[Herpetology • 2010] Bryophryne abramalagae & B. flammiventris • Two New Species of Bryophryne (Anura: Strabomantidae) from High Elevations in Southern Peru (Region of Cusco)

Abstract

We describe two new species of Bryophryne from the Region of Cusco, Provincia de La Convención in southern Peru, increasing the number of currently known Bryophryne to eight. One of the new species is the second known species of Bryophryne that has a tympanic annulus and tympanic membrane. Males of this species have vocal slits, a vocal sac, and call from inside moss. It is readily distinguished from all its congeners by having a blackish-brown venter with yellow, orange, or pale pink blotches. This species is found at elevations of 3800–3850 m in the puna along the road from Vilcabamba to Pampascona. The second new species has the venter orange and pale gray mottled in males, whereas the venter is black and pale gray mottled in the single female. It was found inside bunches of the Peruvian feather grass (Stipa ichu) in the puna of Abra Málaga next to the road at an elevation of 4000 m. A map showing the type localities of all currently known species of Bryophryne is presented.

Keywords: Anura, Bryophryne, New species, Puna, Strabomantidae

FIG. 3.—Map illustrating the type localities of Bryophryne in Peru.

Holotype of Bryophryne abramalagae in life (MUSM 27631, male, SVL 17.8 mm)

Holotype of Bryophryne flammiventris in life (MUSM 27613, male, SVL 19.8 mm)

2010. Two New Species of Bryophryne (Anura: Strabomantidae) from High Elevations in Southern Peru (Region of Cusco). Herpetologica. 66(3):308-319.

http://www.bioone.org/doi/abs/10.1655/09-038.1

http://www.ruffordsmallgrants.org/files/Herpetologica,%2066(3),%202010,%20308%E2%80%93319.pdf

[Herpetology • 2009] Bryophryne gymnotis, B. hanssaueri & B. zonalis • Three New Species of Bryophryne (Anura: Strabomantidae) from the Region of Cusco, Peru

Bryophryne hanssaueri Lehr and Catenazzi, 2009

Bryophryne gymnotis Lehr and Catenazzi, 2009

Bryophryne hanssaueri Lehr and Catenazzi, 2009

Bryophryne zonalis Lehr and Catenazzi, 2009

Abstract

We describe three new species of Bryophryne from the Region of Cusco in southern Peru, increasing the number of currently known Bryophryne to six. One of the new species differs from all species of Bryophryne in having a tympanic annulus and tympanic membrane. Males of this species have vocal slits, a vocal sac, and produce a call, which we analyze herein. This species is found at San Luis, a montane cloud forest along the road from Abra Malaga to Quillabamba at elevations between 3272 and 3354 m. The second new species has an orange throat and groin and is found along the Ericsson trail that connects Acjanaco to Pillahuata in Manu National Park and near Abra Acjanaco along the Paucartambo-Pilcopata road at elevations between 3266 and 3430 m. The third new species has the throat and chest mottled pale gray and tan, whereas the belly is black with white flecks. This species is only known from the upper Marcapata valley along the road from Abra Huallahualla to Quincemil at elevations between 3129 and 3285 m. Ecological observations for all new species are described and a map showing the type localities of all currently known species of Bryophryne is presented.

Keywords: Anura, Bryophryne, new species, Strabomantidae, vocalization

Bryophryne gymnotis Lehr and Catenazzi, 2009

Bryophryne hanssaueri Lehr and Catenazzi, 2009

Bryophryne hanssaueri Lehr and Catenazzi, 2009

Bryophryne zonalis Lehr and Catenazzi, 2009

photo: http://calphotos.berkeley.edu/cgi/img_query?enlarge=0000+0000+0809+1551

Lehr and Catenazzi 2009. Three New Species of Bryophryne (Anura: Strabomantidae) from the Region of Cusco, Peru. South American Journal of Herpetology. 4(2):125-138. http://bioone.org/doi/abs/10.2994/057.004.0204

[Herpetology • 2008] Bryophryne nubilosus • A new species of Bryophryne (Anura: Strabomantidae) from southern Peru

Abstract

A new species of Bryophryne from southern Peru (Cusco Region) is described. Specimens were found in the leaf litter of cloud forest at elevations of 2350–3215 m. The new species has a maximum snout-vent length of 21.9 mm in adult females, 18.9 mm in adult males and is the smallest species of the genus. It lacks a tympanum and dentigerous processes of vomers, has dorsolateral folds, and males without vocal slits and without nuptial pads. The new species is most similar to B. bustamantei but differs in being smaller, having discontinuous dorsolateral folds, the males lacking vocal slits, and an overall darker ventral coloration. Bryophryne contains three species all of which lack a tympanum. The deep valley of the Río Apurímac as a distributional barrier separating Phrynopus from Bryophryne is discussed.

