[Diplopoda • 2026] Enghoffosoma chiangmaiense, ER. minutum, E. chanabunae • Integrative Taxonomy of the Millipede Genus Enghoffosoma Golovatch, 1993 (Polydesmida: Paradoxosomatidae) in Thailand and Myanmar, with Descriptions of Three New Species

Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn; 
E. minutum Srisonchai & Likhitrakarn; 
 E. chanabunae Srisonchai & Likhitrakarn, 
 

 in Srisonchai, Benchapong, Golovatch, Sutcharit, Panha et Likhitrakarn, 2026. 

DOI: doi.org/10.37828/em.2026.99.4 

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Abstract

Although comprehensive studies on the Southeast Asian flat-backed millipede genus Enghoffosoma Golovatch, 1993 in Thailand have begun quite recently, in 2014, continuous field surveys have since revealed a steadily increasing species diversity. Based on extensive new material collected from Thailand and Myanmar, three new species are described using an integrative taxonomic approach that combines morphological characters and mitochondrial COI DNA barcoding: Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn sp. n. from northern Thailand, Enghoffosoma minutum Srisonchai & Likhitrakarn sp. n. from northeastern Thailand, and Enghoffosoma chanabunae Srisonchai & Likhitrakarn sp. n. from the Tanintharyi Region, southern Myanmar. COI sequence comparisons, available for E. chanabunae Srisonchai & Likhitrakarn sp. n. and E. chiangmaiense Srisonchai & Likhitrakarn sp. n., reveal significant interspecific genetic divergences (K2P distances) from their congeners, ranging from 13.53–16.75% and 13.53–18.14%, respectively. While DNA data were unavailable for E. minutum Srisonchai & Likhitrakarn sp. n., its distinctive morphological characters, particularly the gonopod structure, clearly support its status as a new species. To provide a clear morphological baseline for comparison, the type species, Enghoffosoma spinipleurum (Carl, 1941), from Myanmar, originally described as Sundanina spinipleura Carl, 1941, is redescribed and illustrated in detail based on the lectotype. With these additions, the genus Enghoffosoma now comprises 21 valid species, confirming its status as a highly diverse and widespread diplopod lineage in the Indochinese region. An updated identification key to all known species and a distribution map are provided.

Enghoffosoma chiangmaiense Srisonchai & Likhitrakarn sp. n. 

 Enghoffosoma minutum Srisonchai & Likhitrakarn sp. n. 

 Enghoffosoma chanabunae Srisonchai & Likhitrakarn sp. n. 

Ruttapon Srisonchai, Theemaporn Benchapong, Sergei I. Golovatch, Chirasak Sutcharit, Somsak Panha, Natdanai Likhitrakarn. 2026. Integrative Taxonomy of the Millipede Genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae) in Thailand and Myanmar, with Descriptions of Three New Species. Ecologica Montenegrina. 99; 40-71. DOI: doi.org/10.37828/em.2026.99.4 [2026-06-24] 

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[Botany • 2026] Begonia quadricornualata (Begoniaceae, sect. Platycentrum) • A New Species from Arunachal Pradesh, India

 

Begonia quadricornualata R.Maity, U.L.Tiwari & S.S.Dash, 

in Maity, Tiwari et Dash, 2026. 

DOI: doi.org/10.1002/njb.05000 

 

Abstract

A new species of Begonia L. (Begoniaceae) belonging to sect. Platycentrum is described from the Papum Pare District of Arunachal Pradesh, India. This new species is distinguished by its dense, long, red to brown hispid hairs on the adaxial leaf surface of female plants, caducous stipules and bracts, and solitary-flowered female inflorescence. Additionally, it features dense, woolly brown ovaries and fruits, the latter bearing four fully developed wings. This paper includes a detailed description, photographic plate, line drawings, a comparative morphological table and a dichotomous key to distinguish the new taxon from closely allied species.

Keywords: Arunachal Pradesh, Begonia sect Platycentrum, Eastern Himalaya, four-horned begonia, northeast India

  Begonia quadricornualata sp. nov. (female plant).
 (A) General habit, (B) leaf showing red hispid hairs, (C) habit showing open female flower, (D) habit showing position of female inflorescence from side view, (E) bracts, (F) outer tepal, (G) inner tepal, (H) gynoecium, (I) fruit, (J) T.S. of fruit.

Illustration of Begonia quadricornualata sp. nov.

 female plant
 (A) Habit, (B) bract, (C) outer tepal, (D) inner tepal, (E) gynoecium, (F) fruit.

male plant
(A) Habit, (B) inflorescence, (C) male flower, (D) bract, (E) outer tepal, (F) inner tepal, (G) stamen.

