Saturday, February 28, 2026

[Botany • 2026] Nepenthes siamensis (Nepenthaceae) • A New Species of Section Montanae from Surat Thani Province, Peninsular Thailand

 

Nepenthes siamensis Nuanlaong 

in Nuanlaong, Yodthammarat, Chiwan, Laewan et Pohsap, 2026.
หม้อแกงสยาม  ||  DOI: doi.org/10.1007/s12225-025-10336-2
Drawn by Chayan Yodthammarat
 
Summary
Nepenthes siamensis, a new species from Khlong Phanom National Park in southern Thailand, is described. This species, closely related to others in section Montanae, stands out due to its distinct inflorescence characteristics, including bracts present on both male and female flowers, and unique pitcher morphology. It is found approximately 650 km north of the most similar species based on morphological comparison, N. sanguinea and N. thai, which are also both terrestrial and lower montane. It shares characters with them, such as a climbing habit and oblong to subspathulate leaves with 2 – 3 longitudinal nerves on each side of the midrib. This discovery significantly extends the known range of section Montanae.

 Key Words: Carnivorous plants, pitcher plant, taxonomy

Nepenthes siamensis.
A stem with leaves and upper pitchers; B lower pitcher; C male inflorescence; D female flowers; E male flowers; F lower surface of lid with upper part of peristome; G upper surface of lid; H – J nectar glands.
From live specimens. Drawn by Chayan Yodthammarat



Nepenthes siamensis Nuanlaong sp. nov. 
Type: Thailand, Surat Thani Province, Phanom Distr., Khlong Phanom National Park, ...

Recognition. This species differs from Nepenthes thai in leaf base margins clasping the stem for ½ of the circumference, not decurrent (vs clasping stem ¾ – ⅚ of circumference, bases decurrent down the stem for 3 – 12 mm in N. thai); lower pitcher cylindrical in lower ¾ of length, infundibular in upper ¼ (vs ovoid-cylindrical); spur filiform (vs branched); bracts present on all flowers at the bases of pedicels in the male inflorescence and on the lower half of the pedicel in the female inflorescence (vs bract absent).




  Sunya Nuanlaong, Chayan Yodthammarat, Purin Chiwan, Teerayut Laewan and Suphansiri Pohsap. 2026. Nepenthes siamensis (Nepenthaceae), A New Species of Section Montanae from Surat Thani Province, Peninsular Thailand. Kew Bulletin. DOI: doi.org/10.1007/s12225-025-10336-2 [27 February 2026]


[Botany • 2025] Zeuxine drukyulensis (Orchidaceae) • A New jewel orchid from Bhutan Himalaya

 

Zeuxine drukyulensis P.Gyeltshen, Rabgay & Kumar,

in Gyeltshen, Rabgay, Tobgay et Kumar, 2025. 
 
Abstract
An interesting new species of jewel orchid, Zeuxine drukyulensis is described and illustrated from warm broadleaved forests in western Bhutan. It is closely allied to Zeuxine nervosa and Zeuxine rolfeana, however, differs in various morphological characters, primarily the structure, size and colour of the flower. The new species is assessed as Data Deficient (DD) as per the IUCN Red List guidelines.

Conservation, Data Deficient, Drukyul, Orchidoideae, Rimchu, taxonomy, Monocots




Phub GYELTSHEN, Kinley RABGAY, Kezang TOBGAY and Pankaj KUMAR. 2025. Zeuxine drukyulensis (Orchidaceae), A New jewel orchid from Bhutan Himalaya.  Phytotaxa. 690(1); 123-129. DOI: doi.org/10.11646/phytotaxa.690.1.11 [2025-02-26]
 facebook.com/PGyeltshen2022


[Herpetology • 2026] Dibamus irregularis • A New Species of Blind Skink (Squamata: Dibamidae: Dibamus) from Chu Yang Sin National Park, southern Vietnam, with the first description of a Dibamus clutch

 

Dibamus irregularis Kliukin, Poyarkov, Bragin, Krone & Nguyen, 

in Kliukin, Bragin, Nguyen, Vu, Le, Gorin, Krone & Poyarkov, 2026
Chu Yang Sin Blind Skink  |  DOI: doi.org/10.11646/zootaxa.5760.5.1  
Thằn lằn giun Chư Yang Sin  ||  Researchgate.net/publication/401227726  