Key words: Andes, biogeography, Bryophryne cophites, Bryophryne bustamantei

Lehr, E. & A. Catenazzi. 2008: A new species of Bryophryne (Anura: Strabomantidae) from southern Peru. Zootaxa. 1784: 1-10. http://www.mapress.com/zootaxa/2008/f/z01784p010f.pdf

http://www.ruffordsmallgrants.org/rsg/projects/alessandro_catenazzi_0

https://sites.google.com/site/acatenazzi/

[Herpetology • 2008] Terrarana (new taxon) | New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation

Abstract

New World frogs recently placed in a single, enormous family (Brachycephalidae) have direct development and reproduce on land, often far away from water. DNA sequences from mitochondrial and nuclear genes of 344 species were analyzed to estimate their relationships. The molecular phylogeny in turn was used as the basis for a revised classification of the group. The 882 described species are placed in a new taxon, Terrarana, and allocated to four families, four subfamilies, 24 genera, 11 subgenera, 33 species series, 56 species groups, and 11 species subgroups. Systematic accounts are provided for all taxa above the species level. Two families (Craugastoridae and Strabomantidae), three subfamilies (Holoadeninae, Phyzelaphryninae, and Strabomantinae), six genera (Bryophryne, Diasporus, Haddadus, Isodactylus, Lynchius, and Psychrophrynella), and two subgenera (Campbellius and Schwartzius) are proposed and named as new taxa, 13 subspecies are considered to be distinct species, and 613 new combinations are formed. Most of the 100 informal groups (species series, species groups, and species subgroups) are new or newly defined.  Brachycephalus and Ischnocnema are placed in Brachycephalidae, a relatively small clade restricted primarily to southeastern Brazil. Eleutherodactylidae includes two subfamilies, four genera, and five subgenera and is centered in the Caribbean region. Craugastoridae contains two genera and three subgenera and is distributed mainly in Middle America. Strabomantidae is distributed primarily in the Andes of northwestern South America and includes two subfamilies, 16 genera, and three subgenera. Images and distribution maps are presented for taxa above the species level and a complete list of species is provided. Aspects of the evolution, biogeography, and conservation of Terrarana are discussed. 

Key words: Amphibia, Brachycephalidae, Craugastoridae, DNA sequence, Eleutherodactylidae, evolution, Strabomantidae, systematics, taxonomy

Hedges, S.B., W.E. Duellman & M.P. Heinicke. 2008: New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737: 1-182.

http://www.mapress.com/zootaxa/2008/f/z01737p182f.pdf

http://species.wikimedia.org/wiki/Bryophryne

http://en.wikipedia.org/wiki/Bryophryne

[Herpetology • 2008] Bryophryne (Phrynopus) cophites • New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation

Bryophryne cophites (Lynch, 1975)

Distribution.— The genus occurs at elevations of 2900–4120 m in the Cordillera Oriental in the Departamento de Cusco in southern Peru (Fig. 93).

Etymology.— The generic name is derived from the Greek bryon meaning moss and the Greek phrynos, meaning toad. The name is feminine in gender and refers to a common habitat of these species.

Hedges, S.B., W.E. Duellman & M.P. Heinicke. 2008: New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa, 1737: 1-182.

http://www.mapress.com/zootaxa/2008/f/z01737p182f.pdf

http://species.wikimedia.org/wiki/Bryophryne

http://en.wikipedia.org/wiki/Bryophryne

[Herpetology • 2007] Bryophryne (Phrynopus) bustamantei • A new species of Phrynopus (Anura: Brachycephalidae) from Departamento Cusco, southern Peru

Bryophryne bustamantei 

(Chaparro, De la Riva, Padial, Ochoa & Lehr, 2007)

Abstract

We describe a new species of Phrynopus (Anura: Brachycephalidae) from two close localities at the upper limits of cloudforest in the southern Peruvian Departamento Cusco, between 3555–3950 m a.s.l. The new species is characterized byhaving medium size (maximum snout-vent length 23.4 mm), dentigerous processes of vomers absent, tympanic membrane inconspicuous, dorsal skin coarsely shagreen in life, dorsolateral folds, ventral skin areolate, dorsum tan, venterbold black with conspicuous bluish-gray spots, and a bluish-white iris.