 Drawings by D. K. Sah.

Begonia quadricornualata R.Maity, U.L.Tiwari & S.S.Dash sp. nov. 

Diagnosis: A species morphologically allied to B. handelii Irmsch. and B. tessaricarpa C.B.Clarke, but distinguished by its dense, long, red to brown hispid hairs on the adaxial leaf surfaces of female plants (versus glabrescent in B. handelii, and puberulous in B. tessaricarpa), and its significantly longer petioles, measuring 40–50 cm (versus up to 25 cm in B. handelii and 14–31 cm in B. tessaricarpa). It is further characterized by caducous stipules and bracts (versus persistent in both B. handelii and B. tessaricarpa) and male flowers with 40–60 stamens (versus 70–100 in B. handelii, and 12–20 in B. tessaricarpa). Additionally, female inflorescences are solitary-flowered (versus 2–4-flowered in B. handelii and 2–5-flowered in B. tessaricarpa) and the fruit is 4-winged (versus 4-ridged in B. handelii, and 4-horned in B. tessaricarpa).

Etymology: The specific epithet ‘quadricornualata' refers to the unique development of the four ovary horns into four distinct wings on the fruit. The name is derived from the Latin ‘quattuor' (four), ‘cornu' (horn), and ‘alatus' (winged).

Rohan Maity, Umeshkumar L. Tiwari and Sudhansu Sekhar Dash. 2026. Begonia quadricornualata sp. nov. (Begoniaceae) from India. Nordic Journal of Botany. DOI: doi.org/10.1002/njb.05000 [07 July 2026] 

[Botany • 2026] Corybas apoensis (Orchidaceae: Diurideae) • A New endemic Species from Mt. Apo, Philippines

  

Corybas apoensis Agad, Tandang, R.Bustam. & Salas, 

in Agad, Mallari, Tandang, Bustamante et Salas, 2026.

DOI: doi.org/10.11646/phytotaxa.766.1.6 

facebook.com: Philippine Taxonomic Initiative

Abstract

Corybas apoensis, a new species of tribe Acianthinae endemic to Mindanao, is described herein. Among the currently recognized species of the tribe, it most closely resembles C. viridisepalus in overall floral morphology, particularly in having upright flowers borne on a slender, cylindric, erect ovary and the flower possessing a dorsal sepal and labellum tinged with conspicuous carmine veins. Both species further share a pilose labellum. However, this notable new species is distinct by having white floral background and a dorsal sepal being densely papillate on the abaxial surface. We also provided a summarized key morphological characteristics that distinguishes our new species from the later.

Monocots, Helmet orchid, Rare, Diurideae, Malesia, Taxonomy

 

Corybas apoensis Agad, Tandang, R.Bustam. & Salas, sp. nov.

KIER C. AGAD, ARCHELITO L. MALLARI, DANILO N. TANDANG, RENE ALFRED ANTON BUSTAMANTE and DARYL S. SALAS. 2026. Corybas apoensis (Orchidaceae), A New endemic Species from Mt. Apo, Philippines.  Phytotaxa. 766(1); 84-90. DOI: doi.org/10.11646/phytotaxa.766.1.6 [2026-07-07]

  facebook.com/phtaxa/posts/1419584753561021

 

[Entomology • 2026] Neophlugiolopsis? nigrivertex • One New Species of the Genus Neophlugiolopsis Pan & Bian, 2024 (Orthoptera: Tettigoniidae: Meconematinae) from Guangxi, China

 

Neophlugiolopsis? nigrivertex  

 Xiong, Chen & Shi, 2026

 

DOI: doi.org/10.11646/zootaxa.5845.1.12 

The genus Neophlugiolopsis was established by Pan & Bian (2024), with Neophlugiolopsis longiprocera Pan & Bian, 2024 as the type species. It resembles Phlugiolopsis Zeuner, 1940, Paraphlugiolopsis Bian & Shi, 2014, Aphlugiolopsis Wang, Liu & Li, 2015, Pseudophlugiolopsis Wang, Zhou & Chang, 2020 and Guangxia An, Chen & Shi, 2023, but can be distinguished from them mainly by the following characters: tegmina reach the middle of abdomen, and posterior area of male tenth abdominal tergite is strongly produced posteriorly into a long process. 

 

In this paper, a new species from Guangxi, China is described and tentatively assigned to the genus Neophlugiolopsis. All type specimens are deposited in the Museum of Hebei University.