Abstract
A new species of dibamid lizard, Dibamus irregularis sp. nov. is described based on molecular (three mitochondrial genes) and morphological (external and skeletal) characters of twelve specimens collected from the Chu Yang Sin National Park, Dak Lak Province, southern Vietnam. The new species can be distinguished from other congeners by the following combination of characters: medial rostral suture absent or incomplete, nasal sutures incomplete, labial sutures complete or incomplete; two to three postoculars; three to four scales on the posterior edge of infralabial; 22–24 midbody scale rows; 180–211 ventral scales; 39–46 subcaudal scales; and maximum snout-vent length 147 mm. Moreover, two eggs of a new species were collected in the same locality. The only known clutch of Dibamus was described more than 120 years ago based on a single egg of D. alfredi collected from Bukit Besar, Thailand. Herein, we provide the first description of a complete clutch of the genus Dibamus based on two eggs of Dibamus irregularis sp. nov. Our study brings the total number of species in the genus Dibamus to 30; Dibamus irregularis sp. nov. is the 10th species of Dibamus recorded from Vietnam, highlighting the importance of the country as a hot spot of reptilian diversity in Southeast Asia.

Reptilia, Dibamus irregularis sp. nov., Dak Lak Province, conservation, Southeast Asia, systematics, morphology, reproduction

 Dibamus irregularis sp. nov. in life.
(A) holotype ZMMu re-18313, male; (B) Paratype ZMMu re-18315, female.
Photographs by Andrey M. Bragin




Dibamus irregularis sp. nov.
Kliukin, Poyarkov, Bragin, Krone & Nguyen

Etymology. the new species epithet is a latin adjective in nominative singular, meaning “irregular,” andis given in reference to the high intraspecific variability of external morphological features in the new species,especially to the irregularity and asymmetry of the number of postocular scales (Fig. 6). Since Allen greer’s revision(greer 1985) and in all subsequent papers describing new species of Dibamus, the number of postocular scales wasconsidered as a relatively stable diagnostic character underlying diagnoses for new species of dibamids, which in partcould be explained by generally small sample sizes of Dibamus spp. available for examination. however, Dibamus irregularis sp. nov. demonstrates an unexpectedly irregular and asymmetrical number of postocular scales: fivespecimens have two postoculars, four specimens have three postoculars, and three specimens have two postocularson the left side and three on the right side of the head.

Suggested common names: Chu Yang Sin Blind Skink (English); Thằn lằn giun Chư Yang Sin (Vietnamese),Chuyangsinskaya cherveobraznaya yascheritsa (Чуянгсинская червеобразная ящерица, Russian).  



Nikita S. KLIUKIN, Andrey M. BRAGIN, Tan Van NGUYEN, Duy Dinh VU, Son Xuan LE, Vladislav A. GORIN, Isaac W. KRONE, Nikolay A. POYARKOV. 2026. A New Species of Blind Skink (Squamata: Dibamidae: Dibamus) from Chu Yang Sin National Park, southern Vietnam, with the first description of a Dibamus clutch.  Zootaxa. 5760(5); 501-528. DOI: doi.org/10.11646/zootaxa.5760.5.1 [2026-02-26] 

[Botany • 2025] Tessmannia princeps (Fabaceae: Detarieae) • A New rainforest Tree from the Udzungwa Mountains, Tanzania


Tessmannia princeps A.Bianchi, Tomasi, Mwakisoma, Barbieri & Q.Luke, 

in BianchiTomasiMwakisomaBarbieri et Luke, 2025. 
 
Abstract
Tessmannia princeps, a new canopy emergent rainforest tree species from the Udzungwa Mountains, Tanzania, is described and illustrated. It is a montane, unarmed tree distinguished by its large stature, extensively buttressed bole, and high number of leaflets. Tessmannia princeps shows morphological affinities with another Tanzanian species, T. densiflora, nevertheless the two species can be distinguished by morphological characters as well as by ecology and geography. A diagnostic key to the East African species of Tessmannia is provided. Due to its restricted range and small number of recorded mature individuals, the conservation status of T. princeps is assessed as Vulnerable under the IUCN criteria.

Eastern Arc, Fabaceae, Detarieae, new species, taxonomy, Eudicots 




Tessmannia princeps A.Bianchi, Tomasi, Mwakisoma, Barbieri & Q.Luke, sp. nov. 