Key words: Andes, Peru, Phrynopus, Anura

Chaparro, J. C., I. De la Riva, J. M. Padial, J. A. Ochoa, and E. Lehr. 2007. A new species of Phrynopus from Departamento Cusco, southern Peru (Anura: Brachycephalidae). Zootaxa. 1618: 61–68.

http://www.mapress.com/zootaxa/2007f/z01618p068f.pdf

http://calphotos.berkeley.edu/cgi/img_query?seq_num=247482&one=T

[Herpetology • 2008] Strabomantis aramunha • A large and enigmatic new eleutherodactyline frog (Anura, Strabomantidae) from Serra do Sincorá, Espinhaço range, Northeastern Brazil

Figure 1. Living specimens of Strabomantis aramunha from the state of Bahia, Brazil.

Abstract

We describe a new species of a large eleutherodactyline frog from the mountain rocky meadows (“campos rupestres”) of the Serra do Sincorá, Espinhaço mountain range, Mucugê municipality, State of Bahia, Brazil. The new species is promptly diagnosed from all the other Brazilian eleutherodactylines by its large size (males SVL 40.3–41.1; females SVL 75.2–79.7mm), broad head (head width 43–49% of SVL), presence of frontoparietal crests, pars fascialis of the maxilla deepened, discs absent on fingers, toes with poorly developed discs, first and second toes ridged, and tarsal fold absent. On the basis of these characters the new species is attributed to the genus Strabomantis up to now restricted to southern part of Central America and northwest part of South America. 

Key words: Strabomantis aramunha sp. nov., Eleutherodactylus, Strabomantidae, Amphibia, Atlantic forest, Campos rupestres, Taxonomy

Figure 2. Geographic distribution of Strabomantis aramunha on topographic map. Star, municipality of Mucugê, type locality; new records within the Chapada Diamantina Ecoregion, state of Bahia, Brazil: triangle, Miguel Calmon; lozenge, Iraquara; circle, Palmeiras; square, Lençóis; inverted triangle, Abaíra.

Figure 4. Strabomantis aramunha. (A) Dorsal and (B) ventral views of UFBA 6221, adult female, SVL 66.1 mm. (C) Dorsal and (D) ventral views of MZUEFS 2125, adult male, SVL 36.6 mm (photo by Marcelo F. Napoli)

Abstract

During surveys of anurans in the Chapada Diamantina, a semiarid region in central Bahia, Brazil, we obtained new records of Strabomantis aramunha Cassimiro, Verdade and Rodrigues. The new records extend the geographic distribution of S. aramunha in about 200 km north and 75 km south in relation to the type locality (municipality of Mucugê, state of Bahia, Brazil). We observed males calling at night at the Morro do Pai Inácio, municipality of Palmeiras. This new data is important, as in the original description the authors suggested the possibility of absence of advertisement call in the species. We also provide additions on color and morphometric data.

Cassimiro, J., V.K. Verdade and M.T. Rodrigues. 2008. A large and enigmatic new eleutherodactyline frog (Anura, Strabomantidae) from Serra do Sincorá, Espinhaço range, Northeastern Brazil. Zootaxa. 1761: 59-68. http://www.mapress.com/zootaxa/2008/f/z01761p068f.pdf

Napoli, M. F.; Juncá, F. A.; Cruz, D.; Abreu, R. O. 2010. Amphibia, Anura, Strabomantidae, Strabomantis aramunha Cassimiro, Verdade & Rodrigues, 2008: Distribution extension with notes on natural history, color patterns, and morphometric data. Check List. 6: 275–279.

http://dc249.4shared.com/doc/R5APkA1x/preview.html
http://www.amphibia.ufba.br/Public/Public_artigos_2008-2010.htm

[Ichthyology • 2012] Australoheros mattosi, A. montanus & A. tavaresi • Three new species of Australoheros (Labroidei: Cichlidae) from southeastern Brazil

 Australoheros montanus Ottoni, 2012

Abstract

Three new species of  Australoheros are described from the São Francisco, Paraná and Paraíba do Sul river basins, southeastern Brazil. Australoheros mattosi sp. n., from the rio São Francisco basin, by having anal-fin base squamation beginning at the third anal-fin spine; A. montanus sp. n., from the rio Paquequer drainage, by having a complete red bar on the posterior margin of the caudal fin; and, A. tavaresi sp. n., from the rio Tietê drainage, by having prognathous mouth. Three currently listed synonyms of Australoheros facetus are discussed: Chromys oblonga is considered a nomen dubium. The type specimens of Heros autochton (lectotype herein designated) indicate that the species does not belong to the genus Australoheros. Finally, the status of H. jenynsii as synonym of A. facetus is confirmed.