Neophlugiolopsis? nigrivertex sp. nov., male:
A–B. head and pronotum: A. dorsal view; B. lateral view; C–F. apex of abdomen: C. ventral view; D. latero-ventral view; E. dorsal view; F. lateral view.

Neophlugiolopsis? nigrivertex sp. nov., male in habitat.

Neophlugiolopsis? nigrivertex sp. nov.

 Chinese name: 黑顶新吟螽 

Etymology. The specific name refers to the black dorsal surface of the vertex. It is derived from Latin nigri-, meaning “black”, and vertex, meaning “top of the head”.

ZIJIAN XIONG, KEDA CHEN and FUMING SHI. 2026. One New Species of the Genus Neophlugiolopsis Pan & Bian, 2024 (Orthoptera: Tettigoniidae: Meconematinae) from Guangxi, China.  Zootaxa. 5845(1); 197-200. DOI: doi.org/10.11646/zootaxa.5845.1.12 [2026-07-03]

[Botany • 2026] Nymphoides crucioides, N. miniata, N. thailandica, ... • Integration of Cytogenetics and Morphology for the Taxonomic Revision of Nymphoides (Menyanthaceae) in Thailand including Five New Species

 

Nymphoides miniata Noppornch. & Suwanph.,

in Suwanphakdee, Nopporncharoenkul et Hodkinson, 2026. 

DOI: doi.org/10.1016/j.aquabot.2026.104044 

facebook.com: Flora of Thailand

Highlights: 

• A full revision of Thai Nymphoides based on morphology and cytogenetic evidence is provided.

• Cytogenetic and morphological evidence supports Nymphoides species boundaries.

• Ten Nymphoides species are described in Thailand, including five new endemic species

Abstract 

A taxonomic revision of Nymphoides (Menyanthaceae) in Thailand was undertaken using cytogenetics and morphology. Specimens were collected throughout Thailand. Cytogenetic analyses were conducted on living collections. The morphological characters used in the descriptions of all taxa were measured from living specimens, which were subsequently maintained for ex-situ conservation. We provide a new generic description, a key to species, and descriptions of all species. Line drawings are provided for the five new species and color photographic illustrations are provided for all taxa. Herbarium voucher specimens were prepared and deposited in Thai herbaria. Ten species are recognized in Thailand, including five new species described in the present study for the first time, namely Nymphoides chumphonense, N. crucioides, N. miniata, N. thailandica and N. thungyaiense. Cytogenetic evidence supports the proposed species boundaries of Thai Nymphoides.

 

Keywords: Aquatic plants, Terrarium plants, Tropical plants, Wetland

Nymphoides chumphonense Suwanph. 

Nymphoides crucioides Suwanph. & Hodk. 

Nymphoides miniata Noppornch. & Suwanph. 

Nymphoides thungyaiense Suwanph & Noppornch. 

 Nymphoides thailandica Suwanph. 

 

Chalermpol Suwanphakdee, Nattapon Nopporncharoenkul and Trevor R. Hodkinson. 2026. Integration of Cytogenetics and Morphology for the Taxonomic Revision of Nymphoides (Menyanthaceae) in Thailand including Five New Species. Aquatic Botany. 207, 104044. DOI: doi.org/10.1016/j.aquabot.2026.104044 

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🌸 มหาวิทยาลัยเกษตรศาสตร์ พบบัวบาชนิดใหม่ของโลก 5 ชนิด จากแหล่งน้ำของประเทศไทย บ่งชี้ถึงความร่ำรวยทรัพยากรพืชของไทย และความสำคัญของงานวิจัยวิทยาศาสตร์พื้นฐาน ของสถาบันการศึกษา💚

นักวิจัย มก นำโดย รศ.ดร. เฉลิมพล สุวรรณภักดี จากภาควิชาพฤกษศาสตร์ คณะวิทยาศาสตร์ มหาวิทยาลัยเกษตรศาสตร์ ค้นพบบัวบาชนิดใหม่ของโลกจำนวน 5 ชนิด โดยตีพิมพ์เผยแพร่การค้นพบนี้ในวารสาร Aquatic Botany ฉบับปี 2026 การค้นพบครั้งนี้เป็นการบูรณาการวิจัยร่วมระหว่างหน่วยงานซึ่งประกอบด้วย ดร. ณัฐพล นพพรเจริญกุล นักวิจัย จากองค์การพิพิธภัณฑ์วิทยาศาสตร์แห่งชาติ (อพวช) และ Prof. Dr. Trevor R. Hodkinson จากทรินิตี้คอลเลจ มหาวิทยาลัยแห่งกรุงดับลิน ประเทศไอร์แลนด์ โดยเป็นการบูรณาการวิจัยร่วมทางด้านอนุกรมวิธานพืช สัณฐานวิทยาและเซลล์วิทยา 

.

