Wednesday, October 3, 2012

[Paleontology • 2012] Heterodontosauridae: Taxonomy, morphology, masticatory function and phylogeny of heterodontosaurid dinosaurs

Credit: Photo and sculpting by Tyler Keillor
  Heterodontosaurus flesh model and skull. Skin, scales and quills are added to a cast of the skull of Heterodontosaurus, the best known heterodontosaurid from South Africa. 


Abstract
Heterodontosaurids comprise an important early radiation of small-bodied herbivores that persisted for approximately 100 My from Late Triassic to Early Cretaceous time. Review of available fossils unequivocally establishes Echinodon as a very small-bodied, late-surviving northern heterodontosaurid similar to the other northern genera Fruitadens and Tianyulong. Tianyulong from northern China has unusual skeletal proportions, including a relatively large skull, short forelimb, and long manual digit II. The southern African heterodontosaurid genus Lycorhinus is established as valid, and a new taxon from the same formation is named Pegomastax africanus gen. n., sp. n. Tooth replacement and tooth-to-tooth wear is more common than previously thought among heterodontosaurids, and in Heterodontosaurus the angle of tooth-to-tooth shear is shown to increase markedly during maturation. Long-axis rotation of the lower jaw during occlusion is identified here as the most likely functional mechanism underlying marked tooth wear in mature specimens of Heterodontosaurus. Extensive tooth wear and other evidence suggests that all heterodontosaurids were predominantly or exclusively herbivores. Basal genera such as Echinodon, Fruitadens and Tianyulong with primitive, subtriangular crowns currently are known only from northern landmasses. All other genera except the enigmatic Pisanosaurus have deeper crown proportions and currently are known only from southern landmasses.
Keywords: Dinosauria, Heterodontosauridae, Heterodontosaurinae, Heterodontosaurus, tooth replacement, tooth wear, herbivory


Credit: Drawing by Todd Marshall 
new dinosaur dwarf Pegomastax from South Africa. With jaws only 1-inch in length, plant-eating Pegomastax ("thick jaw") is one of the smallest dinosaurs ever discovered. 

Pegomastax africanus | New fanged dwarf dinosaur from southern Africa, ate plants 
New fanged dwarf dinosaur from southern Africa, ate plants: http://www.eurekalert.org/pub_releases/2012-10/pp-nfd100112.php

Sereno, P.C. 2012. Taxonomy, morphology, masticatory function and phylogeny of heterodontosaurid dinosaurs. ZooKeys. 224: 1-225. doi: http://dx.doi.org/10.3897/zookeys.226.2840