Sunday, February 7, 2016

[Herpetology • 2016] Cyrtodactylus klakahensis • A New Species of Bent-toed Gecko, Genus Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with Taxonomic Remarks on C. fumosus (Müller, 1895)


Cyrtodactylus klakahensis
Hartmann, Mecke, Kieckbusch, Mader & Kaiser, 2016 

Abstract

A new species of the gekkonid lizard genus Cyrtodactylus Gray, 1827 is described from Klakah, Lumajang Regency, Jawa Timur Province, Java, Indonesia. Cyrtodactylus klakahensis sp. nov. can be distinguished from all other congeners by the presence of (1) a deep precloacal groove in males, (2) three rows of enlarged precloacofemoral scales, of which the third row bears 37–38 pores in males, (3) three or four rows of enlarged scales between the precloacofemoral scale rows and the cloaca, forming distinct chevrons, (4) raised and strongly keeled dorsal tubercles in 15–19 rows at midbody, (5) an indistinct lateral fold, (6) 17–20 subdigital lamellae under the 4th toe, and (7) subcaudal scales which are not transversely enlarged. Cyrtodactylus klakahensis sp. nov. is only the third bent-toed gecko species described from Java, indicating that the diversity of this genus on this island has been neglected in the past. Furthermore, we confirm that C. fumosus (Müller, 1895) is a species that possesses a precloacal groove in males and is most likely restricted to northern Sulawesi. That species is defined by a single female holotype (NMB-REPT 2662). Specimens in museum collections catalogued as C. fumosus from localities elsewhere are misidentified and likely represent undescribed species.

Keywords: Reptilia, Cyrtodactylus klakahensis sp. nov., C. fumosus, C. marmoratus, Lacertilia, Gekkonidae, bent-toed geckos, East Java, Indonesia, Greater Sunda Islands, morphology


FIGURE 3. Comparison of the shape and arrangement of dorsal tubercles at midbody, the precloacal region in males, and the postmental and gular scale pattern between Cyrtodactylus klakahensis sp. nov., C. fumosus, and C. marmoratus.
Cyrtodacytlus klakahensis sp. nov.: A) Closely arranged, trihedral, raised, and strongly keeled dorsal tubercles at midbody; B) eidonomy of precloacofemoral scales (three series), precloacal groove, and posterior precloacal scales; C) gular region, showing the presence of enlarged 2nd postmentals (photos and drawings of SMF 22476, holotype).
Cyrtodactylus fumosus: D) Widely scattered, roundish, flat, and smooth dorsal tubercles at midbody; E) eidonomy of precloacofemoral scales (only one series distinctly enlarged), precloacal groove, and posterior precloacal scales; F) gular region, indicating the absence of enlarged 2nd postmentals (D, F = NMB-REPT 2662, holotype; E = BMNH 1896.12.9.3).
Cyrtodactylus marmoratus: G) Closely arranged, slightly raised and keeled dorsal tubercles at midbody; H) eidonomy of precloacofemoral scales (three series) and precloacal groove, with posterior precloacal scales absent; I) gular region, indicating the absence of enlarged 2nd postmentals and the presence of a single pair of enlarged gular scales bordering the single pair of enlarged postmentals posteriorly (G = MTKD 8903; H, I = RMNH.RENA 2710a.1, lectotype).
Photos by Sven Mecke; line drawings by Felix Mader (based on photos by Sven Mecke).

Hartmann, Lukas, Sven Mecke, Max Kieckbusch, Felix Mader and Hinrich Kaiser. 2016. A New Species of Bent-toed Gecko, Genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with Taxonomic Remarks on C. fumosus (Müller, 1895). Zootaxa. 4067(5): 552–568.  DOI: 10.11646/zootaxa.4067.5.2

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