รศ.ดร. เฉลิมพล สุวรรณภักดี  กล่าวว่า พืชสกุลบัวบา (Nymphoides) วงศ์ Menyanthaceae เป็นพืชน้ำที่มีการใช้ประโยชน์ทางด้านการใช้เป็นพืชประดับ และผักพื้นบ้าน ในประเทศไทยเคยมีรายงาน 5 ชนิด จากการศึกษาวิจัยพบว่า บัวบาของประเทศไทยมีทั้งหมด 10 ชนิด และเป็นพืชชนิดใหม่ของโลก 5 ชนิด แต่ละชนิดมีศักยภาพในการนำไปใช้ประโยชน์ทางด้านการใช้เป็นพืชประดับสวนขวด สวนกระจก และตู้ไม้น้ำ อีกทั้งยังเน้นให้ตระหนักถึงการอนุรักษ์แหล่งน้ำจืด ซึ่งเป็นที่อยู่ของพืชชนิดเหล่านี้ บ่งชี้ถึงความร่ำรวยทรัพยากรพืชของประเทศไทย และความสำคัญของงานวิจัยวิทยาศาสตร์พื้นฐาน 

บัวบา 5 ชนิดนี้ประกอบด้วย 

.

1. สายติ้ง หรือ สายติ่ง Nymphoides chumphonense Suwanph. 

 พบในแหล่งน้ำจืดตื้นๆ บริเวณ จ. ชุมพร และสุราษฎร์ธานี  โดยตั้งชื่อตาม จ. ชุมพร สถานที่พบตัวอย่างเป็นที่แรก พืชชนิดนี้มีการใช้ประโยชน์เป็นผักพื้นบ้าน โดยกินสดกับน้ำพริก และมีขายในตลาด ถูกจัดสถานะใกล้สูญพันธุ์ (Endangered: EN) เนื่องจากการสูญเสียแหล่งที่อยู่ และการรุกรานของน้ำเค็ม อีกทั้งติดผลและเมล็ดได้น้อย

.

2. บัวบาหัวลูกศร Nymphoides crucioides Suwanph. & Hodk. 

 พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี และยะลา โดยตั้งชื่อตามลักษณะดอกที่คล้ายกากบาท  พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

.

3. บาจิ๋ว Nymphoides miniata Noppornch. & Suwanph. 

 เป็นบัวบาที่มีขนาดเล็กที่สุดของโลก พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี โดยตั้งชื่อตามลักษณะดอกที่มีขนาดเล็กที่สุดของโลก พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

.

4. บาสยาม Nymphoides thailandica Suwanph. 

บัวบาที่มีขนาดเล็ก พบในแหล่งน้ำจืดตื้นๆ และในนาข้าว บริเวณ จ. ปราจีนบุรี กระบี่ และยะลา โดยตั้งชื่อตามประเทศไทย พืชชนิดนี้มีศักยภาพในการนำไปใช้เป็นไม้ประดับสวนขวด สวนกระจก และตู้ไม้น้ำ มีทั้งที่มีฟอร์มน้ำและฟอร์มบก ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากการสูญเสียแหล่งที่อยู่ และการใช้สารกำจัดวัชพืช  

.

5. ดาวรดา Nymphoides thungyaiense Suwanph & Noppornch. 

เป็นบัวบาที่มีขนาดค่อนข้างใหญ่ พบในแหล่งน้ำจืดตื้นๆ บริเวณ จ. ตาก โดยตั้งชื่อตามแหล่งที่พบ ถูกจัดสถานะใกล้สูญพันธุ์อย่างยิ่ง (Critically Endangered - CR) เนื่องจากพบประชากรเพียงแหล่งเดียว  

[Mammalogy • 2026] Revalidation of Manis aurita based on integrative genomic and morphological evidence

 

Manis aurita Hodgson, 1836

in Koju, Zeng, Zhang, Yao, Huang, Wang, Hawkins, Hinckley, Flores, Guo, Li, Maharjan, Byanjankar, Huang, Yu, Leng, He, Feijó et Hua, 2026. 