Diagnosis:—Superficially comparable to Tessmannia densiflora, but Tessmannia princeps is a taller tree, growing up to 40 m in height; leaves bear 18–24 pairs of leaflets on a rachis 68–94 mm long; leaflets are opposite to alternate, sub-rectangular or oblong in shape; flowers are white, with sepals and tepals up to 5 mm in length. Tessmannia densiflora is a smaller tree, reaching 20–25 m in height; leaves bear 8–13 pairs of leaflets on a rachis 30–60 mm long; leaflets are sub-opposite to alternate and may be oblong, lanceolate, ovate-oblong, or ovate-lanceolate in shape; flowers are red, with sepals and tepals measuring 10–20 mm in length.


ANDREA BIANCHI, LAURA TOMASI, ALOYCE MWAKISOMA, MATTEO BARBIERI and QUENTIN LUKE. 2025. Tessmannia princeps (Fabaceae), A New rainforest Tree from the Udzungwa Mountains, Tanzania.  Phytotaxa. 694(2); 109-118. DOI: doi.org/10.11646/phytotaxa.694.2.1 [2025-03-20]
https://news.mongabay.com/2025/03/3000-year-old-trees-in-tanzania-are-new-species/

Friday, February 27, 2026

[Paleontology • 2026] The ontogenetically youngest known pachycephalosaur (Dinosauria: Ornithischia) postcranium

 

juvenile Pachycephalosauria indet. (CMNFV 22039)
Life reconstruction in an environment typical of the upper Maastrichtian Frenchman Formation, Saskatchewan. 

in Moore, Evans, Ryan, Patterson et Mallon, 2026. 
Illustration by Kaitlin Lindblad. 
 
ABSTRACT
Pachycephalosaurian dinosaurs are represented in the fossil record primarily by their taphonomically resistant frontoparietal domes that developed fully only at maturity. Consequently, the postcranial record of Pachycephalosauria is poor, particularly for immature forms. Here, we describe the ontogenetically youngest known, and Canada’s second-most complete, pachycephalosaur postcranium, collected from the uppermost Maastrichtian Frenchman Formation, Saskatchewan. Despite its small size (estimated total length ∼90 cm), the skeleton shows several characters diagnostic of Pachycephalosauria, including a double ridge-and-groove articulation on the pre- and postzygapophyses of the dorsal neural arches, a medial flange on the postacetabular process of the ilium, and a highly reduced pubis that contributes only minimally to the acetabular margin. Histological sectioning of the crural bones shows an immature woven bone texture. The absence of cyclical or annual growth indicators suggests that this specimen was less than a year old at the time of death. Phylogenetic analysis recovers the specimen as a member of Pachycephalosaurinae, perhaps related to Prenocephale, but the lack of cranial data for our specimen and the complementary lack of postcranial data for other pachycephalosaurs make this determination tenuous. Spatiotemporal considerations suggest a possible but currently untestable affiliation with Sphaerotholus buchholtzae. The relatively long hindlimbs of the juvenile compared with those of adult pachycephalosaurs indicate probable negative ontogenetic allometry in the hindlimbs.

 Preserved elements (in red) of juvenile Pachycephalosauria indet. (CMNFV 22039).
 The preserved left ischium does not show in this view. Diagram illustrates preserved elements only and does not accurately reflect the proportions of CMNFV 22039.
Skeletal drawing modified after original illustrations of Pachycephalosaurus (postcranium) and “Dracorex” (cranium) by Gregory Paul. Used with permission.

Life reconstruction of CMNFV 22039 in an environment typical of the upper Maastrichtian Frenchman Formation. 
Illustration by Kaitlin Lindblad. Used with permission.
 

Bryan R. S. Moore, David C. Evans, Michael J. Ryan, R. Timothy Patterson and Jordan C. Mallon. 2026. The ontogenetically youngest known pachycephalosaur (Dinosauria: Ornithischia) postcranium. Journal of Vertebrate Paleontology. e2616325. DOI: 10.1080/02724634.2026.2616325 [26 Feb 2026]

[Botany • 2025] Microchirita baolamensis (Gesneriaceae) • A New Species from Vietnam

 

Microchirita baolamensis T.S. Hoang, W.G. Wang & H.C. Xi, 

in Hoang, Xi, Shen et Wang, 2025. 
 