Key words : Biodiversity, Cichlinae, coastal basins, fishes, Heroini, Neotropical cichlids, systematics, taxonomy.

Australoheros mattosi Ottoni, 2012

Australoheros montanus Ottoni, 2012

Australoheros tavaresi Ottoni, 2012

Ottoni F. P. 2012. Three new species of Australoheros from southeastern Brazil, with taxonomic notes on Chromys oblonga, Heros autochton and H. jenynsii (Teleostei: Labroidei: Cichlidae). Vertebrate Zoology. 62(1): 83 – 96.

http://www.vertebrate-zoology.de/vz62-1/04_Vertebrate_Zoology_62-1_Ottoni_83-96.pdf

http://mahengechromis.blogspot.com/2012/04/mi-si-e-allargato-il-genere-tre-nuovi.html

[Ichthyology • 2011] Australoheros ykeregua & A. angiru • Two new species of Australoheros (Teleostei: Cichlidae), with notes on diversity of the genus and biogeography of the Río de la Plata basin

Abstract

Two new species of Australoheros Říčan and Kullander are described. Australoheros ykeregua sp. nov. is described from the tributaries of the río Uruguay in Misiones province, Argentina. Australoheros angiru sp. nov. is described from the tributaries of the upper rio Uruguai and middle rio Iguaçu in Brazil. The two new species are not closely related, A. ykeregua is the sister species of A. forquilha Říčan and Kullander, while A. angiru is the sister species of A. minuano Říčan and Kullander. The diversity of the genus Australoheros is reviewed using morphological and molecular phylogenetic analyses. These analyses suggest that the described species diversity of the genus in the coastal drainages of SE Brazil is overestimated and that many described species are best undestood as representing cases of intraspecific variation. The distribution patterns of Australoheros species in the Uruguay and Iguazú river drainages point to historical connections between today isolated river drainages (the lower río Iguazú with the arroyo Uruguaí, and the middle rio Iguaçu with the upper rio Uruguai). Molecular clocks are used to date these and other biogeographic patterns.

Key words: Australoheros, new species, Cichlidae, phylogeny, South America, biogeography, Brazilian shield

Australoheros ykeregua 

Etymology. The Guaraní word ykeregua means neighbor (vecino in Spanish). The etymology is based on the fact that A. ykeregua and A. forquilha have been preliminarily treated as conspecific (Říčan & Kullander 2008). New data have however demonstrated that they are two sister group species living in the same river drainage (río Uruguay), though not sympatrically.

Distribution. Australoheros ykeregua is so far known only from Argentinean territory in the tributaries of the río Uruguay below the Salto Moconá, province of Misiones.

Australoheros angiru

Etymology. The Guaraní word angirû means friend, partner (amigo or compañero in Spanish). The etymology is based on the fact that A. angiru and A. kaaygua have been confused as one species (Říčan & Kullander 2008). New data have however demonstrate that they are two non-sister group species living in the same river drainage (río Iguazú), though not sympatrically.

Distribution. Australoheros angiru has a disjunct distribution in the rio Iguaçu and in the upper rio Uruguai. One locality is so far known from the middle río Uruguay in Misiones province, Argentina (Fig. 10).

Two new species of Australoheros (Teleostei: Cichlidae), with notes on diversity of the genus and biogeography of the Río de la Plata basin

http://www.mapress.com/zootaxa/2011/f/zt02982p026.pdf

http://www.cichlidae.com/forum/viewtopic.php?f=17&t=12702

[Ichthyology • 2011] Australoheros perdi • new species (Teleostei: Labroidei: Cichlidae) from the lacustrine region of the Doce River Valley, southeastern Brazil, with biological information

Australoheros perdi 2011

Abstract

Australoheros perdi is a new species herein described from the lacustrine region of the middle Doce River basin, Doce River Valley, southeastern Brazil. The new species is distinguished from all its congeners by having only 25 vertebrae and by a combination of characters states listed below: three abdominal bars in all stages of life, a conspicuous, rounded caudal-fin base spot, a conspicuous wide longitudinal stripe, head with depression in the region above the eyes, tip of pelvic fin reaching vertical through third to sixth anal fin spine base, fewer dorsal-fin spines, fewer pectoral-fi n rays, fewer caudal vertebrae, fewer proximal radials on dorsal-fin base, more proximal radials on anal-fi n base, more pleural ribs, a longer caudal peduncle, a deeper caudal peduncle, a wide ectopterygoid, a longer last anal-fi n spine and a longer lower jaw. The available phylogenetic tree of the genus does not include the species from southeastern Brazil, thus impeding any discussion of the phylogeny of these species. 