 DOI: doi.org/10.1038/s42003-026-10314-9

Abstract

Pangolins face severe conservation threats globally, necessitating accurate taxonomy for effective conservation. Previous genomic research on the Chinese pangolin (Manis pentadactyla) identified two deeply divergent lineages (MPA and MPB), suggesting underestimated species diversity. The recent description of M. indoburmanica (corresponding to MPB), however, did not assess its relationship with historical names, particularly the subspecies M. p. aurita, leaving the group’s taxonomic status uncertain. To resolve this issue, we employed an integrative framework, analyzing genomic and morphological data from museum specimens including the lectotype of M. p. aurita to clarify phylogenetic relationships and taxonomy within M. pentadactyla sensu lato. Our results demonstrate that MPB, which includes both M. indoburmanica and M. p. aurita, is geographically restricted to the southern Himalayas and thus distinct from other M. pentadactyla populations. Genomic analyses indicate the two clades diverged ~1.8 Ma and have remained largely isolated, with only minimal gene flow with congeners. Furthermore, morphometric analyses of both cranial and external features reveal consistent and significant differentiation between the Himalayan lineage (MPB) and M. pentadactyla sensu stricto. Collectively, these congruent findings provide unequivocal support for the revalidation of Manis aurita Hodgson, 1836, thereby establishing it as a distinct extant species of Asian pangolin.

Manis aurita Hodgson, 1836

 

 

Narayan Prasad Koju, Zeling Zeng, Guihua Zhang, Zhicheng Yao, Xia Huang, Xiaoyun Wang, Melissa T. R. Hawkins, Arlo Hinckley, Mary Faith C. Flores, Ce Guo, Jun Li, Devendra Maharjan, Saraswoti Byanjankar, Lianghua Huang, Wenhua Yu, Liang Leng, Kai He, Anderson Feijó and Yan Hua. 2026. Revalidation of Manis aurita based on integrative genomic and morphological evidence. Communications Biology. 9: 770. DOI: doi.org/10.1038/s42003-026-10314-9 [01 July 2026]

  

[Crustacea • 2025] Cambarus ocoeensis • A New Species of Crayfish (Decapoda: Cambaridae) from the Ocoee River Basin of Tennessee, USA

 

Cambarus ocoeensis 

 Thoma & Williams, 2025 

DOI: doi.org/10.11646/zootaxa.5618.1.7  

  Researchgate.net/publication/390392403 

  facebook.com/RogerFThoma

Abstract

Cambarus ocoeensis sp. nov. is described from the Ocoee River Gorge area of Tennessee. The species is morphologically most similar to Cambarus hiwasseensis Hobbs, 1981, but differs from C. hiwasseensis in the following ways: first pereopod chelae in general less sculpted, usually with one row of palmer tubercles, if second row present very weakly developed and consisting of six or fewer tubercles, lateral margin not costate, lateral impression weak at most but usually absent; areola length 2.5–4.1 times greater than width, having 5–8 punctations in narrowest part; size at maturity smaller with MI ranging from 21.8 to 30.9 mm total carapace length (24.6–40.0 mm in C. hiwasseensis); and antennal scale broadest at midpoint. It can be distinguished from other morphologically similar species—those with acuminate rostra—from the Hiwassee River basin of northern Georgia, southeast Tennessee, and western North Carolina area in lacking lateral rostral spines/tubercles and cervical spines. Cambarus ocoeensis sp. nov. is genetically most closely related to an undescribed member of the Cambarus bartonii (Fabricius, 1798) species complex.

 

Crustacea, C. hiwasseensis, C. bartonii, C. howardi, Hiwassee River basin, species complex

Type material of Cambarus ocoeensis sp. nov.:
 A–C, E, G–I, holotype MI (OSUMC #10897); F, allotypic paratypefemale (OSUMC #10898); D, morphotypic paratype MII (OSUMC #10899).
A–carapace, dorsal view; B–gonopod, lateralview; C–gonopod, mesial view; D–gonopod, lateral view; E–antennal scale, dorsal view; F–annulus ventralis; G–pereopods3–5, right side ventral views; H–epistome; I–right chela, dorsal view. 

Color image of Cambarus ocoeensis sp. nov., holotype (OSUMC #10897).

Image captured in lab under artificial light.

Cambarus ocoeensis sp. nov. 