Abstract 
Microchirita baolamensis T.S. Hoang, W.G. Wang & H.C. Xi (Gesneriaceae) is described and illustrated as a new species from Vietnam. It resembles M. involucrata and M. rupestris, but differs from them by its cristate inflorescence, purple corolla tube with yellow stripes and eglandular pubescence, and free anthers. It further differs from M. involucrata by a partly pubescent ovary and an eglandularly pubescent style. Based on IUCN categories and criteria, the species is assessed as Critically Endangered (CR).


Microchirita baolamensis T.S. Hoang, W.G. Wang & H.C. Xi sp. nov.
 

Son Thanh Hoang, Hou-Cheng Xi, Jian-Yong Shen and Wen-Guang Wang. 2025. Microchirita baolamensis (Gesneriaceae), a New Species from Vietnam. Annales Botanici Fennici. 62(1):135-138. DOI: doi.org/10.5735/085.062.0123  [9 May 2025]


Thursday, February 26, 2026

[Herpetology • 2026] Cyrtodactylus kiriromensis • A New Species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia

 
Cyrtodactylus kiriromensis
Grismer, Neang, Samorn, Song & Stuart, 2026


Abstract
A new species of the Cyrtodactylus intermedius group, Cyrtodactylus kiriromensis sp. nov., from Kirirom National Park of Phnom Sruoch in the Cardamom Mountains of Cambodia is delimited based on a mitochondrial (ND2) phylogeny and statistically diagnosed using meristic, morphometric, and color pattern characters. Cyrtodacytylus kiriromensis sp. nov. is the sister species of C. septimontium of southern Vietnam and forms part of a clade with six other site-specific endemic species from the mountains and islands of southeastern Cambodia and southwestern Vietnam. The ongoing discoveries of localized endemic species in the Cardamom Mountains underscores their fragmented nature and the importance of this landscape as a source of speciation as well as its notable contribution to the herpetological diversity of Cambodia. As such, it should be given special consideration as a conservation priority.

Reptilia, Indochina, Southeast Asia, integrative taxonomy, gecko, endemism, conservation


 Cyrtodactylus kiriromensis sp. nov.



L. Lee GRISMER, Thy NEANG, Vireak SAMORN, Det SONG and Bryan L. STUART. 2026. A New Species of the Cyrtodactylus intermedius group (Squamata: Gekkonidae) continues to underscore the high degree of site-specific endemism in the Cardamom Region, Cambodia.  Zootaxa. 5741(3); 519-538. DOI: doi.org/10.11646/zootaxa.5741.3.5 [2026-01-07]


[Botany • 2025] Peliosanthes chenmingsongii (Asparagaceae) • A New Species from Yunnan, Southwestern China

 

Peliosanthes chenmingsongii H.Z. Feng,

in Li, Pan, Sun, Yuan et Feng, 2025. 
 
Abstract
Peliosanthes chenmingsongii is described and illustrated as a new species from southwestern China. The new species is morphologically closest to P. cupuliformis, but differs in its erect, woody and hard rhizome, longer petioles and larger leaf blades, more widely expanded flowers with recurved perianth segments, non-revolute margins and a horizontally inward-growing annular appendage in the corona. Careful comparisons with other closely related species are also made.

pistil, corona, Liliaceae, Peliosanthes cupuliformis, taxonomy



Peliosanthes chenmingsongii H.Z. Feng, sp. nov.


Xiao Hong LI, Xiang Ru PAN, Yi Fei SUN, Juan Juan YUAN and Hui Zhe FENG. 2025. Peliosanthes chenmingsongii (Asparagaceae), A New Species from Yunnan, Southwestern China.  Phytotaxa. 732(3); 295-300. DOI: doi.org/10.11646/phytotaxa.732.3.7 [2025-12-05] 

[Botany • 2024] Peliosanthes yangchunensis (Asparagaceae) • A New Species from Guangdong, China

 

Peliosanthes yangchunensis  F.Z. Feng & W.B Liao,
 
in Feng, Wu, Zeng, Fan et Liao, 2024. 
 