Key words: Cichlinae, Heroini, middle Doce River basin, Minas Gerais state, Parque Estadual do Rio Doce, Pisces, South-American cichlids, Taxonomy, biology, diet.

Ottoni, F.P., Lezama, A.Q., Triques, M.L., Fragoso-Moura, E.N., Lucas, C.C.T., and Barbosa, F.A.R. 2011. Australoheros perdi, new species (Teleostei: Labroidei: Cichlidae) from the lacustrine region of the Doce River Valley, southeastern Brazil, with biological information. Vertebrate Zoology, 61 (1): 137-145 .

http://www.vertebrate-zoology.de/vz61-1/09_Vertebrate_Zoology_61-1_Ottoni.pdf

[Ichthyology • 2010] Australoheros capixaba • a new species of Australoheros from south-eastern Brazil

Australoheros capixaba Ottoni, 2010

Abstract
Australoheros capixaba, new species, is distributed along the rio Itaúnas, rio Barra Seca, rio São Mateus and the lower rio Doce basins. The new species is distinguished from its congeners in the rio Paraná-Uruguay system and Laguna dos Patos system by having 12 caudal vertebrae (vs. 13 – 15). Australoheros capixaba differs from the other species of Australoheros from south-eastern Brazil by its coloration in life (a reddish chest, large spots on the dorsal region of the trunk and a green iridescence on the pelvic fins). It also differs from some of its congeners by having a longer caudal peduncle, a longer anal-fin spine, fewer dorsal-fin spines and fewer anal-fi n rays. Australoheros capixaba sp. n. is the fi rst species from the genus described for the Estado do Espírito Santo, south-eastern Brazil. The phylogenetic placement of the species in the genus cannot be discussed, because there is no phylogenetic work about the Australoheros species from south-eastern Brazil.

Key words: Australoheros facetus, coastal basins, Estado do Espírito Santo, Heroini, Linhares, rio Barra Seca, rio Doce, rio Itaúnas, rio São Mateus, South American cichlids, taxonomy.

Etymology. From the Brazilian idiomatic expression capixaba, meaning people who were born in the Estado de Espírito Santo.

Distribution. Rio São Mateus basin, rio Itaúnas basin, rio Barra Seca basin and lower rio Doce basin, all in south-eastern Brazil (Fig. 2).

Ottoni, F.P., 2010. Australoheros capixaba, a new species of Australoheros from south-eastern Brazil (Labroidei: Cichlidae: Cichlasomatinae). Vertebrate Zoology. 60 (1): 19 – 25

http://www.vertebrate-zoology.de/vz60-1/03_Vertebrate_Zoology_60-1_Ottoni.pdf

[Ichthyology • 2009] Australoheros taura • A new species of Australoheros from the upper Rio das Antas, Laguna dos Patos system, southern Brazil

Australoheros taura 

Ottoni & Cheffe, 2009

Ottoni, Felipe & M. M. Cheffe. 2009. A new species of Australoheros from the upper Rio das Antas, Laguna dos Patos system, southern Brazil. Spixiana. 32 (1): 153-159.

http://www.docstoc.com/docs/44577237/A-new-species-of-Australoheros-from-the-upper-rio#

http://flickr.com/photos/26042632@N03/4040583352/

[Ichthyology • 2008] Australoheros ribeirae • A new species of the genus Australoheros from the rio Ribeira do Iguape basin, So Paulo, Brazil

Australoheros ribeirae Ottoni, Oyakawa & Costa, 2008

Brazilian scientists have described a new species of cichlid 

in the genus Australoheros from southeastern Brazil.

The description of Australoheros ribeirae by Felipe Ottoni, Osvaldo Oyakawa and Wilson Costa is published in the most recent issue of the journal Vertebrate Zoology.