Roger F. THOMA and Bronwyn W. WILLIAMS. 2025. Cambarus ocoeensis, A New Species of Crayfish (Decapoda: Cambaridae) from the Ocoee River Basin of Tennessee, USA.  Zootaxa. 5618(1); 106-118. DOI: doi.org/10.11646/zootaxa.5618.1.7 [2025-04-01] 

 Researchgate.net/publication/390392403_Cambarus_ocoeensis_a_new_species_of_crayfish_from_the_Ocoee_River_Basin_of_Tennessee_USA

  facebook.com/RogerFThoma/posts/10228373600190065

[Paleontology • 2026] New Fossil unveils the Dentary Anatomy of the rare lepidosauromorph Cargninia enigmatica from the Upper Triassic of Southern Brazil

 Cargninia enigmatica Bonaparte, Schultz, Soares & Martinelli, 2010, 

in Damke, Dalle-Laste,  Kerber, Roberto-da-Silva et Müller, 2026. 

DOI: doi.org/10.1002/ar.70268

Artwork by Márcio L. Castro.

Abstract

Nowadays, Lepidosauria is a diverse group represented by squamates and the rhynchocephalian tuatara. However, their origins date back to the end of the Permian Period, when the stem lineages Lepidosauromorpha and Archosauromorpha diverged. Within the lepidosauromorph stem lineage, a few species have been described with unstable phylogenetic placements. In Brazilian strata, the only representative of this lineage is the rare species Cargninia enigmatica. The taxon was found in Norian beds (upper portion of the Candelária Sequence) and is known from the posterior end of a dentary and a tentatively referred partial maxilla. Here, we describe a new specimen (CAPPA/UFSM 0519) referred to Cargninia enigmatica, consisting of a partial dentary recovered from the type locality. The specimen provides new anatomical information on the anterior portion of the dentary. In addition, for the first time, the phylogenetic placement of Cargninia enigmatica was investigated in a computational framework, and the results are congruent with previous suggestions that it represents a non-lepidosaur lepidosauromorph. Finally, the new specimen reveals details of the anatomy of the mandibular ramus of the trigeminal nerve, demonstrating that it shares a similar condition with its extant relatives.

Keywords: Caturrita Formation, Gondwana, Lepidosauromorpha, Norian, Pangea

Left dentary of CAPPA/UFSM 0519 in lateral view. (a) Photograph. (b) Interpretative drawing. (c) Detail of the rostralmost preserved teeth. (d) Additional details of the second and third preserved teeth. fo, foramina.

SYSTEMATIC PALEONTOLOGY

DIAPSIDA Osborn, 1903

SAURIA Macartney, 1802

LEPIDOSAUROMORPHA Gauthier, 1984 (sensu Gauthier et al., 1988)

Cargninia enigmatica Bonaparte et al., 2010

 Holotype: UFRGS-PV-1027-T, posterior end of a left dentary with five preserved teeth.

 Type locality, age, and horizon: Linha São Luiz site (29°33′45″S, 53°26′48″W), municipality of Faxinal do Soturno, Rio Grande do Sul, Brazil. Candelária Sequence of the Santa Maria Supersequence (Horn et al., 2014; Zerfass et al., 2003). These layers are early Norian in age (225.42 ± 0.37) (Langer et al., 2018).

New referred material: CAPPA/UFSM 0519, partial left dentary with several teeth. The specimen was collected from the type locality.

  Amended diagnosis: In addition to the unique traits of Cargninia enigmatica listed by Bonaparte et al. (2010) and Romo de Vivar et al. (2020), we recognize the following additional features based on the study of CAPPA/UFSM 0519 that distinguish the species from closely related taxa: the symphyseal region is medially recurved and formed by the Meckelian groove, and the presence of foramina restricted to the anterior region of the dentary.

A landscape from the Late Triassic of southern Brazil showing Cargninia enigmatica being observed by the sauropodomorph dinosaur Macrocollum itaquii. Artwork by Márcio L. Castro.

Lísie Vitória Soares Damke, Vitória Zanchett Dalle-Laste, Leonardo Kerber, Lúcio Roberto-da-Silva and Rodrigo Temp Müller. 2026. New Fossil unveils the Dentary Anatomy of the rare lepidosauromorph Cargninia enigmatica from the Upper Triassic of Southern Brazil. The Anatomical Record. DOI: doi.org/10.1002/ar.70268  [04 July 2026]

[Botany • 2024] Thrixspermum yei (Orchidaceae: Epidendroideae: Aeridinae) • A New Species from Xizang, China

Thrixspermum yei M. K. Li, Z. Xing & Y. Luo, 

in Li, Pang, Chen, Deng, Wang, Xing et Luo, 2024. 

叶氏白点兰 | taiwania.ntu.edu.tw/abstract/1988

Abstract

A new species Thrixspermum yei M. K. Li, Z. Xing & Y. Luo is described and illustrated from Xizang, China. It is similar to T. fantasticum and T. duplocallosum but differs from them in having a yellowish, ovate, indistinctly 3-lobed lip, disk with 4 yellow papillary front calli densely covered with yellow powder, and base with a cone-like appendage.

