are redescribed: Allocosa alticeps (Mello-Leitão, 1944), A. brasiliensis (Petrunkevitch, 1910) and A. senex (Mello-Leitão, 1945). The female of A. senex is described for the first time and the species is revalidated. A new species, Allocosa marindia sp. nov. from southern Uruguay and southern Brazil is described. The new species is distinguished by the flattened terminal apophysis of the male bulb and the conspicuous pointed projections on the posterior margin of the female epigynum. The species inhabits in sandy estuarine and oceanic coasts with psammophile vegetation.

Keywords: Araneae, Neotropical, wolf spiders, redescription, new species

Allocosa marindia

Etymology. The specific epithet is a noun in apposition taken from the type locality, where, for a long time, several studies on Uruguayan wolf spiders have been developed.

Distribution. Southern Uruguay and coastal Rio Grande do Sul, Brazil. 

Taxonomy

Lycosidae Sundevall, 1833

Allocosinae Dondale, 1986

Allocosa Banks, 1900. 

Type species by monotypy, A. funerea (Hentz, 1844)

• Allocosa marindia sp. nov. 

• Allocosa brasiliensis (Petrunkevitch, 1910)

• Allocosa senex (Mello-Leitão, 1945) revalidated  

• Allocosa alticeps (Mello-Leitão, 1944)  

Miguel Simó, Arno A. Lise, Gabriel Pompozzi and Álvaro Laborda. 2017. On the Taxonomy of southern South American Species of the Wolf Spider Genus Allocosa (Araneae: Lycosidae: Allocosinae). Zootaxa. 4216(3); 261–278. DOI:  10.11646/zootaxa.4216.3.4

[Ichthyology • 2008] Australoheros forquilha, A. charrua, A. minuano & A. guarani • The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages

Australoheros forquilha Rícan & Kullander, 2008

http://www.pecescriollos.de/go/australoheros-forquilha

Australoheros charrua Rícan & Kullander, 2008 

Australoheros forquilha Rícan & Kullander, 2008 

Australoheros guarani Rícan & Kullander, 2008 

Australoheros minuano Rícan & Kullander, 2008 

Abstract

Australoheros Říčan and Kullander, 2006 includes four described species 

— Australoheros facetus (Jenyns, 1842), Australoheros tembe (Casciotta, Gómez & Toresani, 1995), Australoheros scitulus (Říčan & Kullander, 2003) and Australoheros kaaygua Casciotta, Almirón & Gómez, 2006. 

Four additional species are newly described in this paper based on results presented in Říčan and Kullander (2006): Australoheros forquilha sp. nov., from the tributaries of the Upper Río Uruguay in Brazil; Australoheros charrua sp. nov. and Australoheros minuano sp. nov., from the tributaries of the Middle Río Uruguay in Brazil and Uruguay, and Australoheros guarani sp. nov. from the Río Paraná tributaries in Paraguay. Sympatric species are not found closely related and all sister species have allopatric distributions. 

Four species groups are recognized based on phylogenetic hypotheses generated from morphological and cyt b characters; the forquilha group—A. forquilha, A. tembe; the scitulus group—A. scitulus, A. charrua; the kaaygua group—A. kaaygua, A. minuano; and the facetus group—A. facetus, A. guarani.

Key words: ´Cichlasoma´ facetum group, Australoheros, new species, Cichlidae, South America, biogeography, Río Uruguay, Río Paraná

Rícan, O. & Kullander, S.O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724: 1-51. 

http://www.mapress.com/zootaxa/2008/f/z01724p051f.pdf

http://www.practicalfishkeeping.co.uk/content.php?sid=1585

[Ichthyology • 2003] Australoheros ‘Cichlasoma’ scitulum • a new species of cichlid fish from the Río de la Plata region in Argentina, Brazil, and Uruguay

Australoheros ‘Cichlasoma’ scitulum

Rícan & Kullander, 2003

‘Cichlasoma’ scitulum, new species, is described from the Rı´o Rosario drainage in the Río de La Plata region of Uruguay, and from lower Río Uruguay tributaries in Argentina, Uruguay and Brazil. It is easily distinguished from the other two species of the ‘Cichlasoma’ facetum group in meristic and coloration characters. ‘Cichlasoma’ scitulum has the highest anal-fin spine counts of the three species and it is also

characterized by having black-spotted dorsal, anal and caudal fins and flank and opercular scales.

Rícan, O. & S. O. Kullander. 2003. ‘Cichlasoma’ scitulum: a new species of cichlid fish from the Río de la Plata region in Argentina, Brazil, and Uruguay. – Copeia 2003, 4: 794–802.: http://www.aqvaterra.com/docs/scitulum.pdf

http://www.fishbase.org/summary/Australoheros-scitulus.html