Abstract
A new species named Peliosanthes yangchunensis are described, which is endemic to Yangchun, western Guangdong, South China. The new species is closely related to P. macrostegia, but differs from the latter in that the flowers smaller, star-like and with strongly revolute, spreading and narrowly oblong tepals, style conical, ovary hexagonal in cross section, and fruit ovoid-globose, flower period September to October. Comparisons with other closely related species are also provided.

Asparagaceae, Peliosanthes, South China, western Guangdong, Monocots

Peliosanthes yangchunensis.
A. Habit; B & D. Inflorescence; C. Inflorescence & fruits; E & F. Flower, front view; G & H. Flower, views from side; I, J, K & L. Flower, longitudinally sectioned; L. Corona with three anthers; N. Corona, viewed from below; O. Basal portion of flower, with corona cross-sectioned; P. Basal portion of flower except corona and pistil; Q. Basal portion of flower, corona and ovary cross-sectioned; R. Flower, views from side and with corona removed. The the left photo in D and F were taken by Mr. Meng De Chang, and others photos, correction and design by H. Z. Feng.

Peliosanthes yangchunensis F.Z. Feng & W.B Liao, sp. nov.


Hui-Zhe FENG, Rang-Min WU, Qiu-Gen ZENG, Qiang FAN and Wen-Bo LIAO. 2024. Peliosanthes yangchunensis (Asparagaceae), A New Species from Guangdong, China.  Phytotaxa. 650(3); 262-268. DOI: doi.org/10.11646/phytotaxa.650.3.6 [2024-05-29] 

[Botany • 2025] Peliosanthes purpureiflora (Asparagaceae) • A New Species from South China


Peliosanthes purpureiflora H.Z. Feng,  

in K. K Meng, QU, Li, Lu, T. Meng, Wang et Feng, 2025. 
紫花球子草  ||  DOI: doi.org/10.11646/phytotaxa.690.1.10 
 
Abstract
A new species, Peliosanthes purpureiflora, endemic to South China, has been described. The new species exhibits several distinguishing characters including the terete pedicel, perianth segments spreading in the same plane, the truncate-conical corona with a 6-subdentate distal margin, the smaller ovary with a noticeable gap between the corona, and a longer style, approximately 1.2 mm in length, with three wing-like projections.

Asparagaceae, Peliosanthes, South China, corona, taxonomy, Monocots

Peliosanthes purpureiflora.
A. Habitat. B, C, D & E. Inflorescences. B. Young inflorescence. C, D & E. Old inflorescences. F. Young flower, front view. G. Old flower, front view. H. Old flower, dorsal view. I. Young flower, front view with corona and ovary cross-sectioned. J & L. Flower, views from side and corona removed. K & M. Flower, longitudinally sectioned. N. Old flower, front view with corona and ovary cross-sectioned. O. Flower, front view and corona removed. P. Adaxial leaf surface. Q. Abaxial leaf surface.
Photos, photo correction and design by H.Z. Feng.

Peliosanthes purpureiflora H.Z. Feng, sp. nov.


Kai Kai MENG, Jun Ru QU, Ren Kun LI, Li Yan LU, Tao MENG, Li Ping WANG and Hui Zhe FENG. 2025. Peliosanthes purpureiflora (Asparagaceae), A New Species from China.  Phytotaxa. 690(1); 116-122. DOI: doi.org/10.11646/phytotaxa.690.1.10 [2025-02-26] 

[Paleontology • 2026] Haolong dongi • Cellular-level Preservation of cutaneous spikes in an Early Cretaceous iguanodontian Dinosaur


 Haolong dongi 
Huang, Wu, Mao, Bertozzo, Dhouailly, Robin, Pittman, Kaye, Manucci, He, Wang & Godefroit, 2026  


Abstract
The near-complete and articulated skeleton of a new iguanodontian dinosaurHaolong dongi gen. et sp. nov., from the Lower Cretaceous of northeastern China, preserves exquisitely fossilized skin. The integument includes large overlapping scutate scales along the tail and tuberculate scales around the neck and thorax markedly different from the scale pattern described in other iguanodontians. Remarkably, these scales are interspersed with cutaneous spikes preserved at the cellular level. Tomographic and histological analyses reveal a hollow, cylindrical structure composed of a cornified stratum corneum overlying a pluristratified epidermis with keratinocytes preserved to the level of nuclei, surrounding a porous central dermal pulp. These spikes differ structurally from known protofeathers in non-avian dinosaurs and scaly spines in extant squamates, suggesting a distinct evolutionary origin. Their morphology and distribution imply a primary role in predator deterrence, with potential secondary functions in thermoregulation or mechanoreception. This discovery provides unprecedented insight into the microanatomy of non-avian dinosaur skin and highlights the complexity of skin evolution in ornithischian dinosaurs.