Keyword: Thrixspermum fantasticum, Thrixspermum duplocallosum, taxonomy, Xizang, China

Thrixspermum yei M. K. Li, Z. Xing & Y. Luo  
 A-B. plant and inflorescence; C. inflorescence; D-F. flower; G-H. split flower; IK. close-up of lip; L. lip and column. M-N. column and ovary; O-P. anther cap; Q. pollinia. (Photos: by Shen-shen Pang).

Thrixspermum yei M. K. Li, Z. Xing & Y. Luo
A. plant and inflorescence; B. inflorescence; C. flower; D. split flower; E1-E2. Column. F1-F2. close-up of lip; G. anther cap; H. pollinia.

 (Drawn by Jian-ping Deng).

Thrixspermum yei M.K.Li, Z.Xing & Y.Luo, sp. nov. 

 Type: CHINA. Xizang, Mêdog County, Dexing village, 95°14'42.89"E, 29°13'48.19"N, 1184 m, epiphytes on tree trunks in evergreen broad-leaved forest, 08 June 2022, M. K. Li 2022160 holotype: T000167542 (TAAHUC!); isotype: 022016 (XZE!). 

Diagnosis: Thrixspermum yei is morphologically similar to T. fantasticum and T. duplocallosum, but can be clearly distinguished from them by having a yellowish, ovate, indistinctly 3-lobed lip, disk with 4 yellow papillary front calli densely covered with yellow powder, and base with a cone-like appendage.

  

Etymology: The specific epithet "yei" is in honor of Mr. De-Ping YE (1977-2020), who dedicated over twenty years to the botanical surveys and conservation of wild orchids in China. 

Vernacular name: Ye Shi Bai Dian Lan, 叶氏白点兰

Meng-Kai Li, Shen-Shen Pang, Xue-Da Chen, Jian-Ping Deng, Wei Wang, Zhen Xing and Yan Luo. 2024. Thrixspermum yei (Aeridinae, Vandeae, Epidendroideae, Orchidaceae) - A New Species from Xizang, China. Taiwania. 69(2); 138-141. DOI: 10.6165/tai.2024.69.138

 taiwania.ntu.edu.tw/abstract/1988 

[Botany • 2022] Gentiana ranae (Gentianaceae) • A New Species from Himachal Pradesh, India revealed by morphology and molecular phylogeny

Gentiana ranae M.Shabir & M.D.Dwivedi, 

in Shabir, Pal, Dwivedi et Tiwari, 2022.

 Rheedea. 32(1);  

rheedea.in/IAAT/article/view/898 

 

Abstract 

A new species of the genus Gentiana, G. ranae sp. nov., from Rohtang Pass in Himachal Pradesh (India) is described here. The species shows morphological resemblances with G. glauca Pall. and G. venusta Wall. of sect. Monopodiae, but differs in key morphological characters such as a stem with 4–6 fine lines, upper stem leaves more densely enveloping the inflorescence, a light blue corolla with margins of the corolla lobes scabrous to crenulate, horizontally truncate plicae, dentate to erose margins, and ovate to sub-orbicularseeds. Along with the morphological characters the species is supported as a new member of sect. Monopodiae in a molecular phylogenetic analyses using the nuclear ribosomal DNA internal transcribed spacer (ITS) and chloroplast trnL-F intergenic-spacer regions. The species is described, illustrated and discussed.

Keywords: cpDNA trnL-F, Gentiana glauca, nrDNA ITS, sect. Monopodiae, western Himalaya

 

Gentiana ranae M.Shabir & M.D.Dwivedi:
b. Flowering plant; c. Habit; d. Seeds

 (Photos by Mohd Shabir).

Gentiana ranae M.Shabir & M.D.Dwivedi: a. Habitat

 (Photos by Mohd Shabir).

Gentiana ranae sp. nov.