Australoheros ribeirae is distinguished from other members of the genus in having a combination of the following characters: 9"10 dorsal-fin rays, 6"7 anal-fin spines, 8 anal-fin rays, 14 precaudal vertebrae, 12 caudal vertebrae, 26 total vertebrae, 24"26 scales in longitudinal series, 16"18 scales in the superior lateral line, head depth 95.4"98.9 % head length, last dorsal-fin spine length 16"16.8 % standard length, body depth 47.4"51.3 % standard length, pelvic fin length 32.4"37.2 % standard length, predorsal length 47.1"49.6 % standard length, preorbital depth 64.2"73.3 % head length, head width 48.9"53.6 % head length, narrow ectopterygoid narrow, truncate process of the second vertebra, two long tubular processes on the second epibranchial, long anterior arm of the first epibranchial, trunk bars never ventrally forked, fifth trunk bar dorsally straight, straight connection above upper lateral line between the fifth and sixth trunk bars, dorsal portion of the second to fourth trunk bars inclined posteriorly, slightly curved first trunk bar lacking an expanded anterior margin, opercle without spots and usually red sides of the body usually red.

The new species is known from the Ribeira do Iguape drainage in southeastern Brazil, after which it is named.

Ottoni, FP, OT Oyakawa and WJEM Costa. 2008. A new species of the genus Australoheros from the rio Ribeira do Iguape basin, So Paulo, Brazil (Labroidei: Cichlidae: Cichlasomatinae). Vertebrate Zoology. 58: 75 - 81.

http://www.vertebrate-zoology.de/vz58-1/58_1-10_Ottoni.pdf

http://www.practicalfishkeeping.co.uk/content.php?sid=1660

[Ichthyology • 2008] Australoheros forquilha, A. charrua, A. minuano & A. guarani • The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages

Australoheros forquilha Rícan & Kullander, 2008

http://www.pecescriollos.de/go/australoheros-forquilha

Australoheros charrua Rícan & Kullander, 2008 

Australoheros forquilha Rícan & Kullander, 2008 

Australoheros guarani Rícan & Kullander, 2008 

Australoheros minuano Rícan & Kullander, 2008 

Abstract

Australoheros Říčan and Kullander, 2006 includes four described species 

— Australoheros facetus (Jenyns, 1842), Australoheros tembe (Casciotta, Gómez & Toresani, 1995), Australoheros scitulus (Říčan & Kullander, 2003) and Australoheros kaaygua Casciotta, Almirón & Gómez, 2006. 

Four additional species are newly described in this paper based on results presented in Říčan and Kullander (2006): Australoheros forquilha sp. nov., from the tributaries of the Upper Río Uruguay in Brazil; Australoheros charrua sp. nov. and Australoheros minuano sp. nov., from the tributaries of the Middle Río Uruguay in Brazil and Uruguay, and Australoheros guarani sp. nov. from the Río Paraná tributaries in Paraguay. Sympatric species are not found closely related and all sister species have allopatric distributions. 

Four species groups are recognized based on phylogenetic hypotheses generated from morphological and cyt b characters; the forquilha group—A. forquilha, A. tembe; the scitulus group—A. scitulus, A. charrua; the kaaygua group—A. kaaygua, A. minuano; and the facetus group—A. facetus, A. guarani.

Key words: ´Cichlasoma´ facetum group, Australoheros, new species, Cichlidae, South America, biogeography, Río Uruguay, Río Paraná

Rícan, O. & Kullander, S.O. (2008) The Australoheros (Teleostei: Cichlidae) species of the Uruguay and Parana River drainages. Zootaxa, 1724: 1-51. 

http://www.mapress.com/zootaxa/2008/f/z01724p051f.pdf

http://www.practicalfishkeeping.co.uk/content.php?sid=1585

[Ichthyology • 2008] Taxonomic revision of the genus Australoheros Rícan & Kullander, 2006 (Teleostei: Cichlidae) with descriptions of nine new species from southeastern Brazil

Australoheros autrani Ottoni & Costa, 2008

Australoheros barbosae Ottoni & Costa, 2008

Australoheros ipatinguensis Ottoni & Costa, 2008

Australoheros macacuensis Ottoni & Costa, 2008

Australoheros macaensis Ottoni & Costa, 2008

Australoheros muriae Ottoni & Costa, 2008

Australoheros paraibae Ottoni & Costa, 2008

Australoheros robustus Ottoni & Costa, 2008

Australoheros saquarema Ottoni & Costa, 2008

Ottoni, F.P. and W.J.E.M. Costa. 2008. Taxonomic revision of the genus Australoheros Rcan & Kullander, 2006 (Teleostei: Cichlidae) with descriptions of nine new species from southeastern Brazil. Vertebrate Zoology. 58:207 - 232.

http://www.vertebrate-zoology.de/vz58-2/05_Ottoni.pdf