 Haolong dongi gen. et sp. nov.



Jiandong Huang, Wenhao Wu, Lei Mao, Filippo Bertozzo, Danielle Dhouailly, Ninon Robin, Michael Pittman, Thomas G. Kaye, Fabio Manucci, Xuezhi He, Xuri Wang and Pascal Godefroit. 2026.  Cellular-level Preservation of cutaneous spikes in an Early Cretaceous iguanodontian Dinosaur. Nature Ecology & Evolution. 10; 203–210. DOI: doi.org/10.1038/s41559-025-02960-9 [06 February 2026]


Wednesday, February 25, 2026

[Crustacea • 2026] Alpheus madhusoodanai • A New Species of Mangrove associated Snapping Shrimp of Genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from southwest coast of India


Alpheus madhusoodanai  
Vishnu, Deepak, Nidhin, Krishna & Harikrishnan, 2026 

 
Abstract
The present account describes a new species of alpheid shrimp, Alpheus madhusoodanai sp. nov., belonging to the brevirostris group, collected from the Cochin estuary, the south west coast of India. This represents the first species of alpheid shrimps described from the estuary. The morphological and molecular characteristics of the new species are compared with those of its closely related congeners. The newly described species is separated from its morphological congener A. rapax, by its wider major chela and longer merus of the first cheliped. Molecular data also confirmed the delimitation of A. rapax with A. madhusoodanai sp. nov. Habitat and distribution details are also discussed, highlighting the potential for further taxonomic exploration in the Cochin estuary and the importance in uncovering its hidden biodiversity.

Keywords: alpheid shrimp, estuary, integrative taxonomy, new species, south east Arabian Sea


Alpheus madhusoodanai sp. nov. Colour pattern, male, non-type, recently deceased (Cl- 10.9 mm, SIF/HK/CR/25/122), Cochin estuary, Kochi, Kerala, India.

 Alpheus madhusoodanai sp. nov.

Etymology: The new species is named after Dr. B. Madhusoodhana Kurup, for his significant contributions to fish biodiversity, fisheries taxonomy and ecology, and sustainable management of fisheries resources in the state of Kerala, south India.


Kaimakulangara Vishnu, Jose Deepak, Balachandran Nidhin, Unnikrishnan Abhai Krishna, Mahadevan Harikrishnan. 2026. Alpheus madhusoodanai sp. nov.: A New Species of Mangrove associated Snapping Shrimp of Genus Alpheus Fabricius, 1798 (Decapoda: Caridea: Alpheidae) from southwest coast of India. Journal of the Marine Biological Association of the United Kingdom. 106; e12. DOI: doi.org/10.1017/S0025315426101052 [26 January 2026]
 

[Botany • 2026] Medinilla calanasan (Melastomataceae) • A New Species from Apayao Province, Philippines

 
Medinilla calanasan  

in Pelser, Dela Cruz, Suetos, Ibañez et Barcelona, 2026. 

Abstract
Medinilla calanasan is here described as a new species from Calanasan in Apayao Province in northern Luzon (Philippines). It is morphologically distinct from all other known Medinilla species in having horn-like protrusions at the base of the leaf lamina. Morphological similarities with M. clementis Merr. and M. dolichophylla Merr. suggest that M. calanasan may be a member of the M. magnifica-alliance.

 Luzon, Medinilla magnifica-alliance, Myrtales, taxonomy, Eudicots



Medinilla calanasan sp. nov.

Pieter B. PELSER, Breiden DELA CRUZ, Kristian J. E. SUETOS, Jayson C. IBAÑEZ and Julie F. BARCELONA. 2026. Medinilla calanasan (Melastomataceae), A New Species from Apayao Province, Philippines.  Phytotaxa. 742(1); 99-104. DOI: doi.org/10.11646/phytotaxa.742.1.7 [2026-02-18]
https://www.philippineeaglefoundation.org/post/new-horn-leaf-medinilla-discovered-inside-apayao-eagle-forest