Shabir M., Pal A.K., Dwivedi M.D. and J.K. Tiwari. 2022. Gentiana ranae (Gentianaceae), A New Species from India revealed by morphology and molecular phylogeny.  Rheedea. 32(1); 30-45. DOI: 10.22244/rheedea.2022.32.01.01 https://rheedea.in/IAAT/article/view/898 

 

[Herpetology • 2026] Boulenophrys dongli & B. raoping • Survival in Fragmented Landscapes: Two New Boulenophrys Species (Anura: Megophryidae) from the coastal mountains of eastern Guangdong, China

Boulenophrys raoping 

Wang, Zhan, Xiao, Tan, Chen, Li, Lin, Lyu & Zeng, 2026

Raoping horned toad | 饶平角蟾  ||  DOI: doi.org/10.3897/vz.76.e195550 

Abstract

Two new species, Boulenophrys dongli sp. nov. and Boulenophrys raoping sp. nov., are described from the coastal mountains of Raoping County, Chaozhou City, eastern Guangdong, China, based on morphological and molecular evidence. Currently, these two new species are known only from low-altitude mountainous areas (below 500 m a.s.l.), where they inhabit fragmented patches of secondary forest interspersed with bamboo plantations. Given their extremely limited distribution, ongoing habitat loss, and isolated populations, we recommend listing these two new species as Critically Endangered (CR) in accordance with IUCN Red List criteria B1ab(i,ii,iii)c(i,ii)+B2ab(i,ii,iii)c(i,ii). Continued research is critical to fill existing knowledge gaps and to guide coordinated conservation efforts aimed at protecting these unique amphibians within the critical montane refugia of eastern Guangdong.

Keywords: Conservation, highly threatened, horned toads, morphology, mountain refugia, phylogeny

Life aspect of the holotype (GEP a514) of Boulenophrys dongli sp. nov.:
 A dorsolateral view, B dorsal view, C ventral view, D ventral view of hand, E ventral view of foot.

 

Boulenophrys dongli sp. nov. 

Dongli horned toad (in English) 

dōng lǐ jiǎo chán (东里角蟾 in Chinese)

 

Etymology. The specific epithet dongli is derived from the ancient name “东里 (dōng lǐ)” of the region where the new species is distributed, meaning the eastern boundary of Guangdong, indicating the distribution area of the new species.

Diagnosis. (1) Small size (SVL 30.3–35.6 mm in six adult males); (2) snout rounded in dorsal view, canthus rostralis well developed, tongue not notched distally; (3) tympanum visible, tympanic margin indistinct, temporal region bearing conical spines except tympanum; (4) vomerine ridges and vomerine teeth present; (5) dorsal skin rough and highly granular, discontinuous X-shaped or V-shaped ridge on center of dorsum, dorsolateral ridges absent, sparse large tubercles on flanks, dorsal limbs with tubercles bearing conical spines, sparse rounded tubercles on belly, dense tubercles ...

Life aspect of Boulenophrys raoping sp. nov.
the holotype (GEP a519): A1 dorsolateral view, A2 dorsal view, A3 ventral view, A4 ventral view of hand, A5 ventral view of foot;
female paratype (GEP a523): B1 dorsolateral view, B2 dorsal view, B3 ventral view, B4 ventral view of hand, B5 ventral view of foot.

Boulenophrys raoping sp. nov. 

Raoping horned toad (in English) 

 ráo píng jiǎo chán (饶平角蟾 in Chinese)

Etymology. The specific epithet raoping is derived from its type locality, Raoping County. The name “Raoping (饶平)” combines the characters “饶” (ráo), meaning “rich” or “abundant,” symbolizing prosperous harvests and natural resources, and “平” (píng), meaning “peace” or “stability,” reflecting aspirations for social harmony. Together, they convey the traditional Chinese ideal of material prosperity and enduring peace.

Diagnosis. (1) Small size (SVL 34.1–36.6 mm in four adult males, 39.2–41.8 mm in three adult females); (2) snout rounded in dorsal view, canthus rostralis well developed, tongue not notched distally; (3) tympanum distinct, tympanic margin distinct and raised, temporal region bearing conical spines except tympanum; (4) vomerine ridges present, vomerine teeth absent; (5) dorsal skin rough and highly granular, discontinuous X-shaped or V-shaped ridge on center of dorsum, dorsolateral ridges absent, sparse large tubercles on flanks, dorsal limbs with tubercles bearing conical spines, dense rounded tubercles on belly, dense tubercles bearing conical spines on ...

 Jian Wang, Bin-Bin Zhan, Wei-Wen Xiao, Lin Tan, Hong-Hui Chen, Yuan-Hang Li, Shi-Shi Lin, Zhi-Tong Lyu and Zhao-Chi Zeng. 2026. Survival in Fragmented Landscapes: Two New Boulenophrys Species (Anura: Megophryidae) from the coastal mountains of eastern Guangdong, China. Vertebrate Zoology. 76: 381-399.  DOI: doi.org/10.3897/vz.76.e195550 [02-07-2026]